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SMITHSONIAN INSTITUTION
UNITED STATES NATIONAL MUSEUM

Bulletin 132

REVISION OF THE
NORTH AMERICAN MOTHS OF THE
SUBFAMILIES LASPEYRESIINAE
AND OLETHREUTINAE

CARL HEINRICH

Of the Bureau of Entomology, United States Department of Agriculture

WASHINGTON
GOVERNMENT PRINTING OFFICE
1926

ADDITIONAL COPIES

OF THIS PUBLICATION MAY BE PROCURED FROM
THE SUPERINTENDENT OF DOCUMENTS
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75 CENTS PER COPY

The scientific publications of the National Museum consist of two
series—Proceedings and Bulletins.

The Proceedings, the first volume of which was issued in 1878, are
_intended primarily as a medium for the publication of original
papers based on the collections of the National Museum, setting
forth newly acquired facts in biology, anthropology, and geclogy
derived therefrom, or containing descriptions of new forms and
revisions of limited groups. One or two volumes are completed
annually and copies of each paper, in pamphlet form, are dis-
tributed as soon as published to libraries and scientific organizations
and to specialists and others interested in the different subjects. The
dates at which these separate papers are poeeee are recorded in
the table of contents of the volume.

The Bulletins, the first of which was issued in 1875, consist of a
series of separate organizations comprising chiefly monographs of
large zoological groups and other general systematic treatises (occa-
sionally in several volumes), faunal works, reports cf expeditions,
and catalogues of type specimens, special collections, etc. The
majority of the volumes are octavos, but a quarto size has been
adopted in a few instances in which large plates were regarded as
indispensable.

Since 1902 a series of octavo volumes containing papers relating to
the botanical collections of the Museum, and known as the Contri-
butions from the National Herbarium, has been published as
bulletins.

The present work forms No. 132 of the Bulletin series.

Wasuineton, D. C., December 7, 1925.

Wiccan WETMORE,
Assistant Secretary,
In charge of the United States National Museum.

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CONTENTS

Page

TE HUEY TCL BIE) 0 i = NSS ae ee See eee eee a
VE PSS URTV ECTE Le PSN 7a nae Nd ot & i Et ee ge Le Ee EO 2
ASST Ga ptOleee te: Seems Ae. ei TRY Ee A ees 3
Characters of the subfamilies of Olethreutidae_________-______-________ 4
anvaihabits and economic importance. 22-8 5
Comparative table of structural characters: Laspeyresiinae_____________ 6
Key tosune: renera, ob Laspeyresiinaes: 2 26 te 6
Paylorenetie tree of the Haspeyresiinae:—- = 2 ee a eee 7
BREVIS} Tas OK oY 1 Fl 0: Yes Sa ene es One ED EYAL DE a tg ee ee we NSA UE OR 8
EMAC HOT AID [pees ee weet eee Seas ee OE ee ee ee 9

See A Tso eee ee Pn ee RE SPREE ee ee 17

ADE RTC UNL peer ae een aes ee Shee et eek ee 18

7, “ANEW i covey ty) ees ee SEER 2 cies ee Ee Ape See) Ree ee ae eae Ee 19

Game miimen ese ha 2 ie pe ES ne) eee 20

Fel DY EN ER OY gts SA Sa Bae ee ea nae SP 23

Sb ASSET BW REV aes el ET a Ie ND eS a a I eee Aan 25

ee ETO Ese aes ee eens eee Seen ee ee ee 26

OR Ona Oe esis cee Sow Sa Pes het Set, WR CES i A ae 39

MMS STC UEOS ae uae 2S el epee tare a Bee ok 8 ee eee 41

UALS 61 8 eC Ie Wee a ee SS ee ee On ee Se Eee, ete 65

BLES VEC ESCO T) ELS eer ee ne teen). pe ge RS gy Se aa 66

Lied bs sol QTE OTULECIN GSE ssa a al ok A a a ee A a 69

MC VINMATIOTOSO IM chen a eee eke Re eS ee eee ee Tal

GSR MeH VOLO pines eee SAS Noe Be De ER ees Te eas oe 73
Comparative table of structural characters: Olethreutinae__________ 76
MersLOumerseneracot, OTe re tia Can Pe ee St ley 76
Ehylosenekie wree (0b the lOlethreutinae 2.8 a ee rir
EA BVERERS): Tes ol DOES TITUS pes we el cael 8 ne ee a8 a ep ea ee ee eo 78
Fi, TESA Ty pce he a a Sa a Se eae tO ee ed we RP et 81
SEOLYGHTOSIS IIIs .6Ne Eh ei ie fot eee, Ae eee sy ee Ae 87

PA ee ACY INN () She eae Se er As Beh eo Vee: A a pes Ve ais sees SEED 97

eo Send CEG aE SAi- ms mie gee TRS OE ag i ee 100
Geen yc) ees ere ep neaee fy Pees Pie A ee a ae a 106

Ty ARIE es pte if Bl ele Pi eS ee a SRLS 108

Same ENE) Gl ee ees na ee erate Om Cee Seen e eta ne SR IO ere UE ee 108
eS i eee are ee OEY BP eA tele LN eet a. 2 eae ks 109

AD TESTU CDV Abel ee See Do, DERE rk TEE en ee Ly Rae PS Oe 110

LOL OMA Ne eee eee Gee a eh oe tee i a 111

TAN OUST eek aN als 8 SS Rk a eve ener y 113

Serr SOLA EN are seen we eee yaa NM ROS cde rel eho Se 123

NEOs 1 CC Clemente see elon tan RAP ie eo os See ee ee 124

AE ACCASIO piI@E eee maven ning paar! ete ree he ESP cee Nae ee a 126

HG Se ORT eS TARA epee Mace A Hel aie eel lie A ial B= oe 128

LUT SELES TE ee gD cP UP a ea ge ORC CEB pI eA 161

SS Ole ATAU G Mere etl Sedat UF ele etey te ital meen all ib 166

OME Chir eee ROM Ne Gee i eh as ee eae 189
SDEGIeSerebenra hl eCaGisewiite re ten eh at wee ies De eee on he her eee 191
TRV OUR MANET INC OV RY conte og GRE ELS ll 1 = yo Ce yg va rT RE IESG CORI Re SULA Rea Ca Se 191
JOY G YE Se A pt ll 2 AT Le ol ORCI IO AEN Ce ag ee PB CIS aN 209

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adi

REVISION OF THE NORTH AMERICAN MOTHS OF THE SUB-
FAMILIES LASPEYRESTINAE AND OLETHREUTINAE

By Cari HetnricH
Of The Bureau of Entomology, United States Department of Agriculture.

INTRODUCTION

This paper is a continuation of the revisionary work of the family
Olethreutidae, begun in United States National Museum Bulletin
123,1 and treats of the two remaining subfamilies, Laspeyresiinae
and Olethreutinae. It is based chiefly upon the collections of the
United States National Museum,? the American Museum of Natural
History, the Canadian Department of Agriculture, and of Dr.
William Barnes, of Decatur, Ill. In addition I have examined the
types of the Museum of Comparative Zoology and the Philadelphia
Academy of Sciences. I have also received considerable material
from E. H. Blackmore and Dr. W. T. M. Forbes, and Dr. J. McDun-
nough has kindly loaned the genitalia slides of his types.

The classification here proposed follows that of my revision of
the Eucosminae and the method of treatment of genera and species
is the same except that characters of the female genitalia are in-
cluded and a drawing of these organs given for every species of
of which a female could be secured. For those species common to
Europe and North America purely European synonymy is omitted.
Otherwise generic and specific synonymy is given as fully as possible
and, except where so noted in the text, has been verified by com-
parison with types or other authentic specimens. In each case the
more important systematic references are given; and for species of
economic importance, citations to the papers giving the fullest ac-
counts of life history, distribution, and control.

Under the heading Distribution are given (unless otherwise stated
in the text) only the localities by States of the specimens which
I have examined and which are now in the National, Canadian

1 Revision of the North American Moths of the Subfamily BPucosminae of the Family
Olethreutidae, by Carl Heinrich, 1923.

2 Including the Fernald collection recently purchased by the United States Bureau of
Entomolegy and now incorporated in the national collection.

Oy BULLETIN 182, UNITED STATES NATIONAL MUSEUM

National, American Museum, and Barnes collections. Published
records which I have been unable to verify are omitted.

Food plant records are also given for each species, where known.
When such references are omitted the food plant is presumed to
be unknown.

Thirty-five genera, 223 species, and 9 varieties are recognized as
belonging to the two subfamilies. Of these, 16 genera, 34 species,
and 8 varieties are described as new. Five species now listed in the
Olethreutinae, but which must be referred elsewhere, are briefly
treated at the end of the paper.

HISTORICAL REVIEW

Tt is necessary here to call attention to three important works deal-
ing with the family and not mentioned in Bulletin 123. While the
latter was in press an extremely interesting paper by F. N. Pierce
and J. W. Metcalfe * appeared, figuring the male and female geni-
talia of the British Tortricoidea, describing these organs in detail
and proposing a classification for the group upon purely genetalic
characters. It is a valuable work and I would acknowledge my in-
debtedness to it. Unfortunately the authors ignored all other charac-
ters of the insect except genitalia and as a result have made some un-
natural groupings. In the Olethreutidae they make six group divi-
sions corresponding roughly to our three subfamilies, as follows:
Their Olethreutidii to our Olethreutinae; Ancylisidii and Epiblemi-
dii to part of the Eucosminae; Lipoptychidii to part of the Laspey-
resiinae; and their Ephippiphoridii (a heterogenous group) to the
remaining genera and species of our Eucosminae and Laspeyresiinae.
Their divisions are made entirely upon the form and number of the
signa of the female bursa, characters of generic rather than sub-
family value, and which do not hold even for the groups as they de-
fine them. Thorn-like signa are found in the Laspeyresiinae, Ole-
threutinae, and Eucosminae (Epiblimidii) ; and the transitions from
pocket to thorn-like and from thorn-like to “ pilleate” shapes are
gradual and nowhere clearly marked except between species or, at
most, genera.

Recently Dr. W. T. M. Forbes* has published a handbook of The
Lepidoptera of New York and Neighboring States, with generic and
specific keys and descriptions of the species occurring in the North-
eastern United States. In this the Olethreutidae are treated as a
subfamily of the Tortricidae. Most of the genera defined in my Re-
vision of the Eucosminae are retained and the classification there pro-
posed is more or less followed. The general treatment of the Tor-

* Genitalia of the group Tortricidae of the Lepidoptera of the British Islands, 1922, 34
pls., 101 pp.
*Cornell Univ. Agric. Exp. Sta., Memoir 68, 1924, pp. 376-476.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 3

tricoidea, however, is eclectic and the classification incorporates the
systems of several authors.

Kennel’s monograph of the European Tortricoidea,® mention of
which was unfortunately omitted from Bulletin 123, deals only with
such American species as are common to the two continents. It is an
elaborate work with full specific descriptions and fine color figures
of the moths. There is a detailed discussion of phylogeny and struc-
tural characters and numerous figures of the latter, but no keys
specific or other. The genitalia receive only occasional mention and
the classification departs but little from the Heinemann system.

CLASSIFICATION

In as much as a full family description and a key to the subfamilies
are given in Bulletin 123, it will be necessary here to note only a few
emendations. In my definition of the family I described vein 2 of
fore wing as “from the cell before outer three-fourths.” This char-
acter holds for all the American species except hemidesma Zeller
which I am making the type of a new genus (/vora). Here 2 comes
from cell well beyond three-fourths as in the Phaloniidae. On gene-
talic and other characters, however, Hvora is a perfectly good Ole-
threutid and goes in the Olethreutinae. Absence of the pecten on the
lower median vein of hind wing should also be noted in three genera
of the Laspeyresiinae (Goditha, Satronia, and Sereda). In males of
the laspeyresiin genera Hemimene and Balbis vein 8 of hind wing is
not free, but either fuses with 7 beyond cell (Hemimene) or (in
Blabis) anastomoses with it from slightly beyond base to well beyond
cell, somewhat as in the Pyralidae. Such exceptions make it practi-
cally impossible to frame a definition that will hold throughout for
the family unless we include characters of the male genitalia. On
these organs, however, the definition is clear-cut and permits of no
confusion. The Olethreutidae is a natural, well-marked family, much
easier to recognize than to describe in categorical terms.

The subfamilies also appear to be natural groups, in the main
clearly defined on hind wing venation; though here it is also neces-
sary to note a few exceptions. Some of the genera (Pseudogalleria
in the Eucosminae, /'ndothenia, Fsia, and E'pistmus in the Olethreu-
tinae, and Gymnandrosama and E'cdytolopha in the Laspeyresiinae)
are obviously transitional and on some of their characters could go in
other groups. Pseudogalleria has the reduced genetalia of the Las-
peyresiinae and the hind wing venation of the Eucosminae. On the
sum of its characters it seems to go better in the latter subfamily than
elsewhere. Another genus of the Eucosminae (Gwendolina) has
veins 3 and 4 of hind wing connate and 5 approximate to 4 at base,

5 Die Palaearktischen Tortriciden, Zoologica, Heft 54, vol. 21, Lfg. 1-4, 1908-1918.
54346—26——2

4 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

characters which would place it in the Olethreutinae did not genitalia
and the notched termen of fore wing show its clear relationship to
the EZ pinotia group of the Eucosminae. In the Olethreutinae, /'ndo-
thenia, and sia exhibit the venational characters of the Laspeyresiiae
(vein 5 or hind wing parallel to 4). They also have the general hab-
itus of the latter; but the genitalia are typically Olethreutin, Zndo-
thenia having a strongly developed uncus (not found in Laspeyres-
jinae), and both genera, the heavy spine cluster on sacculus of harpe
typical of the Olethreutiinae. Hndothenia forms (with Gymnan-
drosma and Ecdytolopha in the Laspeyresiinae) the connecting link
between the two subfamilies. /pisimus, which is here referred to the
Olethreutinae might go, on the other hand, as easily into the Eucos-
minae were it not for the normally connate condition of veins 3 and
4 of hind wing. Both the subfamilies Laspeyresiinae and Eucos-
minae seem to be derived directly from the Olethreutinae which ap-
pears to be the primitive group of the family.

The trees shown on pages 7 and 77 illustrate my conception of
the relations and phylogeny of the genera of the two groups here
treated, and the tables opposite pages 6 and 76 give comparative
analyses of the various structural characters upon which the genera
are classified. It will be necessary here only to expand the original
key to the subfamilies® by a brief summary of the characters dis-
tinguishing them.

CHARACTERS OF THE SUBFAMILIES OF OLETHREUTIDAE

Olethreutinae—Hind wing normally with vein 5 bent at base and
approximate to 4, veins 3 and 4 connate. (Where 5 is straight and
parallel with 4, harpe of male genitalia has at least one strong spine
cluster (Spc') always present; and where 3 and 4 are stalked
(E'pisimus tyrius) there are long flat spines on base of sacculus).
Thorax normally with strong posterior tuft (absent only in /’pzs7-
mus and Bactra). Fore wing with termen normally convex, rarely
concave (Hpisimus) and never notched; no costal fold in male.
Male genitalia with uncus normally present and well developed,
usually simple; socii usually well developed; harpe with at least one
and frequently two strong spine tufts (Spc, Spc?) from or near
sacculus; cuculli usually narrowly elongate; sacculus often with
strong basal spining (ScSp).

Eucosminae.—Hind wing with vein 5 always somewhat bent and
approximate to 4 at base; veins 3 and 4 stalked or united (connate
only in Gwendolina where termen of fore wing is notched, and sac-
culus of harpe is haired but without strong spine cluster). Thorax
seldom with posterior tuft. Fore wing with termen convex, con-

‘Bull. U. S. Nat. Mus. 128, p. 10, 1923.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 5

cave or notched; often with costal fold in male. Male genitalia
with uncus present or absent, when present simple, bifid or bifur-
cate; socii usually well developed; harpe rarely with strong spine
tufts on or near sacculus; cucullus usually rather broad in propor-
tion to its length, sometimes (in /’pinotia and allied genera) nar-
rowly elongate; sacculus smooth at base or weakly haired.

Laspeyresiinae—Hind wing with vein 5 always straight and par-
allel with 4; veins 3 and 4 connate or stalked. Fore wing with
termen convex or concave, rarely notched; costal fold usually absent
(present in a few species of Dichrorampha). Thorax without pos-
terior tuft (except in Gymnandrosoma, Ecdytolopha, and a few
tropical genera). Male genitalia with uncus absent; socii usually
absent; harpe simple, without spine clusters on or near sacculus;
cucullus rather broad in proportion to its length, very rarely nar-
rowly elongate; sacculus smooth at base or very weakly haired.

In female genitalia there are no definitive subfamily characters.
The differences (chiefly in the shape and number of the signa, the
shape of the genital plate and the chitinization and curvature of the
ductus) are of generic and specific rather than of larger group sig-
nificance. There is a certain habitus that tells one experienced with
the genitalia of the group whether a specimen belongs in one sub-
family or another; but it does not seem possible to express this in
any satisfactory description.

LARVAL HABITS AND ECONOMIC IMPORTANCE

A variety of larval habits prevail in the two subfamilies. In the
Olethreutinae the majority of the species feed externally on the leaves
or flowers of trees, shrubs, and low plants, either exposed or as leaf
tiers, folders, or rollers. Some are seed feeders in the capsules or
fruits and a few are stem or root borers in low plants. In the
Laspeyresiinae a larger percentage are internal feeders in fruits, nuts,
or seed capsules. Some are stem borers and a goodly number leaf
folders.

The latter subfamily contains several of our most important
economic insects, notably: The notorious codling moth (Carpocapsa
pomonella), the oriental peach moth (Grapholitha molesta), two
other serious fruit pests (G. packardi and G. prunivora), the pea
moth (Laspeyresia nigricana), the acorn moth (Melissopus latifer-
veanus), the Robinia gall maker (Hcadytolopha insiticiana) and a

.. group of spruce cone moths that do serious damage by the destruc-

tion of seeds (Laspeyresia piperana and allies).

In the Olethreutinae we have a spruce defoliator of importance
(Taniva albolineana) and the destructive grape berry moth (Poly-
chrosis viteana).

BULLETIN 132, UNITED STATES NATIONAL MUSEUM

There are several other species in the two subfamilies that are

potential enemies of serious import; but the above will serve to indi-
cate the economic importance of the family..

10.

13.

14.

Subfamily LASPEYRESIINAE

KEY TO THE GENERA OF LASPYRESIINAEB

. Thorax with posterior tuft_____________-_----------_-_----___--__- a eas
INNOTaAx without POSteLION tiitt= === 3}
. Male with hind tibia dilated, broadly tufted, and with heavy dorsal hair

pencile from base; female with ductus bursae short.
(15) Gymnandrosoma.
Male with hind tibia loosely scaled, but otherwise simple, no hair pencile

from base; female with ductus bursae long__-------~- (16) Ecdytolopha.

. Hind wing without pecten on lower median vein___-----------~-----~- 4,
Hind wing with pecten on lower median vein__-__-------.-------~------ 6.

. Hind wing with veins 6 and 7 parallel or subparallel___------------_- 5.

Hind wing with veins 6 and 7 approximate toward base (tortriciform).
(8) Sereda.

. Hind wing with veins 6 and 7 parallel; fore wing of male with 12 veins.

(1) Goditha.
Hind wing with veins 6 and 7 subparallel; fore wing of male with 11 veins.
(3) Satronia.

. Hind wing with veins 6 and 7 parallel or subparallel__--_----~----~----- 4.
Hind wings with veins 6 and 7 approximate toward base (or sometimes
in male with 7 running into BS Spe se ep hoe ge Bip ep thet Fey ete 10.
mEingi wine with veins) 6G-and (-parallel= === 8.
Hind wing with veins 6 and 7 subparallel______________-_____-------- 9.

. Male genitalia with socii developed, long, finger like; female with two signa

in bursa; COpwlatrixe=-4 204 vee be cen eters at eek eed fae ek (5) Talponia.
Male genitalia without socii; female with single signum.
(2) Dichrorampha.

. Male genitalia with socii developed ; female with single signum in bursa.

(4) Ricula.

Male genitalia without socii; female with two signa________ (7) Ethelgoda.
Head, palpi, legs, and underside of thorax covered with fine long hairs.

(12) Hedulia.

Head, palpi, legs, and underside of thorax normally scaled______--___- iat

. Male with veins 7 and 8 of hind wing fusing beyond cell____ (6) Hemimene.

Male with vein 8 of hind wing free; 7 approximate to 6 toward base____ 12.

. Male with hind tibia dilated and broadly tufted___--____-_ (13) Melissopus.

Maleswath: hind tubias smooth ;scaledai-s5=-._ fh. 34ers) eee Pee 13.
Abdomen of male with a pair of lateral tufts on eighth segment.
(9) Grapholitha.

PANG OTE O1TMY OL TTA ss SUN sa Se 14.
Male genitalia with a row of strong outer surface marginal spines near anal

angle ior cucullusso£ harpe 2 22) sh to) Srrieier eaeeeee (10) Ofatulena.
Male genitalia with outer surface of harpe unspined____~ (11) Laspeyresia.

(14) Carpocapsa.

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5 Talponia AT Sis ooo eS w---|l--- Ot eee ete (ee nes x x ---- known. :
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NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE

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8 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

1. GODITHA, new genus

(Fig. 24.)

Genoty pe-—Goditha bumeliana, new species (North America).

Thorax smooth.

Fore wing smooth; termen notched below apex, decidedly slant-
ing; 12 veins, all separate; 7 to termen; 11 from cell slightly before
middle; 10 well separated from 9; upper internal vein of cell from
between 10 and 11; 3, 4, and 5 remote at termen; 2 from cell
near 14, straight; no costal fold in male.

Hind wing without pecten on lower median vein; 8 veins; 6 and 7
separate and parallel; 3 and 4 connate; inner angle in male simple.

Hind tibia smooth scaled.

Male genitalia with outer surface of harpe unspined; cucullus
well defined, crescentiform; neck incurvation pronounced; neck
smooth; sacculus weakly haired. Uncus absent. Socii absent.
Gnathos a simple weakly chitinized band. Aedoeagus moderately
long, slender, tapering, slightly bent; cornuti a cluster of short
deciduous spines.

Abdomen of male with lateral hair tufts from sternite of eighth
segment.

A development from Dichrorampha. Monotypic and probably
tropical in origin.

Female unknown.

GODITHA RUMELIANA, new species

(Figs. 24, 293.)

Antenna fuscous. Palpus and face fuscous, faintly dusted with
sordid white. Head and thorax fuscous brown. Fore wing fuscous
with white markings, with distinct shading of ocherous on outer
half and with some obscure blackish streaking (visible only under
magnification); from mid dorsum three narrow irregular white
bands partially divided by blackish dusted streaks of the ground
color and forming a somewhat broken, outwardly curved white
dorsal patch extending to cell; costa faintly strigulated with white,
with somewhat broader spots of blackish fuscous between, the dark
spots extended (especially on outer half) into narrow black lines
upon a pale ocherous brown ground; outer third of wing brownish
ocherous dusted with black and with a few definable longitudinal
black streaks from end of cell; ocelloid patch a pair of closely
appressed obscure dull metallic vertical bars connecting above with
fainter metallic streaks from costa and inwardly margined by a
narrow white line; along termen below vein 6 four black dots on an
ocherous ground; at apex a short fine vertical black dash; cilia pale

_ NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 9

leaden fuscous with a whitish subbasal line. Hind wing pale smoky
fuscous, slightly paler toward base; silia sordid whitish with a dark
basal band.

Male genitalia of type figured.

Alar expanse —10-13 mm.

Type and paraty pes.—Cat. No. 28010, U.S.N.M.

Type locality —Dallas, Tex.

Food plant—Bumelia rigida (B. lanuginosa).

Described from male type and two male paratypes from the type
locality numbered, respectively, 626, 676, and 152, and the type
labeled in Fernald’s handwriting as follows: “ Dichrorampha bou-
- melliana Boll mss., two generations, June and Oct. in the rolled
leaves of Boumellia languinosa.” 'These specimens are from the Fer-
nald collection.

2. Genus DICHRORAMPHA Guenée
(Figs. 8, 28)

Dichrorampha GuENEE, Ann. Soe. Ent. France, ser. 2, vol. 3, 1845, p. 185.
Genotype.—Grapholitha plumbagana Treitschke (Europe).

Lipoptycha LEDERER, Wien. Entom. Monats., vol. 3, 1859, p. 370.
Genotype.—Phalaena plumbana Scopoli (Europe).

Hemimene Frernatp and AvutrHors (not Hiibner), Tortricidae and Their

Types, 1908, p. 8.
Genotype.—Phaleana Tortrix petiverella Linnaeus (Europe).

Thorax smooth.

Fore wing smooth; termen slightly concave below apex; 12 veins,
all separate; 7 to termen; 11 from cell before middle; 10 more or
less approximate to 9; upper internal vein of cell from between 10-
11; 3, 4, and 5 well separated at termen; 2 from cell at or near 44,
straight; male with or without costal fold.

Hind wing with normal pecten;.8 veins; 6 and 7 separate and
parallel; 3 and 4 connate or very short stalked; inner margine sim-
ple in male.

Hind tibia of male smooth scaled.

Male genitalia with harpe simple; outer surface unspined; cuculus
well defined, finely and evenly spined; neck incurvation usually pro-
nounced; neck smooth; succulus small, weakly spined. Tegumen a
narrow band. Uncus absent. Socii absent. Gnathos a simple,
weakly chitinized band. Aedoeagus curved; long or moderately
long; slender and scarcely tapering; cornuti a cluster of several weak
deciduous spines.

Abdomen of male simple.

Female genitalia with single thorn-like signum. Ductus bursae
moderately long; strongly chitinized toward genital opening and
sometimes with a slight chitinization near junction with bursa
copulatrix.

10 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

A genus about equally represented in North and South America
and Europe. The costal fold appears to be an acquired character—
not primative for the group. In North America it separates the
species into two groups‘ differing decidedly in the shape of the
harpe of the male genitalia; those with the fold have the cucullus
bent back on costa in the form of a broad hook, and these without
the fold have it crescentiform or semitrigonate. This difference
however does not hold for the European species (most of these with
the fold having the cucullus crescentiform). Busck, ignoring the
fold, suggests in his revision of the genus’ a possible division upon
the degree of obliqueness of the termen, the relative positions of
veins 9 and 10 and the relative width of areas above and below
the median vein of forewing, the protrusion of apex of hind wing
relative to tornus of forewing, and the color of the palpi. These
characters do not hold with any consistency, and, as far as they do,
separate into two groups forms with almost identical genitalia (like
kana and capitana), and group together others with radically dif-
ferent genitalia (like kana and banana). It is impossible to hold
Lipoptycha as a separate genus upon any character or set of
characters. The division I make here into two groups upon the
costal fold is apparently a natural one as far as our American species
are concerned, but no farther.

So far as known the larvae are root and shoot borers in Com-

positae.
KEY TO THE SPECIES OF DICHRORAMPHA

1. Thorax and fore wing striped in orange yellow and blackish fuscous.
(12) leopardana

MehOnax And: LOLe wil & OUMCL WSC sa ee a ee ee Pu
2. Fore wing with three or more black dots on termen near tornus________ 3
Mores wala oe withome Suchet ss. -o eis ee ee ee eee 9.
8. Outer half of fore wing heavily dusted with golden yellow___—_ (5) bittana.
Outer half of fore wing with faint yellow dusting or none____-------_- 4,
4. Termen of fore wing decidedly slanting, angle with costa less than 60°___ 5.
Termen of fore wing less slanting, angle with costa over 60°__-------- 6.
5. Fore wing with strong white dorsal spot_______________-___- (2) capitana.
Fore wing with very faint whitish dorsal spot or none__________ (1) kana.

6. Fore wing with no trace of pale dorsal spot; no costal fold in male.

(10) sedatana.

(11) dana.
Fore wing with pale dorsal spot at least indicated, normally strongly marked ;
MIALE WIth: COSTAL ONG ASE e ly egeh TIEN Se CDT 8 Se PE ey OE a hs
7. Alar expanse 10 mm. and less; dorsal spot of fore wing white, divided by a
Strone dark ime 5 he Be wie oe Ee et (6) ineanana.
Alar expanse 11 mm. and over; dorsal spot normally yellow, if white
unmarked ior wveny Lainthy Wined= 2220 ee ee ee ee 8.

7 Proc. Biol. Soe. Washington, vol. 19, 1906, pp. 1738-176.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 11

8. Distribution, Eastern United States and Canada____________ (4) simulana.
WISTEIPUTION Ee aAcine Coast Statesuce aap ee (3) britana,

OR LOLrem wine fawar COLT. 2 —— Sa War in ee (7) radicicolana.
Fore wing fuscous with very slight-ochreous dusting__________________ 10.

10. Vein 10 from cell over twice as far from 9 as 9 is from 8_____ (9) piperana.
Vein 10 from cell not over one and one-half times as far from 9 as 9 is

1B 005 Ape ot Meg Me ee ree ed be On a Re ee rn es (8) banana.

Group A. MALE WITH CosTAL FoLtp on Fore WING
1. DICHRORAMPHA KANA (Busck)

(Figs. 8, 273)

Lipoptycha kana Buscx, Proce. Biol. Soc. Washington, vol. 19, 1906, p. 182.

Lipoptycha planiloqua Meyrick, Ent. Mo. Mag., vol. 48, 1912, p. 36.

Hemimene kana BARNES and McDonnoueH, Check List Lepid. Bor. Amer.
no. 7266, 1917.

This and the two following species are very close. All are some-
what variable and show more or less of a dorsal patch on fore wing.
It is least obvious in some specimens of britana from California and
in kana. Specimens from the type localities can be distinguished
by the characters given in our key; but it is often difficult to place
those from new localities. D. kana differs from the others chiefly
in the more slanting termen of its fore wing.

' Male genitalia figured from paratype in National Collection from
the type locality (“ Dyar 23577”). Female genitalia as in capitana.

Distribution.—British Columbia, Washington, California (Tuo-
lumne Meadows).

Alar expanse.—17-20 mm.

Type.—In National Collection.

Type locality.— Kaslo, British Columbia.

2. DICHRORAMPHA CAPITANA (Busck)
(Figs. 111, 274)

Hemimene capitana Buscx, Proc. Biol. Soc. Washington, vol. 19, 1906, -p.
178.—BarneEs and McDunnovuen, Check List Lepid. Bor. Amer., no.
7262, 19177.
A Rocky Mountain species with a strong white dorsal patch on
fore wing.
Male genitalia figured from type; female from specimen in
National Collection from Silverton, Colo. (“July 16-23”).
Distribution.—Colorado, Utah.
Alar expanse.—i3-19 mm.
Type.—In National Collection.
Type locality—South Park, Colo.

12 BULLETIN 132, UNITED'’STATES NATIONAL MUSEUM

3. DICHRORAMPHA BRITANA (Busck)
(Fig. 275)

Hemimene britana Buscxk, Proc. Biol. Soc. Washington, vol. 19, 1906, p.
178.—BarNEs and McDuNNovucH, Check List Lepid. Bor. Amer., no.
7263, 1917.

Hemimene alpinana FERNALD (not Treitschke), in Dyar List N. Amer.
Lepid., no. 5290, 1903.

A Pacific coast species resembling kana and with the same dis-
tribution. Differs chiefly in having termen of fore wing less slant-
ing. Has been confused with the European alpinana which it re-
sembles superficially, but from which it differs strikingly in shape
of harpe of male genitalia.

Male genitalia figured from specimen in National Collection from
Goldstream, British Columbia (“2-VI-21, E. H. Blackmore No.
417”). Female genitalia as in capitana.

Distribution.—British Columbia, Oregon, California.

Alar expanse.—14-18 mm.

Type.—tiIn National Collection.

Type locality.—Kaslo, British Columbia.

4. DICHRORAMPHA SIMULANA (Clemens)

(Fig. 276)
Halonota simulana CLEMENS, Proc. Acad. Nat. Sci. Philadelphia, 1860,
p. 351.
Dichrorampha aurisignana ZELLER, Verh. Zool.-bot. Ges. Wien, vol. 25,
1875, p. 319.

Hemimene simulana FreRNALD, in Dyar List N. Amer. Lepid., no. 5289,
1903.—Busck, Proc. Biol. Soc. Washington, vol. 19, 1906, p. 179.—
BaRnES and McDuNNouGH, Check List Lepid. Bor. Amer., no. 7255,
1917.—ForBEs, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 389.

An Eastern species resembling britana, but with quite different
male genitalia; the chitin of the outer margin of the basal opening
of the harpe being produced into a short tongue or spur. In all the
other species of the costal-fold group (with the exception of bit-
tana) this outer margin is evenly rounded. The pattern and color
are somewhat variable; specimens from Mount Washington having
a much fainter dorsal patch and considerably darker shading on
outer half of forewing. They may possibly represent a distinct
race; but I do not feel justified in so designating them at this time.

Male genitalia figured from specimen in National Collection from
Ottawa, Canada (C. H. Young, “ 7—-VITI-1906 ”). Female genitalia
similar to those of bzttana.

Distribution.—Virginia, District of Columbia, Pennsylvania, New
Jersey, New Hampshire, Maine, Ontario, Quebec.

Alar expanse.—11-16 mm.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 13

Types—tIn Academy Natural Sciences, Philadelphia (simulana) ;
British Museum (aurisignana).
Type localities—Baltimore, Md. (simulana); Washington, D. C.
(aurisignana).
5. DICHRORAMPHA BITTANA (Busck)

(Figs. 104, 277)

Hemimene bittana Busck, Proc. Biol. Soc. Washington, vol. 19, 1906, p.
180.—Barnes and McDunnNouGH, Check List Lepid. Bor. Amer., no.
7256, 1917.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 389.
Close to and possibly a pale race of stmulana; differs chiefly in the
much heavier dusting of ochreous scales on outer half of fore wing.
Genitalia figured from specimens in National Collection from Pitts-
burgh, Pa. (Henry Engel, “ V—29—65,” male type) and Cincinnati,
Ohio (A. F. Braun, “ VI—24—04, female).
Distribution.— Pensylvania, Ohio, Wisconsin.
Alar expanse.—14—15 mm.
Type.—tIn National Collection.
Type locality —Pittsburgh, Pa.

6. DICHRORAMPHA INCANANA (Clemens)
(Figs. 108, 279)

Halonota incanana CLEMENS, Proc. Acad. Nat. Sci. Philadelphia, 1860,
p. 301.

Hemimene incanana FERNALD, in Dyar List N. Amer. Lepid., no. 5288,
1903.—Busck, Proce. Biol. Soc. Washington, vol. 19, 1906, p. 179.—
Barnes and McDuNnnoucH, Check List Lepid. Bor. Amer., no. 7254,
1917.—IorsBEs, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 389.

Hemimene nigromaculana Krarrott, Bull. Amer. Mus. Nat. Hist., vol. 23,
1907, p. 159.—Barnes and McDunnovueH, Check List Lepid. Bor. Amer.
no. 7264, 1917.

The type of Clemens’ species, a small male with distinct costal
fold (yellow label, No. 1437), is in Philadelphia. It agrees with
Kearfott’s nigromaculana except that the latter is somewhat larger.
Kearfott described from two specimens, but I am able to locate only
one (the female type in the American Museum). He had other
specimens under his name, but these are incorrectly determined.
They are Grapholitha prunivora Walsh. Aside from the Clemens
and Kearfott types I have seen only one other specimen of the true
incanana, a male in the National Collection from Falls Church, Va.
(Heinrich, “6—6—17”).

Male genitalia of this last figured; female genitalia figured from
type of nigromaculana.

Alar expanse—8—10 mm.

14 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Types—In Academy National Sciences (incanana); American
Museum (nigromaculana).

Type localities—Pennsylvania? (incanana); Black Mountains,
N. C. (nigromaculana).

Group B. MALE witTHouT CosTAL FoLp ON FORE WING
7. DICHRORAMPHA RADICICOLANA Walsingham
(Figs. 109, 282)

Dichrorampha radicicolana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus.,
vol. 4, 1879, p. 75.

Hemimene radicolana Frrnatp, in Dyar List N. Amer. Lepid., no. 5292,
1903.—Buscx, Proc. Biol. Soc. Washington, vol. 19, 1906, p. 180.

Hemimene radicicolana BARNES and McDuNNouGH, Check List Lepid. Bor.
Amer., no. 7257, 1917.

A large fawn-colored species. Walsingham gives the expanse of
his type (which according to Busck is a female) as 17 mm. All the
specimens in our American collections have an expanse of 20 mm. or
more.

Male genitalia figured from specimen in National Collection from
Deer Park Springs, Lake Tahoe, Calif. (“July 1-7”); female from
specimen in American Museum from Colfax, Placer County, Calif.

All specimens in National Collection, American Museum, and col-
lection Barnes from California.

Alar expanse.—17-22 mm.

Type.—tn British Museum.

Type locality —“*Camp Watson, on John Days River,” Oreg.

Food plant—‘Scophularia?” (larva feeding in roots according to
Walsingham).

8. DICHRORAMPHA BANANA (Busck)

(Fig. 278)

Lipoptycha banana Busck, Proc. Biol. Soc. Washington, vol. 19, 1906, p. 182.

Tipoptycha sordescens Meyrick, Ent. Mo. Mag., vol. 48, 1912, p. 36.

Hemimene banana Barnes and McDuNnnoucH, Check List Lepid. Bor.
Amer., no. 7267, 1917.

Close to radicicolana and with similar genitalia, but darker, and
grayish rather than fawn colored. In some specimens there isa faint
indication of a pale shade on mid dorsum; but in most it is not distin-
guishable. This dorsal pale marking is rather variable even in spe-
cies where it is normally present and sharply contrasted against the
ground color.

Male genitalia figured from type. (The harpes are somewhat bent
on the slide which has caused a foreshortening of the cuculli in the
photograph. They are shaped very much like those of radicicolana.)
I have seen no females.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 15

Specimens in National Collection, American Museum, and collec-
tion Barnes from Colorado.

Alar expanse.—20-24 mm.

Type.—tIn National Collection.

Type locality —South Park, Colo.

9. DICHRORAMPHA PIPERANA (Busck)

(Fig. 281)
Hemimene piperana Buscx, Proc. Biol. Soc. Washington, vol. 19, 1900
p. 177.—BarNES and McDunnovueH, Check List Lepid. Bor. Amer.,
no. 7261, 1917.

Superficially resembling both banana and sedatana. Differs from
the latter very distinctly in genitalia and from both in the venation
of fore wing (vein 10 is over twice as far from 9 at base as 9 is
from 8 in piperana, and much nearer 9 in the other two species).

Male genitalia figured from type.

In addition to the type, I have seen only one other specimen, a
male from Washington in the American Museum.

Alar expanse.—18 mm.

Type.—tIn National Collection.

Type locality—Pullman, Wash.

10. DICHRORAMPHA SEDATANA (Busck)

(Figs. 107, 283)

Hemimene sedatana Buscx, Proc. Biol. Soc. Washington, vol. 19, 1906,
p. 177.—Barnes and McDunnovueH, Check List Lepid. Bor. Amer.,
no. 7260, 1917.

Hemimene plumbana FERNALD (not Scopoli), in Dyar List N. Amer. Lepid.,
no. 5291, 1903.

Very close to the European plumbana and probably only a race
of that species. The two have been distinguished on pattern and
color; plumbana having a somewhat stronger irroration of yellow
scales on fore wing and sedatana a more abundant dusting of black
on the inner margin of the metallic lines. These characters are
somewhat variable. In genitalia I see no appreciable difference
between the two. For the present, however (at least until the Ameri-
can form is reared), they had best be kept separate.

Genitalia figured from specimens in National Collection from
Mount Tzouhalem, British Columbia (“24-V-21, Blackmore No.
424.” male), and Siskiyou Mountains, Calif. (“ 7-V 1-15-1872, Wal-
singham No. 91829,” female).

Distribution Colorado, California, British Columbia, Alaska.

Alar expanse.—12-15 mm.

Type.—tin National Collection.

Type locality.—South Park, Colo.

16 BULLETIN 132, UNITED STATES NATIONAL MUSEUM
11. DICHRORAMPHA DANA (Kearfott)

(Fig. 110)

Enarmonia dana Kearrort, Trans. Amer. Ent. Soc., vol. 33, 1907, p. 65.

Enarmonia aequorea Mryrick, Ent. Mo. Mag., vol. 48, 1912, p. 34.

Laspeyresia dana BARNES and McDuNnNnovueH, Check List Lepid. Bor. Amer.,
no. 72338, 1917.

Hemimene dana Forses, Memoir, 68, Cornell Univ. Agr. Sta., 1924, p.

388.

At most an eastern race of sedatana; but until the status of the
latter can be definitely settled Kearfott’s name may as well be kept
separate. The only difference I have been able to find between the
two (and this is of very doubtful significance) is in the female
genitalia; the bursa copulatrix is nearly twice as large in sedatana
as itisin dana. Superficially dana resembles Laspeyresia nigricana
Stephens, and Kearfott has frequently confused the two.

Female genitalia figured from specimen in American Museum
from New Brighton, Pa.

Distribution—Pemnsylvania, New Jersey, New Hampshire, Quebec.

Alar expanse —12-14 mm.

Type—tn American Museum.

Type locality—Montclair, N. J.

12. DICHRORAMPHA LEOPARDANA (Busck)

(Figs. 105, 280)

Hemimene leopardana Busck, Proe. Biol. Soc. Washington, vol. 19, 1906,
p. 181.—BarNeES and McDuNNovuGH, Check List Lepid. Bor. Amer., no.
7258, 1917.—Forses, Memoir 68, Cornell Uniy. Agr. Exp. Sta., 1924, p.
388.

A striking species, easily distinguished by the leopard-like striping
of forewing and thorax. It had been wrongly identified by Kear-
fott with incanana Clemens. The latter is about the same size, but:
has a costal fold and a white dorsal patch and differs otherwise in
structure and pattern.

Genitalia figured from reared specimens in National Collection
from Falls Church, Va. (Busck, August 1, 1913.)

Distribution.—North Carolina, Virginia, Maryland, Pennsylvania,
Ohio, Kansas, Ontario.

Alar expanse.—8-10 mm.

Type.—In National Collection.

Type locality—Hyattsville, Md.

Food plant.—Verbesina (Larva pupates within the rolled leaf).

. NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE tz

3. SATRONIA, new genus

(Fig. 23)

Genotype.—Satronia tantilla, new species (North America).

Thorax smooth.

Fore wing smooth; termen slightly concave; 11 veins in male
(12 in female), all separate; 7 absent (or united with 8) in male;
11 from cell at middle; 10 approximate to 9; upper internal vein
of cell from between 10-11; 3, 4 and 5 slightly approximate at ter-
men; 2 from cell before 24, straight; no costal fold in male.

Hind wing without pecten on lower median vein; 8 veins; 6 and
‘7 subparallel; 3 and 4 connate; inner margin in male simple.

Hind tibia of male smooth scaled.

Male genitalia with outer surface of harpe unspined; cucullus
elongate, narrow, finely and evenly spined; neck incurvation slight;
neck slender; sacculus weakly haired. Tegumen a narrow chiti-
nized band. Uncus absent. Socii absent. Gnathos a simple weakly
chitinized band. Aedoeagus short, stout, straight; cornuti a dense
cluster of short deciduous spines.

Abdomen of male with lateral hair tufts from sternite of eighth
segment.

_A higher development from #zcula. The male genitalia are simi-
lar in both except for the socii.

Monotypic and probably of tropical origin.

SATRONIA TANTILLA, new species
(Figs. 23, 285)

Palpus, face and head sordid whitish. Thorax and fore wing
grayish fuscous; from costa before middle to mid dorsum a pair of
moderately broad outwardly curved leaden metallic bands with a
faint dusting of white between; on outer half of costa four short,
faint white geminate dashes; ocelloid patch a single, rather wide
vertical metallic bar outwardly margined by 5 or 6 short, faint,
black dashes upon a fuscous ground faintly dusted with white; ter-
minal edge black; cilia leaden fuscous. Hind wing pale smoky
fuscous; cilia sordid whitish with dark basal band.

Male genitalia of type figured.

Alar expanse.—9.5 mm.

Type and paratype—Cat. No. 28011, U.S.N.M.

Type locality —Archer, Fla.

Described from male type (“No. 2630, May 482”); and one
female paratype (without abdomen) from Virginia shore opposite
District of Columbia (May 25, 1882). The type had been in the
National Collection unidentified for several years. It is somewhat

18 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

faded; but otherwise in good condition. The female is from the
Fernald Collection.

4. RICULA, new genus
(Figs. 4, 25, 106)

Genotype.—Lipoptycha maculana Fernald (North America).

Thorax smooth.

Fore wing smooth; termen convex; 12 veins all separate; 7 to
termen; 11 from cell slightly before middle; 10 approximate to 9;
upper internal vein of cell from between 10-11; 3, 4, and 5 remote
at termen; 2 from cell slightly beyond 24, straight; no costal fold
in male.

Hind wing with normal pecten; 8 veins; 6 and 7 somewhat ap-
proximate toward base (subparallel) ; 3 and 4 connate; inner margin
in male simple.

Hind tibia of male smooth scaled.

Male genitalia with outer surface of harpe unspined; cucullus
elongate, narrow, finely and evenly spined; neck incurvation slight;
neck slender; sacculus weakly haired. Tegumen a narrow chitinized
band. Uncus absent. Socii developed, long, fingerlike, flexible.
Gnathos a simple weakly chitinized band. Aedoeagus short, stout,
straight; cornuti a dense cluster of long deciduous spines.

Abdomen of male with lateral hair tufts from sternite of eighth
segment.

Female genitalia with single signum developed as a thornlike
spine. Ductus bursae very short and broad. Bursa copulatrix nar-

row; scobinate at neck.
A monotypic genus presumably of tropical origin. Related to
Talponia, from which it differs chiefly in the more approximate
condition of veins 6 and 7 of hind wing, the convexity of the termen
of fore wing, and the absence of one signum from the bursa of the
female.
RICULA MACULANA (Fernald)

(Figs. 4, 25, 106, 284)

Lipoptycha maculana FERNALD, Journ. New York. Ent. Soc., vol. 9, 1901, p.
51; in Dyar List N. Amer. Lepid., no. 5293, 1903.

Laspeyresia maculana Buscx, Proc. Biol. Soc. Washington, vol. 19, 1906, p.
173.—BaRNbs and McDuNNovuGH, Check List Lepid. Bor. Amer., no.
G211, 1917.

A very small dark fuscous species with several long narrow ob-
lique orange costal streaks on fore wing, a large triangular white
spot on mid dorsum and an ocelloid patch consisting of a single ver-
tical metallic bar outwardly margined by 3 or 4 black-centered
orange spots.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 19

Genitalia figured from paratypes in National Collection from the
type locality (H. G. Dyar “ 833 Fla.,” issued Feb. 25-26, 1900).

Represented in the collections, as far as I know, only by the type
series in the National Collection.

Alar expanse.—7.5 mm.

Type—tIn National Collection.

Type locality —F lorida.

Food plant.—Schoepfia arborescens.

5. TALPONIA, new genus
(Wigs. 114, 286)

- Genotype.—Hemimene plummeriana Busck (North America).

Characters as in /?cula except:

Fore wing with termen incurved below apex; 2 from cell at 2%.

Hind wing with 6 and 7 separate and parallel.

Female genitalia with two thorn-like signa. Ductus bursae long,
slender. Bursa copulatrix large; neck smooth.

Monotypic. Ailied to Ricula, Ethelgoda, and the tropical Balbis
Walsingham. In wing shape, general habitus most like Lthelgoda.
In genitalia (male and female) closest to Balbis. The latter, how-
ever, is quite distinct, having a costal hair pencile on hind wing of
male, strong pecten on underside of vein 1° on fore wing and dif-
ferent venation: 1° in fore wing absent and 8 weak and closely
approximate to cell in hind wing, partially anastomosing with 7
to beyond cell.

TALPONIA PLUMMERIANA (Busck)

(Figs. 114, 286)

Hemimene plummeriana Buscxk, Proc. Biol. Soc. Washington, vol. 19,
1906, p. 181; Proce. Ent. Soe. Washington, vol. 11, 1909, p. 99.—BarNES
and McDunnoucH, Check List Lepid. Bor. Amer., no. 7259, 1917.—
Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 388.
Thorax and basal half of fore wing powdery gray; outer half
of fore wing red brown to the naked eye, under magnification brown-
ish orange dusted and streaked with black and heavily marked with
purplish metallic bands. Hind wing brown with a narrow yellow
border along termen. Larva feeds in flowers of pawpaw.
Genitalia figured from reared specimens in the National Collec-
tion from Plummer Island (male) and Hyattsville, Md. (female).
Distribution —Maryland, Ohio. There is also a specimen in
the National Collection from Guerrero, Mexico.
Alar expanse.—9-10 mm.
Type.—tIn National Collection.
Type locality —Plummer Island, Md.
Food plant.—A siminia triloba.

20 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

6. Genus HEMIMENE Hiibner
(Figs. 6, 27)

Hemimene HUsner, Verz. Schmet., 1826, p. 378. (=Phthorobdlastis
Lederer).
Genotype.—Pyralis populana Fabricius (Europe).
Pammene HtUsner, Verz. Schmet., 1826, p. 378.
Genoypte.—Tortriz trauniana Schiffermiiller (Europe).
Pseudotomia STEPHENS, System, Cat Brit. Ins., 1829, p. 175.
Genotype.—Tortriz argyrana Hiibner (Europe).
Hemerosia STEPHENS, List, Brit. Animals, pt. 10, Lepid., 1852, p. 60.
(=Palla Billberg —Pyrodes Guenée, preoccupied).
Genotype.—Phalaena Tinea rhediella Clerck (Europe).
Strophedra HerricH-ScHAEFER, Schmet. Eur., vol. 5, 1855, p. 94.
Genotype.—Grapholitha flerana Zeller (Europe).

Thorax smooth.

Fore wing smooth; termen straight or slightly concave below
apex (at vein 6); 12 veins, all separate; 7 to termen; 11 from cell
slightly before middle; 10 more or less approximate to 9; upper
internal vein of cell from between 10 and 11; 3, 4, and 5 well
separated at termen (often parallel from beyond cell, rarely 3 and
4 approximate at termen) ; 2 from cell at or before 24, straight; no
costal fold in male.

Hind wing with normal pecten; 8 veins; in male 7 fusing with
8 beyond cell and 6 well separated; in female 8 free and 6 and 7
approximate toward base; 3 and 4 connate or stalked; inner margin
in male simple.

Hind tibia of male smooth scaled.

Male genitalia with outer surface of harpe unspined; cucullus
densely and evenly spined; neck incurvation appreciable, but slight;
neck well spined; sacculus finely haired, sometimes with two or
three strong spines. Uncus absent. Socii absent. Gnathos weakly
chitinized, a simple band, sometimes expanded beneath into an appre-
ciable subanal plate. Aedoeagus moderately long; rather stout;
straight or curved; tapering and often bottle necked; cornuti a
cluster of short stout spines, a few fixed, the rest deciduous.

Abdomen of male simple or with modified dorsal hair tufts (under
superficial scaling) on segments 6, 6 and 7, or 6, 7 and 8; rarely (in
flexana Zeller) with a pair of lateral tufts from sternite of eighth
segment.

Female genitalia with two thorn-like signa. Ductus bursae very
short, unchitinized except near genital pees Bursa copulatrix
large; neck smooth.

The above description is drawn to include the European species
with vein 7 running into 8 in the male hind wing. These vary
greatly in male abdominal characters: populana (type of Hemi-

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 2]

mene), spiniana Duponchel and fimbriana Haworth have the abdo-
men simple; trauniana (type of Pammene), oxycedrana Milliére,
splendidulana Guenée and amygdalana Duponchel have a tuft on
sixth segment only; argyrana (type of Pseudotomia), juliana Curtis,
gallicolona Zeller, christophana Moeschler, regiana Zeller and our
four North American species have tufts on abdominal segments
6 and 7; rhediella Clerck (type of Hemerosia) has them on 6,7 and
8; flexana Zeller (type of Strophedra) has lateral tufts on the eighth
segment like these of Grapholitha, otherwise the abdomen is smooth.
This last has other differences also; the fore wing has termen decid-
edly slanting, costa sharply bent at apex and apex somewhat pointed
and produced and veins 3, 4, and 5 widely separated at termen.
Very likely when larvae and pupae are studied it may be found ad-
visable to divide the genus. In that case all the above synonyms will
take their places as valid genera and the American species be referred
to Hemerosia. At present nothing would be gained by splitting.

It is unfortunate that the name Hemimene, which has hitherto
been used for the moths now under Pichrorampha, must be sub-
stituted for Pammene; but the fixing of populana as type by
Walsingham and Durrant (1901) compels it.

In Europe the larvae are leaf tiers, feeders in galls (inquilines) or
under bark of forest trees. The larve of none of our American
species are known.

Derived from and a higher development of Laspeyresia.

KEY TO THE SPECIES OF HEMIMENB

1. Fore wing with well-marked white patch on dorsum____________-______ 2
Ore wink without’ SuGh2. 2 = so. ey se oy sees bere RE ee (4) paula.
2 aWhite dorsal patch continuous! to base Of (wing + i.e 2-2 es 2
White dorsal patch limited toa spot on mid-dorsum__________ (3) signifera.

3. White costal geminations on outer half of fore wing strong and partially
fused ; metallic markings dull, leaden____________ (1) ocliferia.
White costal geminations on outer half of fore wing faint and well spaced:
meiAlichmarkines: Drieht . bluish. = ee (2) felicitana.

1. HEMIMENE OCLIFERIA, new species
(Fig. 291)

Antenna ocherous. Palpus white. Face white. Head grayish
fuscous. Thorax dark grayish fuscous; posterior margin and tip
of tegula white. Fore wing dark grayish fuscous (almost black)
with white costal strigulations and an elongate irregular white dor-
sal patch extending from base to beyond middle; white dorsal patch
slightly narrower at base than at middle of dorsum; costal strigu-
lation beyond mid costa partially fused forming a whitish shade on
apical half of costa; ocelloid patch obscure, four indistinct black
streaks between two faint vertical metallic bars on a blackish gray

ae BULLETIN 1382, UNITED STATES NATIONAL MUSEUM

ground; edge of termen black, broken below apex, at middle and at
tornus by white spots; cilia leaden fuscous; white dorsal patch and
costal markings repeated on under side of wing. Hind wing dark
smoky brown; cilia shining white with dark basal band; on under
side basal half of wing whitish. Legs white; outer sides dusted
with dark grayish fuscous.

Male genitalia of type figured.

Alar expanse.—12-14 mm.

Type.—tin collection Barnes.

Paratypes.—Cat. No. 28012, U.S.N.M. Also in American Museum
and collection Barnes.

Type locality —Pyramid Lake, Nev.

Described from male type and five male paratypes from type
locality.

Close to felicittana and with similar genitalia. Easily distin-
guished by the characters given in the key.

2. HEMIMENE FELICITANA (Heinrich)
(Figs. 116, 292)

Pammene felicitana HrEtnricu, Proe. Ent. Soe. Washington, vol. 25, 1923,
p. 120.—Forsers, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 389.

An eastern species similar to ocliferia but with much brighter
metallic markings on forewings.

Male genitalia figured from type in American Musetim; female
from paratype in National Collection from Oak Station, Pa. (Mar-
loff, “ June 19-07.”)

Distribution.—Pennsylvania, Quebec.

Alar expanse.—13-14 mm.

Type.—in American Museum.

Type locality —Montreal, Quebec.

3. HEMIMENE SIGNIFERA, new species
(Fig. 290)

Antenna fuscous. Palpus and face fuscous ocherous. Head fus-
cous ocherous, darker at sides. Thorax semilustrous leaden fuscous.
Forewing blackish (or dark grayish) fuscous; on mid dorsum a
conspicuous nearly square white spot reaching up to cell; costa with
a faint antemedian and four postmedian pair of obscure whitish
ocherous germinate marks; ocelloid patch obscure, consisting of four
or five black streaks interspaced with ocherous scaling and between
two dull vertical metallic bars; edge of termen black; cilia leaden
fuscous. Hind wing smoky fuscous; cilia pale fuscous with a dark
basal band. Underside of fore and hind wings concolorous, pale
smoky fuscous. Legs pale ocherous fuscous.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 23

Male genitalia of type figured.

Alar expanse.—12 mm.

Type.—tin Canadian National Collection.

Type locality —Lake of Bays, Ontario.

Described from unique male type (McDunnough, “ 1—-VII-1920”).

4. HEMIMENE PAULA, new species
(Fig. 289)

Antenna blackish. Palpus, face, head, and thorax blackish with
the extreme ends of the scales white. Forewing dark grayish fus-
cous; from costa before middle a pair of narrow dull metallic lines
curving out to a little beyond middle of cell and thence back to mid
dorsum forming a complete fascia; bordering this, outwardly, a
black dusted shading of the ground color, appearing to the naked
eye as a blackish brown fascia; outer half of costa with four pair
of white geminate marks from the first and second of which extend
faint metallic lines to the vertical bars of ocelloid patch; ocelloid
patch obscure, consisting of a couple of indistinct, irregular black
dashes upon an ocherous ground between two dull metallic vertical
bars; edge of termen black; cilia leaden fuscous. Hind wing smoky
fuscous; cilia whitish with a dark basal band; underside of wing
pale (smoky whitish) with a few dark spots at apex. Legs dark
grayish fuscous; tarsi faintly annulated with white.

Male genitalia of type figured.

Alar expanse —10 mm.

Type.—In Canadian National Collection.

Type locality —Aweme, Manitoba.

Described from unique male (N. Criddle, “ 11-V-1921”’).

7. ETHELGODA, new genus
(Figs. 26, 122)

Genotype.—Phthoroblastis tevanana Walsingham (North Amer-
ica).

Thorax smooth.

Forewing smooth; termen incurved below apex; 12 veins, all
separate; 7 to termen; 11 from cell at middle; 10 approximate to 9;
upper internal vein of cell from between 10-11; 3, 4, and 5 parallel
from slightly beyond cell; 2 from cell near 24, straight; no costal
fold in male.

Hind wing with normal pecten; 8 veins; 6 and 7 subparallel; 3
and 4 connate; inner margin in male simple.

Hind tibia of male smooth scaled.

Male genitalia with outer surface of harpe unspined; cucullus
small, rounded, heavily spined; neck incurvation deep; neck slender,

94 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

smooth; sacculus weakly haired. Uncus absent. Socii absent.
Gnathos a simple weakly chitinized band. Aedoeagus long, stout,
tapering, slightly curved; cornuti a cluster of very small slender
deciduous spines.

Abdomen of male with lateral hair tufts from sternite of eighth
segment.

Female genitalia with two thorn-like signa. Ductus bursac mod-
erately long; strongly chitinized toward genital opening and with
a chitinized patch near junction with bursa copulatrix.

Monotypic and probably of tropical origin. The genus has hardly
a single character to define it, yet on the sum of its characters it fits
in none of the other genera. On wing pattern and general habitus it
should go with Zalponia (T. plummeriane and FE’. texanana differ
superficially only in color). On male genitalia and abdominal
characters it could go in Grapholitha. Its female genitalia (except
for the two signa) are those of Dichrorampha. Its hind wing vena-
tion is that of Ricula. A separate designation is therefore necessary
unless we are to confuse the definition of our other genera.

Derived from Grapholitha.

ETHELGODA TEXANANA (Walsingham)
(Figs. 26, 122, 287)

Phthoroblastis texranana WAaALsiIncHAmM, Illus. Lepid. Heter. Brit. Mus.,
vol. 4, 1879, p. 70.

Pammene teranana FERNALD, In Dyar List N. Amer. Lepid. no. 5294, 1903.—
BARNES and McDunnoueH, Check List Lepid. Bor. Amer. no. 7268,
1917.

In pattern and general habitus similar to Z'alponia plummeriana
Busck, but with different genitalia and without the distinctly reddish
shade on outer half of fore wing. Thorax and basal half of fore
wing powder gray; outer half of wing brownish; from mid costa to
dorsum near tornus a straight narrow dull metallic band; from costa
just beyond, a similar narrow band curving out to a point between veins
6 and 7 a short distance from termen thence inward to base of vein 3
and thence down to dorsum; termen finely edged with ocherous,
bordered near tornus by a dull metallic band; a subterminal row of
small black dots or abbreviated dashes. Hind wing brown.

Genitalia figured from specimens in National Collection from
Dallas, Tex. (“ 595,” male) and Biscayne Bay, Fla. (female).

The only specimens I have seen are three males from Texas and
the female from Florida from the Fernald collection in the National
Museum; one of these had evidently been referred to Walsingham
and determined by him.

Alar expanse.—10-13 mm.

Type—tn British Museum.

Type locality —Texas.

“NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 95

8. Genus SEREDA Heinrich
(Figs. 29, 118)

Sereda Hetnricu, Proc. Ent. Soc. Washington, vol. 25, 1923, p. 121.
Genotype.—Halonota lautana Clemens (North America).

Thorax smooth.

Fore wing smooth; termen covex; 12 veins, all separate; 7 to ter-
men; 11 from cell before middle; 10 approximate to 9; upper inter-
nal vein of cell from between 10 and 11; 3, 4 and 5 remote at termen;
2 from cell just before 14; straight; no costal fold in male.

Hind wing without pecten on lower median vein; 8 veins; 6 and 7
approximate toward base; 3 and 4 connate; inner margin simple in

male.

Male genitalia with outer surface of harpe unspined; cucullus trig-
onate; neck very slender; neck incurvation deep; sacculus enlarged,
weakly spined. Tegumen a narrow chitinous band. Uncus absent.
Socii absent. Gnathos a simple, weakly chitinized band. Aedoeagus
long, stout, slightly bent, scarcely tapering; cornuti 3 or 4 short,
stout fixed and a cluster of deciduous spines. ~~
Abdomen of male with a pair of long hair tufts from lateral ex-
tremities of sternite of eighth abdominal segment.

Female genitalia with two signa, developed as thornlike spines.
Ductus bursae short, chitinized and bent.

A monotypic genus derived from Grapholitha. The absence of
pecten is rare for the family, occuring elsewhere, as far as I know,
only in Satronia and Goditha.

SEREDA LAUTANA (Clemens)
(Figs. 29, 118, 288)

Halonata lautana CLEMENS, Proc. Ent. Soe. Philadelphia, vol. 5, 1865, p.
139, (as tautana through typographical error).

Grapholitha perfluana ZELLER, Verh. Zool.—bot. Ges. Wien, vol. 25, 1875,
p. 299.

Enarmonia lautana FERNALD, in Dyar List N. Amer. Lepid., no. 5279, 1903.—
Kerarrort, Can. Ent., vol. 37, 1905, p. 254.

Laspeyresia lautana Krarrort, Ins., New Jersey, 1910, p. 546.—BaArNEsS and
McDunnoueHu, Check List Lepid. Bor. Amer., no. 7231, 1917.—ForBgs,
Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 391.

Sereda lautana HeEtnricu, Proc. Ent. Soc. Washington, vol. 25, 1928, p. 121.

Easily identified by structural characters and the strong black dots
along termen on upper and under side of fore wing. In one specimen
in the National Collection veins 7 and 8 of fore wing are stalked.
This, however, is very unusual. In all other specimens I have seen
they are distinctly separate.

Genitalia figured from specimens in National Collection from Ox-
bow, Saskatchewan (Knab, “15-V—07,” male), and New. Brighton,
Pa. (Merrick, “4-14-03,” female).

26 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Distribution—Massachusetts, New Hampshire, New York, New
Jersey, Pennsylvania, Texas, Manitoba, peewee

Alar expanse.—10-12 mm.

Types.—In Academy National Sciences (lautana) ; Museum Com-
parative Zoology (perfluana).

Type localities—Virginia (lautana) ; Texas (perfluana).

9. Genus GRAPHOLITHA Treitschke
(Fig. 7)
Grapholitha TreIrscHKE, Schmet. Eur., vol. 7, 1829, p. 282 (=Stigmonota
Guenée).

Genotype.—Pyralis dorsana Fabricius (Europe).
Euspila STEPHENS, Syst. Cat. Brit. Ins., 1829, p. 103.

Genotype—Tinea compositella Fabricius (Burope).
Ephippiphora DuproncHEL, Hist. Nat. Lépid. France, vol. 9, 1834, pp. 22,

304.
Genotype.—-Phalaena Tinca jungiella Clerck (Hurope).
Opadia GuENEE, Eur. Microlepid, Index Method., 1845, p. 48.
Genotype.—Grapholitha funebrana Treitschke (Europe).

Thorax smooth.-

Fore wing smooth; termen straight or convex, or with a very
slight concavity at vein 6; 12 veins, all separate (in aberrant speci-
mens 7 and 8 are occasionally short stalked, but this character does
not seem to hold even for species); 7 to termen; 11 from cell at or
a trifle before middle; 9 and 10 well separated; upper internal
vein of cell from between 10-11; 3, 4 and 5 well separted at termen;
2 from cell before 24, straight or very slightly bent; male without
costal fold.

Hind wing with normal pecten; 8 veins; 6 and 7 approximate
towards base; 3 and 4 connate or very short stalked; inner margin
simple in male.

Hind tibia of male smooth scaled.

Male genitalia with harpe simple; outer surface unspined; cucul-
lus well defined, finely and evenly spined; neck incurvation usually
pronounced; neck smooth or very weakly spined; sacculus simple,
weakly spined. Tegumen a rather narrow band. Uncus absent.
Socii absent. Gnathos a simple, weakly chitinized band. Aedoea-
gus straight or very slightly curved; rather long; stout or mod-
erately so and evenly tapering; cornuti a cluster of three or more
spines, one or two fixed, the rest deciduous, rarely all fixed.

Sternite of eighth abdominal segment (in male) developed as a
narrow, chitinized band at the lateral extremities of which are a
pair of long heavy hair tufts (fig. 7.).

Female genitalia with two signa developed as thorn like spines;
signa rarely absent. Ductus bursae short; strongly chitinized and
often scobinate near genital opening.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE BT

Derived from Laspeyresia. Is well represented in Europe and
North America. On the ocelloid patch it divides into what appear
to be two natural groups. I do not think however that these deserve
generic rank, since there are no characters other than the ocelloid
patch on which to separate them.

10.

i is

12.

18.

14.

KEY TO THE SPECIES GRAPHOLITHA

Use patch of fore wing a pair of vertical me tallie bars, well separated

Ocelloid patch a single vertical hae or ak Finders appressed vertical
metallic bars scarcely separated by a vertical line of darker sealing__ 13.

. Fore wing with one or more distinct white geminate marks on mid dor-

SUM 23232 eee ca acs pours ea ee ee ee 3.
Fore wing with pale dorsal markings obscure or absent________-__---- 6.

. Fore wing with a faint, scattered: dusting of white scales at base, on disk,
enc salom er Cermene = ase = Aa ae te (10) imitativa.
Moreswine without such white dusting 225 <2 28 ee ee 4,

. Ocelloid pateh containing six unbroken longitudinal black streaks; white
dorsalugeminations:broad_ifutel Gou W_ bath gow eres f-- 53h (8) fana.
Black longitudinal markings of ocelloid patch irregular, broken, and less
than six; white dorsal geminations narrow__~__~--_---___-________ 5s

. Hind wing distinctly paler than ground cclor of fore wing; post median

area of fore wing darker than basal area________________ (11) lunatana.
Hind wing dark, nearly concolorous with ground color of fore wing; post

- median area of fore wing no darker than basal area.
(9) conversana (part).

. Hind wings hyaline white toward base; broadly margined with dark

fuscews at. Apex. Bie. wes ee het pnt a et (12) eclipsana.
Hind wings sometimes pale toward base but never hyaline white_____~ lee

. Fore wing with white dusting on fore wing forming an obscure, angulate
Dale pInCUTAN! iNCla t=.) ane anes a ON Pe eee ee ae (7) vitrana.
Fore wing without such a pale median fascia___-__--_-_--_---_-_-_-_ 8.

. Fore wing longitudinally streaked on disk and well dusted toward apex
rit Re RUSiVOCHCTOUNS, SCION 5 Fe ee eee 55 tt (4) prunivora.
Fore wing without such ocherous sealing______-_-_-__-___~=----_-_+._ 9.

. Fore wing with six or more distinct white geminations on costa_______ 10.
Fore wing with costal geminations obsolete or distinguishable only as very
Paint, minute! $treaksss sus SI HB iO Gissiiss Die Syke use
Costal-geminations evenly spaced____z2-- +--_---__+_+-=n- (5) angleseana.

Apical pair of costal geminations well separated from the preceding four.
(6) caeruleana.
(9) conversana (part).
Fore wing fully half as broad as long; costa appreciable arched toward

apexciiGis 54. Sis it Bi O21 Sty sas de 2 UO (1) molesta.
Fore wing somewhat less than half as broad as long; costa straight toward

RNID A ee errs ee Re eo ey eee i Oe an eg da ges ge ere pm eee 12.
Male with heavy black scaling on hind wing_----_-~----_-_-__ (3) packardi.
Male without sex, scaling on hing wing- ~~ -- (2) libertina.
Four geminate white streaks on mid dorsum of fore wing_-________-_- 14.
Two geminate white streaks on mid dorsum of fore wing______-_-__--- 15.
Hntire paipus white=-——- so... a ee eet st aren (17) tristrigana.
Palpus strongly dusted with fuscous toward apex _____-~~~ (15) lana (part).

54346—26—_3

28 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

15. Thorax and base of fore wing pale drab brown, paler than ground color of

CEG ee 61 a ei, Se eee ene ae 2 | oe ee (13) interstinctana.
Thorax and base of fore wing fuscous brown, concolorous with ground
MOLT TCE POSIT AV ATES os ee ee ee) Le 16.

16. Hind wing cilia uniformly clear white except for contrasting dark basal
Basen Ghee fs ee ee = I eee (14) edwardsiana.
Hind wing cilia pale fuscous: sometimes shaded with whitish but never
qistinetly white = 22 elOu Ske ere aE ee ee tf:

17. Some faint whitish dusting on terminal area of fore wing; dorsal geminate
markings extending above middle of wing________________ (16) dyarana.

No white dusting on terminal area; dorsal geminations not extending above
WTIGNC OE WINS oo 8 ee ee (15) lana (part).

Group A.—OcELLOID PatTcH CONSISTING OF Two VERTICAL Bars INCLOSING
BriackK Dots oR LONGITUDINAL STREAKS

1. GRAPHOLITHA MOLESTA (Busck)
(Figs. 129, 305)

Laspeyresia molesta Busck, Journ. Agr. Res. U. S. Dept. Agr., vol. 7, 1916,
p. 373.—QUAINTANCE and Woop, Journ. Agr. Res. U. S. Dept. Agr., vol.
7, 1916, pp. 373-377.—CHUKICHI HAarRUKAWA and NoBUuMASA YAGI,
Berichte des Ohara Instit., Juraschiki, Japan, vol. 1, pt. 2, 1917, pp.
151-170; vol. 2, pt. 3, 1923, pp. 235-258.—Yasusur1 Nawa, Insect
World, Gifu, vol. 21, no. 7, pl. 7—-Woop and SeExLkrece, Journ. Agr.
Res. U. S. Dept. Agr., vol. 18, 1918, pp. 59-72—Garuan, Bull. no. 223,
Maryland Agr. Exp. Station, 1918, pp. 103-126—Pao.r, Agr. Col.
Florence, vol. 15, 1921, pp. 572-576.—MeEyricx, Entomologist, vol. 55,
1922, p. 255.—TruveLot, Bull. Soc. Ent. France, 1922, pp. 220-223.—
Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1994, p. 394.

The well-known oriental peach moth of economic literature. It is
an important enemy of peach, apple, and a number of other fruit
trees, the larva boring in the growing shoots and also tunneling the
fruits.

A moderately large, dark, slate-colored species with very obscure
markings. Most closely resembling packardi and libertina but
easily separable from both. Hind wing with veins 3 and 4 connate.

Male and female genitalia figured from reared specimens in Na-
tional Collection from Arlington, Va. (on peach, Kk. R. Selkregg,
“5-6-19”).

Distribution.—District of Columbia, Maryland, Virginia, Penn-
sylvania, Indiana, North Carolina, Tennessee, Arkansas, Mississippi,
Georgia, Alabama, Florida. We also have in the National Collec-
tion reared specimens from Japan and Australia.

These last are part of a series determined by August Busck for
W. W. Frogatt, and which the latter had previously figured and
described * as an unidentified enemy of the peach in Australia under
the popular name peach-tip moth. This is the first reference to
the species in literature.

§ Agr. Gazette, New South Wales, May, 1914, p. 413.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 99

There are also authentic records from France and Italy.
Alar expanse—11-13 mm.
Type.—tn National Collection.
Type locality.—Arlington, Va.
Food plants—Apple, peach, quince, plum, pear, nectarine, apricot,
and cherry.
2. GRAPHOLITHA LIBERTINA, new species
(Fig. 294)

A western species of the same size and color as packardi Zeller
and with similar maculation; but without the characteristic hind
wing sex scaling of the latter and with different genitalia. Palpus,
head, thorax, and fore wing an almost unicolorous fuscous brown;
fore wing very faintly dusted with whitish scales and with an
obscure narrow dark fascia from mid costa to outer fifth of dor-
sum; ocelloid patch obscure, containing four or five faint black
dashes. Hind wing uniformly brownish, but little paler than fore
wing; cilia concolorous, with dark basal band. Male genitalia with
costa of harpe more decidedly curved than that of packardi.

Male genitalia of type figured.

Alar expanse.—9.5 mm.

Type.—In American Museum.

Paratype.—Cat. No. 28013, U.S.N.M.

Type locality —Wellington, British Columbia.

Described from male type and paratype from the type locality
(G. W. Taylor, collector). These two specimens had been included
by Kearfott with a series of the eastern packardi under the latter
name.

3. GRAPHOLITHA PACKARD! Zeller
(Figs. 132, 295)

Grapholitha packardi ZeLi_eR, Verh. Zool.-bot. Ges. Wien, vol. 25, 18785.
p. 300.

Steganoptycha pyricolana MURTFELDT, Bull. 23, U. S. Dept. Agr., 1891,
p. 52.—SaANDERSON, 12th Ann. Rep. Deleware Agr. Exp. Sta., 1901,
p. 195; Can. Ent., vol. 35, p. 159.

Epinotia pyricolana FERNALD, In Dyar List N. Amer. Lepid., no. 5234,
1908.

Enarmonia packardi FERNALD, in Dyar List N. Amer. Lepid., no. 5282, 1903.

Enarmonia pyricolana Kearrott, Ins. New Jersey, 1910, p. 544—BARNES
and McDunnovucH, Check List Lepid. Bor. Amer., no. 7158, 1917.—
GARMAN, Bull. 223, Maryland Agr. Exp. Sta., 1918, pp. 105, 106, 108,
and 109.

Laspeyresia packardi BARNES and McDunnovueH, Check List Lepid. Bor.
Amer., no. 7238, 1917—(not Forbes, Memoir 68, Cornell Univ. Agr.
Exp. Sta., 1924, p. 392.)

Laspeyresia pyricolana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 395.

30 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

A species of some economic importance as an enemy of apple.
The larva is apt to be confused with that of molesta which it much
resembles. It usually attacks the growing twigs, seldom the fruit.
It has also been reared from rose tips and there are a couple of
doubtful records from peach.

The adult is smaller than that of molesta and has a faint but
distinguishable dark median fascia on fore wing. Its most striking
character, however, is a strong patch of blackish sex scaling upon
the upper surface of the hind wing and a similar patch on the under
surface of the fore wing of the male. This character as far as I
know is shared by no other North American species of Grapholitha
or Lespeyresia. Zeller’s type, unfortunately, is a female; but I do
not think there can be any doubt of its synonymy with Miss Murt-
feld’s species.

Hind wing with veins 3 and 4 connate.

Genitalia figured from specimens in National Collection frora
Plummer Island, Md. (Busck, July, 1903, male) and Missouri
(“ 368—M,” “ 5-2-07 ” reared from “rose tips,” female).

Distribution.—Texas, Mississippi, Missouri, Arkansas, Illinois,
Michigan, Maryland, West Virginia, Virginia, New Jersey, Dela-
ware, Massachusetts, New Hampshire.

Alar expanse.—8-10.5 mm.

Types.—Museum Comparative Zoology (packardi); lost (pyri-
colana).

Type localities—Texas (packardi) ; Missouri (pyricolana).

Food plants—Apple, rose, (peach?), Crataegus.

4. GRAPHOLITHA PRUNIVORA (Walsh)
(Figs. 130, 296)

Semasia prunivora WAusH, First Rep. Ins. Illinois, 1868, pp. 105-110.

Enarmonia prunivora FERNALD, in Dyar List N. Amer. Lepid., no. 5269,
1903. QuAINTANCE, U. S. Dept. Agr., Bur. Ent. Bull. no. 68, pt. 5.
1908, pp. 49-60.

Laspeyresia prunivora BARNES and McDuNNoucH, Check List Lepid. Bor.
Amer. no. 7208, 1917.—GarMAN, Bull. 223, Maryland Agr. Exp. Sta.,
1918, pp. 105-107.—WeELLHousE, Memo. 56, Cornell Agr. Exp. Sta., 1922,
p. 1078.—Forses, Memo. 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 392.

Another rather common species, known in economic literature as
the lesser apple worm.

Genitalia figured from specimens in National Collection, reared
from crab apple but without locality labels (“Aug. 19-82,” male and
“Quaintance No. 875,” female).

Hind wing with veins 3 and 4 connate.

Distribution.—District of Columbia, Maryland, Virginia, West
Virginia, Pennsylvania, Indiana, Illinois, Missouri, Arkansas,
Washington, Maine, Ontario.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 8]

Alar expanse.—8-10 mm.

Type.—tIn National Collection.

Type locality.—Ilinois.

Food plants.——¥ruits of apple, plum, peach, cherry, Crataegus,
Amelanchier; also in “black-knot” fungus and in aphid galls on
oak and elm.

5. GRAPHOLITHA ANGLESEANA (Kearfott)

(Figs. 127, 304)

Enarmonia angleseana Kearrott, Trans. Amer. Ent. Soc., vol. 33, 1907,
. 64.
creel angleseana BARNES and McDuNnnNovueH, Check List Lepid. Bor.
Amer., no. 7214, 1917.—Forxbes, Memo. 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 393.
Hind wing with veins 3 and 4 connate.
Male and female genitalia figured from paratypes in National Col-
lection from type locality.
Distribution—New Jersey, Massachusetts.
Alar expanse.—10-11 mm.
Type—tIn American Museum.
Type locality Anglesea, N. J.
Food plant.—‘ Strawberry seeds” (Fernald rearing note).

6. GRAPHOLITHA CAERULEANA Walsingham
(Figs. 125, 299)

Grapholitha caeruleana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol. 4,
1879, p. 66.

Enarmonia caeruleana FERNALD, in Dyar List N. Amer. Lepid., no. 5277,
1908.

Enarmonia zana Krarrott, Trans. Amer. Ent. Soc., vol. 33, 1907, p. 61.

Enarmonia vana Kearrort, Trans. Amer. Ent. Soce., vol. 33, 1907, p. 62.

EHnarmonia xanthospora Meyrick, Ent. Mo. Mag., vol. 48, 1912, p. 34.

Enarmonia eoleuca Meyrick, Ent. Mo. Mag., vol. 48, 1912, p. 34.

Laspeyresia vana BARNES and McDunNovuGH, Check List Lepid. Bor. Amer.,
no. 7227, 1917.

Laspeyresia zana BARNES and McDuNnNoveH, Check List Lepid. Bor. Amer.,
no. 7228, 1917.

Laspeyresia caeruleana BARNES and McDuNNovuGeH, Check List Lepid. Bor.

Amer., no. 7229, 1917.

Male paratype from Fernald collection in the National Museum.

The genitalia (male and female) of Kearfott’s two species and
caeruleana agree in all details. There is a slight difference in some
specimens in the color of the hind wing cilia, but this is not con-
stant for any given locality or collecting date, and the gradation
is gradual from white to pale smoky fuscous.. Kearfott was unable
to distinguish the different forms himself, for he had several speci-
mens of each set aside under another name (“banduna’’) as cotypes
of a new species.

32 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Male abdominal tufts consisting entirely of fine hairs.

Hind wing with veins 3 and 4 connate of very short stalked.

Genitalia figured from specimens in National Collection from
Mount Tzouhalem (“24-V-23, E. H. Blackmore No. 439,” male)
and Wellington, British Columbia (“'T. Bryant, VI,” female).

Distribution —British Columbia, California, Colorado, Nevada.

Alar expanse.—10-14 mm.

Types.—In British Museum (caeruleana); American Museum
(Zana and vana).

Type localities—Rogue River, southern Oregon, (caeruleana) ;
Wellington, British Goce heeaahs Colfax, Placer County,
Calif. Ppa

7. GRAPHOLITHA VITRANA Walsingham

(Figs. 131, 303)

Grapholitha vitrana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol. 4,
1879, p. 65.

Enarmonia vitrana FERNALD, in Dyar List N. Amer. Lepid., no. 5278, 1903.

Laspeyresia vitrana BaRNES and McDunNnovuGeH, Check List Lepid. Bor.
Amer., no. 7230, 1917.

Paratype in National Collection.

Similar and very close to the preceeding but apparently distinct.
The white dusting on the middle of fore wing forms (in normal
specimens) an indistinct pale median somewhat larger transverse
shade which is much fainter or entirely lacking in caeruleana; vi-
trana also has somewhat larger male genitalia and a differently
shaped genital plate in the female.

The cornuti of the two species are similar (one or two rather
long stout attached spines and a cluster of shorter deciduous ones) ;
in caeruleana there are two of the permanently attached spines while
in vitrana there appears to be only one. The latter species is some-
what variable. In some specimens the hind wing is distinctly pale
(whitish) toward base while in others it is evenly dark throughout.

Male abdominal tufts consisting of mixed flattened and cylindrical
hairs.

Hind wing with veins 3 and 4 connate or very short stalked.

Genitalia figured from specimens in National Collection from
Carmel, Calif. (“A. H. Vachell, IV.”).

Specimens in National Collection, American Museum, and collec-
tion Barnes from California.

Alar expense.—15-16 mm.

Type.—In British Museum.

Type locality —Northern Oregon.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 33

8. GRAPHOLITHA FANA (Kearfott)
(Figs. 7, 126, 297)

Enarmonia fana Kerarrotr, Trans. Amer. Ent. Soc., vol. 33, 1907, p. 64.

Enarmonia oenochroa Meyrick, Ent. Mo. Mag., vol. 48, 1912, p. 34.

Laspeyresia fana BARNES and McDuNNouGH, Check List Lepid. Bor. Amer.,
no. 7213, 1917.—Forsrs, Memoir 68, Cornell Univ. Agr. Expt. Sta., 1924,
p. 393.

Hind wing with veins 3 and 4 short stalked.

Genitalia figured from specimens in National Collection from
Anglesea, N. J. (W. D. Kearfott, “ V—29-05,” male paratype) and
Plummer Island, Md. (R. C. Shannon, June 9, 1914, female).

_Distribution—New Jersey, Maryland, North Carolina, Ohio,
Kansas.

Alar expanse.—8-9 mm.

Type—tIn American Museum.

Type locality—Anglesea, N. J.

Food plant—Meibomia (larvae in flower heads and terminal
buds. )

9. GRAPHOLITHA CONVERSANA Walsingham
(Figs. 133, 301)

_Grapholitha conversana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol. 4,
1879, p. 66.

Enarmonia conversana FERNALD, in Dyar List N. Amer. Lepid., no. 5272,
1903.

Enarmonia wana Krarrott, Trans. Amer. Ent. Soc., vol. 33, 1907, p. 60.

Enarmonia cupida Meyrick, Ent. Mo. Mag., vol. 48, 1912, p. 34.

Laspeyresia conversana BaRNES and McDunnoueH, Check List Lepid. Bor.
Amer., no. 7223, 1917.

Laspeyresia wana Barnes and McDunnovueH, Check List Lepid. Bor.
Amer., no. 7226, 1917.

A Pacific coast species much resembling the eastern fana but with
different genitalia; the female genital plate and the chitinization of
the ductus are quite differently shaped and the aedoeagus is much
stouter. The cornuti in conversana are arranged upon a horseshoe
shaped chitinous band and are all deciduous except one rather slen-
der spine which is fixed at one of the tips of the band. The wing
patern is somewhat variable. In a large reared series from Oregon
in the National Museum some specimens show but faint traces of the
dorsal white marks and a few are entirely without them. Normally,
however, the dorsal white geminations are present and sharply de-
fined. The abnormal specimens will run to caeruleana in our key, but
can be separated by their genitalia.

Kearfott’s wana is a straight synonym, agreeing in all details with
a paratype of conversana from the Fernald collection in the National
Museum.

84 BULLETIN 132, UNITED STATES -NATIONAL MUSEUM

Hind wing with veins 3 and 4 connate or very short stalked.

Genitalia figured from paratypes in National Collection from San
Luis Obispo (“ March, A. H. Vachell,” male) and Carmel, Calif.
(“ A. H. Vachell, IV,” female).

Distribution California, Oregon, Washington, British Columbia.

Alar expanse.—8-12 mm.

Types—In British Museum (conversana); American Museum
(wana).

Types localities —“ Camp Watson, on John Day’s River, Oreg.”
(conversana) ; Carmel Calif. (wana).

Food plants—Trifolium fimbriatum, cranberry (U.S. Bureau of
Entomology rearings).

10. GRAPHOLITHA IMITATIVA, new species

(Fig. 134, 298)

A western species similar to conversana and lunatana in pattern
but distinguishable from both by the genitalia andthe characters
given in the key. The sacculus of the harpe is considerably longer
in proportion to the rest of that organ in émétativa than in either
of the other two species.

Antenna brown. Palpus sordid grayish white more or less dusted
with fuscous. Head brownish fuscous. Fore wing semilustrous
dark brownish fuscous with a faint scattered dusting of whittish
scales at base, on disk and along termen; costa with four pairs
of short white geminate dashes, a pair before middle, two pairs
close together beyond middle and a pair close to apex; the first of
the antemedian geminations, the first and fourth of the postmedium
and the outer of the apical are continued for a short distance as
leaden metallic streaks; from mid dorsum a narrow outwardly
curved white geminate mark (sometimes partially divided by a
thin line of brown scaling) extends to middle of cell; ocelloid patch
consisting of a moderately long, vertical, metallic inner bar and a
much shorter, slanting outer bar inclosing two longitudinal black
dashes (in upper part of patch) and two or three small irregular
black dots upon a whitish or whitish ocherous ground; cilia pale
lustrous leaden fuscous with black basal line, outwardly finely edged
with whitish scaling, the black line cut below apex by an obscure
white spot. Hind wing dark brown, almost concolorous with fore
wing; cilia pale, smoky, with dark basal band and the tips of the
hairs shading to white and with a white spot on costa close to apex;
veins 3 and 4 connate.

Genitalia figured from paratypes in National Collection from
joldstream, British Columbia (“31-Y-21, E. H. Blackmore No.
378,” male and San Francisco, Calif. (female). Male abdominal
tufts consisting entirely of fine hair like scales.

- NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 35

Alar Hxpanse.—10.5-12 mm.

Type and paratypes—Cat. No. 28014, U.S.N.M. Paratypes also
in American Museum, Canadian National Collection and collections
Barnes and Blackmore.

Type locality—San Francisco, Calif.

Described from male type, 8 male and 7 female paratypes from the
type locality; 5 male and 1 female paratypes from the Hy. Edwards
Collection of the American Museum labeled “ California,” “3851,”
and “722”; 4 male and 2 female paratypes from Goldstream,
British Columbia (E. H. Blackmore, “ 16-IV-21,” “17-IV-21,” and
“31-IV-21” Blackmore numbers “378,” and “379” and “796”);
1 male and 1 female paratype from Fitzgerald, British Columbia
(E. H. Blackmore, “ 14-V-1922”); 1 male paratype from Salmon
Arm, British Columbia (“15-5-21, W. R. B.”); and 1 female para-
type from Waterton Lakes, Alberta (J. McDunnough, “20-VI-
1923”).

This species has been appearing in our collections as conversana
Walsingham. The true conversana is close to it but distinct and
equal to what Keargott described as wana. G. imitativa is most like
lunatana but has termen of fore wing decidedly less slanting.

11. GRAPHOLITHA LUNATANA Walsingham
(Figs. 128, 300)

Grapholitha lunatana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol. 4,
1879, p. 66.

Enarmonia lunatana FERNALD, in Dyar List N. Amer. Lepid., no. 5274, 1903.

Laspeyresia lunatana BaRNES and McDuNnnoucu, Check List Lepid. Bor.
Amer., no. 7225, 1917.

A somewhat variable species as far as color of hind wings and size
are concerned. In some specimens the hind wing is distinctly whitish
toward base; in others it is evenly dark throughout. The aedoeagus
is very long and stout as compared with the rest of the male genitalia.
The cornuti consist of several deciduous and two slender, straight
fixed spines, the latter nearly as long as the aedoeagus. The decid-
edly slanting termen of forewing distinguishes it from zmétativa and
conversana which it most resembles.

Hind wing with veins 3 and 4 connate.

Genitalia figured from specimens in National Collection from
San Luis Obispo (A. H. Vachell, March, male) and Los Angeles,
Calif. (“ Coquillet, No. 149,” female).

Distribution.—Colorado, California, Washington, British Colum-
bia, Alberta, Saskatchewan, Manitoba.

Alar expanse.—10-15 mm.

Lype.—tn British Museum.

Type locality.—North Oregon.

54346—26——4

86 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

12. GRAPHOLITHA ECLIPSANA Zeller
(Figs. 124, 302)
Grapholitha (Ephippiphora) eclipsana ZELLER, Verh. Zool.-bot. Ges. Wien,
vol.-25, 1875, p. 298.
Enarmonia eclipsana Krarrott, Bull. Amer. Mus. Nat. Hist., vol. 23, 1907,
p. 159.
Laspeyresia eclipsana Barnes and McDuNnNoucH, Check List Lepid. Bor.

Amer., no. 7216, 1917.—Forsres, Memoir 68, Cornell Univ. Agr. Exp.
Sta., 1924, p. 393.

A striking species easily distinguished by its shining white, api-
cally dark dusted hind wings. Not to be confused with anything
else.

Hind wing with veins 3 and 4 connate.

Genitalia figured from specimens in National Collection from Oak
Station, Pa. (F. Marloff, “ TV-17-15,” male and female).

Distribution.—North Carolina, Virginia, Maryland, Pennsylvania,
New York, Ohio.

Alar expanse.—9-15 mm.

Type.—In Museum of Comparative Zoology.

Type locality.—Texas.

Groupe B—OcELLoID PatcH oF ForE WING CONSISTING OF A SINGLE VERTICAL
METALLIC BAR oR A PAIR OF CLOSELY APPRESSED VERTICAL BARS

13. GRAPHOLITHA INTERSTINCTANA (Clemens)
(Figs. 139, 306)

Stigmonota interstinctana CLEMENS, Proc. Acad. Nat. Sci. Philadelphia,
1860, p. 351.

Dichrorampha scitana WALKER, Cat. Lepid. Heter. Brit. Mus., vol. 28,
1863, p. 4138.

Grapholitha distema Grote, Bull. Buffalo Soc. Nat. Sci., vol. 1, 1878, p. 92.
Grapholitha (Ephippiphora) interstinctana ZELLER, Verh. Zool.-bot. Ges.
Wien, vol. 25, 1875, p. 296.

Enarmonia interstinctana FERNALD, in Dyar list N. Amer. Lepid., no. 5270,
1903.—Fotsom, Bull. 134, Illinois Agr. Exp. Sta., 1909, p. 184.

Laspeyresia interstinctana Barnes and McDunNnouGH, Check List Lepid.
Bor. Amer., no. 7210, 1917.—-Forses, Memoir 68, Cornell Univ. Agr.
Exp. Sta., 1924, p. 393.

A well known clover insect. The life history and a bibliography
of the principal references are given in Folsom’s bulletin.

Genitalia figured from specimens in National Collection from Dis-
trict of Columbia (‘“ No. 82, in flower head of red clover, iss. July 2,
79,” male) and Plummer Island, Md. (Busck, Aug. 1903, female).
Bursa of female without signa. Hind wing with veins 3 and 4
connate.

Distribution—North Carolina, Virginia, Maryland, District of
Columbia, Pennsylvania, New Jersey, New York, New Hampshire,

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 37

Vermont, Minnesota, Iowa, Illinois, Missouri, Ontario, Quebec,
Alberta.

Alar expanse.—T-12 mm.

Types.—In Academy Natural Sciences, Philadelphia (tnterstinc-
tana) ; British Museum (scintana and (?) distema).

Type localities—Pennsylvania (énterstinctana); “ North Amer-
ica” (scintana); New York (distema).

Food plants.—Trifolium (larvae in flower heads and stems).

14. GRAPHOLITHA EDWARDSIANA (Kearfott)

Enarmonia edwardsiana Krarrotr, Trans. Amer. Ent. Soc., vol. 33, 1907,
. 58.
Lasbea cab edwardsiana BARNES and McDunNovueH, Check List Lepid. Bor.
Amer., no. 7218, 1917.

Represented so far only by the cotypes in the American Museum
and Barnes collections and a specimen from the Fernald collection
(California) in the National Museum. I am unable to find the cotype
which Kearfott states was deposited in the National Collection.

Very similar in pattern and structure to Jana Kearfott from which
it differs in its distinctly white hind wing cilia; lana is probably
either a synonym or a local race.

Hind wing with veins 3 and 4 connate.

Genitalia as in lana.

Alar expanse.—14 mm.

Type—aIn American Museum.

Type locality.—California.

15. GRAPHOLITHA LANA (Kearfott)
(Figs. 137, 308)

Enarmonia lana Kearrott, Trans. Amer. Ent. Soc., vol. 38, 1907, p. 59.

Enarmonia placerana Kearrort, Trans. Amer. Ent. Soe., vol. 33, 1907, p. 60.

Enarmonia vancouverana Kearrott, Trans. Amer. Ent. Soc., vol. 33, 1907,
p. 63.

Enarmonia chrysotypa Meyrick, Ent. Mo. Mag., vol. 48, 1912, p. 34.

Laspeyresia lana BARNES and McDunNnoucH, Check List Lepid. Bor.
Amer., no. 7219, 1917.

Laspeyresia placerana BARNES and McDuNNouGH, Check List Lepid. Bor.
Amer., no. 7221, 1917.

Laspeyresia vancouverana BARNES and McDuNNovueH, Check List Lepid.

Bor. Amer., no. 7212, 1917.

Variable in size and character of the dorsal geminate markings
of fore wing. Typical /ana has four white dorsal dashes and typical
placerana and vancowverana two; but in a large series of moths in
the Barnes Collection from Loma Linda, Calif. (“July 24-31”),
and several specimens from Mineral King, Calif. (“July 1-7”),
there is every gradation, some specimens showing two distinct lines,

38 BULLETIN 182, UNITED STATES NATIONAL MUSEUM

others two lines partially divided by dark scaling, and still others
with the normal two lines so completely broken as to make four. If
the names were kept separate it would be impossible to decide under
which many specimens would go. In fact, the Kearfott paratype
of placerana in the National Collection has four distinct white marks
and agrees more closely with his type of Jana than with any of the
other types of placerana. There are some differences in the size
and shape of the harpes of the male genitalia between the actual
types of placerana and vancouverana; but in a series of moths from
Goldstream, British Columbia, all collected upon the same day
(June 1, 1921) the intergrades are such that it is impossible to tell
under which name some of the specimens should go if the two
names are to be kept separate. The species is probably a stem borer
or seed-capsule feeder, which habit would easily account for the
variability. A reared series of S. tristrigana shows quite as much
variation. I am of the opinion that Jana is only a race of edwards-
tana, from which it differs chiefly in the darker cilia of its hind
wing, not a good character. This synonym, however, will have: to
wait upon rearings before it can be definitely settled.

Hind wing with veins 3 and 4 connate.

Genitalia figured from specimens in National Collection from
Placer County, Calif. (“June,” male cotype of placerana), and
Goldstream, British Columbia (“1-VI-21, E. H. Blackmore, No.
428,” female).

Distribution.—California, British Columbia.

Alar expanse.—10-15 mm.

Types.—In American Museum.

Type localities—Colfax, Placer County, Calif. (lana and place-
rana) ; Wellington, British Columbia (vancouveruna).

16. GRAPHOLITHA DYARANA (Kearfott)
(Fig. 185)

EHnarmonia dyarana Krarrott, Trans. Amer. Ent. Soc., vol. 33, 1907, p. 58.
Laspeyresia dyarana Barnes and McDunnovuGH, Check List Lepid. Bor.
Amer., no. 7217, 1917.

Females of this species average somewhat larger than lana and
show some slight differences in genitalia, having the chitinization
about genital opening much weaker. It is probable that the two
are only racially distinct. The determination of this, however, will
have to wait upon rearings.

Female genitalia figured from type. Male genitalia as in Jana.

All specimens in the collections are from Colorado localities.

Alar expanse.—16-17 mm.

Type.—In American Museum.

Type locality.—Colorado.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 39
17. GRAPHOLITHA TRISTRIGANA (Clemens)

(Figs. 136, 307)

Stigmonota tristrigana CLEMENS, Proc. Ent. Soc. Philadelphia, vol. 5, 1865,

p. 133.

Enarnonia tristrigana Fernatp, in Dyar List N. Amer. Lepid., no. 5275,
19038.

Enarmonia saundersana Krarrorr, Trans. Amer. Ent. Soe., vol. 33, 1907,
p. 63.

Laspeyresia saundersana BARNES and McDuNnnoucH, Check List, Lepid.
Bor. Amer. no. 7215, 1917.—Forses, Memoir 68, Cornell Univ. Agr.
Exp. Sta., 1924, p. 393.

Laspeyresia tristrigana BaRNES and McDunnoucnH, Check List Lepid.
Bor. Amer., no. 7220, 1917.—Hetnricu, U. S. Journ. Agr. Res., vol.
20, 1921, p. 824.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,;
1924, p. 394.

Kearfott’s types of sawndersana are simply runted specimens of
tristrigana. We have a considerable series of the latter reared from
both Baptista and Lupinus and among the lot are several specimens
agreeing in every detail with typical saundersana as well as specimens
intergrading in size between it and typical tristrigana. There are
no structural differences.

Hind wing with veins 3 and 4 connate.

Genitalia figured from specimens in National Collection from E]1
Vista, Tex. (reared from seed pods of Baptisia May 15, 1919, Hein-
rich, male and female). Bursa of female without signa.

Mstribution.—IUllinois, Kansas, Oklahoma, Alabama, Texas,
Florida, North Carolina, Pennsylvania, New Jersey, New York,
Massachusetts, Ontario.

Alar expanse.—10-16 mm.

Types—In Academy Natural Sciences (tristrigana); American
Museum (saundersana).

Type localities.—Virginia (tristrigana); Toronto, Ontario, Can-
ada (saundersana).

Food plants.—Baptisia, Lupinus (larvae in seed pods and stems).

10. OFATULENA, new genus
(Figs. 30, 119)

Genotype—Grapholitha? duodecemstriata Walsingham (North
America).

Thorax smooth.

Fore wing smooth; termen straight or very slightly concave; 12
veins; 7 to termen; 11 from cell at middle; 10 rather well separated
from 9; upper internal vein of cell from between 10-11; 3, 4 and
5 parallel from beyond cell, not approximate at termen; 2 from cell
at 24, straight; no costal fold in male.

40 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Hind wing with normal pecten ; 8 veins; 6 and 7 approximate
toward base; 3 and 4 connate; inner mar are simple in male.

Hind iia smooth scaled.

Male genitalia with a row of long stout, flattened marginal spines
at lower outer angle of harpe; cucullus densely spined; basal open-
ing greatly enlarged; no appreciable neck incurvation; sacculus
reduced, sparsely haired. Uncus absent. Socii represented as
heavy hair tufts at extremity of tegumen. Gnathos poorly defined,
weakly chitinized. Aedaeagus long, slender, straight; conuti absent.

Abdomen of male simple.

Female genitalia with two signa developed as short thorn-like
spines. Ductus bursae moderately long, ‘unchitinized. Bursa copu-
latrix roughly triangular, smooth. Genital plate absent.

A small North American genus affiliated with Laspeyresia.

KEY TO THE SPECIES OF OFATULENA

1. Basal half of fore wing whitish gray, finely cross marked with fuscous;
ground color of ocelloid patch whitish; hind wing blackish fuscous.

(1) duodecemstriata.

Basal half of fore wing grayish fuscous, unmarked by transverse lines;

ground color of ocelloid patch ocher yellow; hind wing dark smoky brown.

(2) luminosa.

1. OFATULENA DUODECEMSTRIATA (Walsingham)
(Figs. 30, 119, 309)

Grapholitha? duodecemstriata WALSINGHAM, Trans. Ent. Soc. London,
1884, p. 146.

Enarmonia duodecemstriata FrrNaLp, in Dyar List N. Amer. Lepid., no.
5284, 1903.

Laspeyresia duodecemstriata BARNES and McDunnovueH, Check List Lepid.
Bor. Amer., no. 7240, 1917.

A beautiful species easily recognized by its whitish gray, finely
cross-striped fore wing with large many striped ocelloid patch and
its black brown hind wing.

Genitalia figured from specimens in National Collection from
Tempe, Ariz. (“July 19, 20,” male) and La Puerta, Calif. (“July,
1911, Wright and Field,” female).

Distribution.—Texas, Arizona, Utah, Nevada, California.

Alar expanse.-—14-17 mm.

Type—Iin British Museum.

Type locality —Arizona.

Food plant.—Mesquite beans.

2. OFATULENA LUMINOSA, new species

(Figs. 120, 310)

Palpus white. Face white. Head pale ocher yellow shading to
white in center. Thorax fuscous gray; tegula paler, whitish gray;

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 41

collar white. Fore wing grayish fuscous; at extreme base a few
whitish scales and on mid dorsum a pair of very faint pale gray out-
wardly curved dashes extending upward to cell; ocelloid patch con-
sisting of two vertical leaden metallic bars (the outer slanting
inward a little and parallel to termen) inclosing a varying number
of fine irregular black dots or streaks upon an ocher yellow ground;
subcostal area above ocelloid patch ocher yellow; from costa just
beyond middle a narrow metallic outwardly slanting bar, joining
inner vertical bar of ocellus; beyond this four pairs of very short,
narrow, white, geminate dashes, from the second of which extends a
thin metallic bar (parallel to the inner one from costa) and joining
outer vertical bar of ocellus; a similar, short, curved metallic line
from outer pair of white costal dashes; apex and terminal area
beyond ocellus, grayish fuscous with scale ends white; cilia leaden
fuscous, semimetallic. Hind wing dark smoky brown, paler toward
base; cilia white with a dark basal band.

Male genitalia figured from type; female from paratype in Na-
tional Collection.

Alar expanse.—12-14 mm.

Type.—tn American Museum.

Paratype.—Cat. No. 28015, U.S.N.M.; also in collection Barnes.

Type locality Brownsville, Tex.

Described from male type from Brownsville, Tex. (Townsend, “19
June, 95”) ; and four female paratypes from San Benito, Tex., dated
“Apr. 1-7,” “Apr. 24-30,” and “Sept. 8-15.”

Close to duodecemstriata but distinct and easily separated on color
and structure; in /wminosa the aedoeagus of the male is shorter and
stouter than in duodecemstriata; the large outer spines of harpe are
also more thickly clustered (in the photograph they do not show
well as most of them were broken off before the slide was made).

The male type had been identified by Kearfott as “ Pammene texa-
nana Walsingham” and was in the American Museum under that
name.

11. Genus LASPEYRESIA Hiibner

Laspeyresia Htener, Verz. Schmet., 1826, p. 381.
Genotype.—Tortrix corollana Hiibner (Hurope).

Endopsia GUENEE, Eur. Microlepid. Index Method., 1845, p. 48.
Genotype.—Pyralis nigricana Stephens (Europe).

Cerata STEPHENS, List Brit. Animals, pt. 10, Lepid., 1852, p. 77.
Genotype.—Penthina servillana Duponchel.

Characters as in Grapholitha except:

Fore wing with termen concave, straight or convex; veins 3, 4 and
5 separate or slightly approximate at termen.

42 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Hind wing with 8 and 4 connate, stalked or united; inner margin,

in male, sometimes with rought sex scaling; male rarely with a slight
fold along vein 1c inclosing or partially inclosing the pecten from
lower median vein.

10.

ali ls

Abdomen of male simple.

KEY TO THE SPECIES OF LASPEYRESIA

Fore wing with a broad well-defined median or antemedian transverse
fascia; or a whitish suffusion at middle extending from dorsum to costa
and much extended on latter; or with basal half of wing whitish__-__ 2.

Here AwiNe GtHeIwise st tts 208s be BAN 28 NS ee ee 9.

. Antemedian pale area of fore wing distinguished as a broad fascia bounded

inwardly and outwardly by narrow metallic bands extending from
COStR tO GONSUM oe 2 ee pe ES pe ee ee 3
Pale area covering a greater part of basal half of wing; when defined as
an antemedian fascia, not bounded by metallic bands__-_--------_- 4,

. Antemedian fascia whitish ocherous, strongly contrasted against dark basal

and median’ areas_—_-..-_ BORG 2ReRe RG Boys eee (21) tana.
Antemedian fascia grayish ocherous only slightly paler than basal band and
MeCIANAATCAS— ss -seskbe, SN aes Pe ee a ees (22) cupressana.

. Thorax and extreme base of fore wing concolorous with antemedian pale

SCE eee ea AA a ie Se eS eee (20) fletcherana.
Thorax and extreme base of fore wing darker than antemedian area__- 5.

. Dark basal patch of fore wing distinct only to top of cell; costa at base

white and but faintly marked by dark scaling; hind wing white

Basal patch complete to costa; costa at base dark or very strongly aigrked
with dark scaling; hind wing not white at base____________-_-_____ 1@

. Thorax white with a dark narrow transverse median band; entire patagia

white; alar expanse less than 14 mm_____-~--_----+_ (25) gallaesaliciana.
Thorax black with white dusting on posterior half; patagia with a blackish
shade on anterior margin; alar expanse over 15 mm____ (26) lautiuscula.

. Basal patch leaden fuscous or semilustrous fuscous drab__-__-__----~_ 8.

Basal patchy PlaCkiShe seco melon eee eS 2 oe ee (27) flexiloqua.

. Dark terminal area of fore wing strongly dusted with black; longitudinal

black streaks of ocelloid patch not sharply defined, more or less fused into
the black dusting of the entire terminal area_____-______~_ (24) leucobasis.
Dark terminal area with only scattered black dustings; longitudinal black
streaks of ocelloid patch sharply defined and interspaced with ocherous

JEOESYO OU (Se oe ee ee ek ee ee (23) prosperana.

. Fore wing with greater part of costal area white__________ (14) populana.
Fore wing with greater part of costal area gray or brown- —~-----~--- 10.
Terminal area-of fore, wing pink;or red... -<c 2Sece) sfeeehs 1
No pink or red shading, in terminal area. 2 —- oye tt. Sete 12.

Terminal area pink; ocelloid patch consisting of a single vertical leaden
metallic bar inwardly margined by a narrow blackish fuscous shading and
(on its upper half) by a second narrow metallic bar__--__~_ (30) ninana.

Terminal area red; ocelloid patch consisting of two vertical bronzy metallic
bars inclosing six or seven longitudinal blackish fuscous streaks upon a
White ground 22220 yeep sb he cpt pyepecet te NEE bel (29) flavicollis.

12.

13.

14.

15.
16.

18.

19.
20.
21.

22.

23.
24.
25.

26.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 43

Fore wing with a single, conspicuous, undivided white spot on mid
CNS a ee apaapgeg ee ry Mag ta he A ays beets a betes eg 138.
Fore wing without such; white dorsal markings when present consisting
of a pair of geminate marks or a white line bordering a narrow trans-
verse hmetallic) barse) 2 a4 eae oh tgs ee tn gil eee ee OE 14.
White dorsal spot not extending above vein 1b; ground color of fore wing
to naked eye, dark gray; a conspicuous black spot on mid costa; hind
wing! white! toward basew2i2 jru! acid eee! 5 eee (13) albimaculana.
White dorsal spot extending up into cell; ground color of fore wing to naked
eye, dark purplish brown; no such black spot on mid costa; hind wing

brown. throvghontis1 otetiyeisp vfterts lees we fteih. et bbe (28) americana.
Fore wing grayish ocherous, vertically marked by broken leaden-metallic

ease Ye ag hem Dy erin 8 ere egg Hf Be (81) colorana.
Hore wing otherwise. {4st 24 eters pb fed tt nay jiu, wires Fs 15.
General-color of fore wing; gray ocs2- ween eta ee sy 16.
General color of fore wing;| brownas: 2. 3 led wets sh by bee 20.
Hind wing distinctly white toward base.u_-.. 2. Lf.
Hind wing not appreciably whitish toward base____________-______ 18

Three pairs of outer-costal white geminate marks equidistant.

(11) multilineana.
Third (apical) pair of outer-costal white geminations well separated from
Ober C30: eres adi ie E piijeyy tow edep Si, porn (12) ingrata.
Dorsum of fore wing at extreme base dark, unmarked by white. gemina-
GIONS! Sep seceeryeey Arete tee, FA Bo ppelere pa eye. wip cer sais (10) membrosa.
Several white geminate marks on base of dorsum, or a white shade extend-
ing along dorsum to base from median dorsal geminate marks_______ 19.

Ocelloid patch containing three distinct longitudinal black streaks.
(8) larimana.

No black streaks in ocelloid patch 20220 2 (9) garacana.
Fore wing with a widely spaced pair of narrow, strongly marked, metallic

transverse bands; one near and the other beyond middle___-_________ 21.
Forel wing without .suéhieit sec i ae ere mei obese 23.

First transverse metallic band inwardly margined with white.
(32) erotella.

First transverse band not margined with white_____._____-___+_-_____ 22.
Hind wings very dark brown; cilia cream white, strongly contrasted against
SY Ena Se ree Pees Da we thc oe al en Roe ee Se ee (36) miscitata.

Hind wing rather pale brown; cilia whitish but not strongly contrasted.
(33) toreuta.
(34) ingens.
(35) piperana.
Fore wing with a pair of white geminate dashes on dorsum near middle. 24.
Fore wing without white markings on dorsum; dorsal geminate marks
Wiens presents leaden. Meta WiG@ ws sat eo ee 29.
Thorax and base of fore wing as dark as darkest outer shade of wing.
(3) laricana.
Thorax and base of fore wing paler than outer dark shading of wing__- 25.
Ocelloid patch containing only two longitudinal black dashes.
(7) parmatana.
Ocelloid patch containing four or more longitudinal black dashes______- 26.
Fore wing heavily dusted with black beyond base______________ (6) obnisa.
Fore wing with little or no black dusting beyond base_____-_-_-______-- at.

44 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

27. Seven white geminate marks on costa________-_----+---=-4_-__-_- (4) rana.
Ten--white-geminate marks: on- costas.-2—=2. See eee eo 28.
26) ‘Alar ‘expanse ‘under’15 mimi $2200 olube 2 costes Siri (1) bracteatana.
Alar expanse Over!15 mmUL Oley fF: Se) AACE OPT (2) cornutana.

29. Hind wing blackish brown, much darker than fore wing; hind wing cilia
snow white with no dark basal band_-_----_--______-___ (18) grandicula.

Hind wing brown, sometimes quite dark, but not distinctly darker than
fore wing; cilia more or less white but.always with dark basal band_ 30.

30. Fore wing with a pair of mid dorsal metallic geminate marks (or a single
metallic spot) fusing above with a similar pair from costa beyond base

to form a faint but distinguishable angulate metallic fascia___-____ 31.
Horewine without: SuehUvATL AE FUL Otay ery raiog. lary go irae al 38.

31. Hind wing with veins 3 and 4 united_______________________- (5) inopiosa.
Hind wing with veins 3 and 4 connate or stalked_____-________________ 32.

32. Fore wing with median brown area between costa and dorsum heavily
dusted with black; outer half of costa with four to six whitish geminate
dashes; alar expanse less than 12 mm__________________ (15) youngana.

Fore wing with scales of median brown area between costa and dorsum
tipped with ochreous, no appreciable black dusting; outer half of costa
with eight or nine whitish geminate dashes; alar expanse 12 mm. and
OVERSLG 9s SUT BOS RSS OTT UR IAOS testis Te Teas tie (17) candana.

33. Termen of fore wing concave; veins 3 and 4 bent upward at middle; hind
wane whitish on Costh#G_.2TbD S280 Soy bi aie spt (19) caryana.

Termen of fore wing convex; veins 3 and 4 not bent upward at middle;
hind ‘wing’ not whitish''on' costa 224) oo Sa aie siyiies 9 (16) nigricana

1. LASPEYRESIA BRACTEATANA (Fernald)
(Figs. 311, 312)

Grapholitha bracteatana FERNALD, Rep. U. S. Dept. Agr. for 1880, 1881,
p. 265.

Enarmonia bracteatana FERNALD, in Dyar List N. Amer. Lepid., no. 5271,
1903. :

Laspeyresia bracteatana BARNES and McDunnoueH, Check List Lepid.
Bor. Amer., no. 7222, 1917.

Laspeyresia pallidibasalis HetnricH, Proc. U. S. Nat. Mus., vol. 57, 1920,
p. 60.

When I described pallidibasalis I distinguished it from bractea-
tana by its size and a difference in genitalia. In a large reared series
the specimens averaged larger than Fernald’s types and showed a
deeper emargination of the harpe. These differences, I am now con-
vinced are not significant. The Fernald types are runted specimens;
and while they differ from typical, or average pallidibasilis, in the
characters mentioned, they do agree very well with occasional small
specimens of the latter. I am therefore sinking my name.

Male genitalia figured from paratypes of bracteatana and pallidi-
basalts in the National Collection. These photographs show the ex-
treme differences in harpe shape. Cornuti a dozen or more short
fixed spines arranged in a row. The female genitalia are as in varie-
ty cornutana Dyar.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 45

Veins 3 and 4 of hind wing stalked.

Distribution.—California and Oregon.

Alar expanse.—9-14 mm.

Types—tIn National Collection.

Type localities —Jolon, Calif. (bracteatana) ; Kaolin Beds, Oreg.
(pallidibasalis).

Food plant.—Abies concolor (Larvae feed in cones or bracts and
seeds).

2. LASPEYRESIA BRACTEATANA CORNUTANA (Dyar)
(Fig. 151)

Epinotia cornutana Dyar, Proc. Ent. Soc. Washington, vol. 5, 19038, p. 231.
Enarmonia cornutana BARNES and McDuNnNouGH, Check List Lepid. Bor.
Amer., no. 7166, 1917.

The only specimen of this I have seen is the female type. It has
nothing to separate it from bracteatana except its larger size and
darker banding (somewhat heavier chitinization) of the abdominal
segments, a character not visible until the abdomen is completely
denuded. For the present the name should be retained. It probably
designates a good food plant or local race.

Genitalia figured from type.

Alar expanse.—17 mm.

' Type—tin National Collection.

Type locality —Williams, Ariz.

3. LASPEYRESIA LARICANA (Busck)
(Figs. 155, 317)

Laspeyresia laricana Buscx, Proc. Ent. Soc. Washington, vol. 18, 1916, p.
152.— Barnes and McDunnoucHu, Check List Lepid. Bor. Amer., no.
7251-1, 1917.

This and the following two species are cambium miners in the
bark of coniferous trees. They are very similar and possibly only
food plant races of a single variable species. Further rearing will
be necessary to determine this. The pattern differences are shown
in the figures. The female of /aricana has the genital opening larger
and the genital plate more strongly chitinized than those of the
others. The male has the aedoeagus quite stout, straight, and not
perceptibly tapered; the cornuti are arranged in three clusters of
stout fixed spines, some very short and some moderately long.

Hind wing with veins 3 and 4 connate or stalked.

Genitalia figured from specimens in the National Collection from
Missoula (male, reared under Hopk. U. S. No. 11551, May, 1913,
from larva in cambium of Douglas fir, J. C. Brunner, collector) and
Evaro, Montana (female type, reared under Hopk. U. S. no. 12830,
May 5, 1914).

46 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Specimens in National Collection and collection Barnes from
Montana.

Alar expanse.—14.5-16.5 mm.

Type.—tn National Collection.

Type locality.—Evaro, Mont.

Food plants.—Lariz occidentalis, Pseudotsuga taxifolia.

4, LASPEYRESIA RANA Forbes
(Figs. 153, 314)

Laspeyresia rana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 394.

Kearfott had a single female under the above name in his collec-
tion but had never published a description; so the species will have
to be credited to Forbes. It is similar to Javicana Busck; but with
paler thorax and base of forewing; ocelloid patch containing 4
longitudinal black streaks; veins 3 and 4 of hind wing connate.

Genitalia figured from specimens in National Collection from
Missoula, Mont. (reared under Hopk. U. S. Nos. 11081 and 11082,
May 15 and June 1, 1915, from larvae feeding in bark of Picea engel-
manni, B. T. Harvey, collector). Male genitalia like those of brac-
teatana except: aedoeagus stout, bottle necked toward apex; cornuti
a double row of 8 to 10 short, weak, fixed spines.

Distribution.—North Carolina, Montana, Alberta, Ontario. The
specimens from Alberta (Nordegg) are a trifle darker than the
others but otherwise agree.

Alar expanse.—13-14 mm.

Type.—titn American Museum.

Type locality —Black Mountains, N. C.

Food plant.—Picea engelmannt.

5. LASPEYRESIA INOPIOSA, new species

(Fig. 165)

Similar to the preceding (rana) except:

Fore wing with geminate marks on dorsum, broad, leaden scaled ;
eight (rather than seven) white geminate marks on costa; ocelloid
patch with two broken black longitudinal dashes set close together
near top or with two or three black dots in place of the usual black
lines; veins 4 and 5 closely approximate at termen.

Hind wing with veins 3 and 4 united; underside of wing (in male)
rough scaled toward base.

Female genitalia of type figured.

Alar expanse.—9-11 mm.

Type and paratype.—Cat. No. 28016, U.S.N.M.

Type locality Coeur d’Alene, Idaho.

Food plant—Pinus contorta.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 47

Described from female type and male paratype (latter without
abdomen) reared May 11, 1916, under “ Hopk. U. S. No. 18958-2,”
from larvae feeding in twigs of Pinus contorta infested by Pe-
trova albicapitana, taken at the type locality by J. C. Evendon.

6. LASPEYRESIA OBNISA, new species
(Fig. 164)

Similar to bracteatana Fernald but with darker wings, dorsal
geminations of fore wing broader and more fused, a narrow border
of ocherous fuscous scaling along termen, and different genitalia.

Palpus and face sordid whitish ocherous. Head, thorax, and fore
- wing a semilustrous leaden drab. Fore wing with termen slightly
concave below apex; blackish fuscous, with some dusting of ocherous
fuscous on outer half; from mid dorsum a broad pair of white
geminate dashes, fused at base and extended as bluish metallic
streaks to top of cell; from costa before middle a pair of short,
obscure, white dashes ending in bluish metallic scales but not fusing
with the dorsal patch; on costa beyond middle three pair of short
white streaks (repeated upon under surface of wing), and continued
in bluish metallic streaks to the vertical bars of ocellcid patch, the
outer pair well separated from the other two and close to apex;
ocelloid patch consisting of two vertical metallic bars inclosing five
or six longitudinal black lines interspaced with ocherous fuscous
scaling; termen narrowly bordered with ocherous fuscous; cilia
leaden fuscous with a black basal line; this black line and the
inner ocherous margin of termen cut below apex and above tornus
by white spots reproduced on under surface of wing. Hind wing
dark brown; cilia whitish with dark basal band; undersurface of
wing distinctly paler toward base; veins 3 and 4 stalked.

Female genitalia of type figured.

Alar expanse.—15-16 mm.

Type and paratype——Cat. No. 28017, U.S.N.M.

Type locality.—¥raser Mills, British Columbia.

Described from female type and paratype from the type locality
(“16-V1-1922” E. H. Blackmore and “30-VII-22” L. E. Marmont) ;
one female paratype from Brentwood, British Columbia (“14-VII-—
23” Blackmore); and one female paratype from Mount Newton,
British Columbia (“1—-VIII-20” Blackmore) ; all received under
Blackmore No. 472.

7. LASPEYRESIA PARMATANA (Clemens)

Ephippiphora parmatana CLEMENS, Proc. Acad. Nat. Sci. Philadelphia,
1860, p. 352.

Enarmonia parmatana FererNnatp, in Dyar List N. Amer. Lepid., no.
5283, 1903.

Laspeyresia parmatana BARNES «and McDunnovucH, Check List Lepid.
Bor. Amer., no. 7239, 1917.

48 BULLETIN 182, UNITED STATES NATIONAL MUSEUM

In the Academy of National Sciences at Philadelphia there is a
male which Fernald had labeled as possibly the Clemens type. It
is a Thiodia of the crispana-alterana group. This, however, can
not be the type, as Clemens takes pains to give details of the vena-
tion, which show plainly that his species is a Laspeyresia: “in the
medium vein” of hind wing, he says, “the upper nervula is forked
remotely from the insertion of the medio discal.” This rules out the
aforementioned 7’ hiodias which all have vein 5 very closely approxi-
mate to the stalk of 3 and 4 at base. Forbes® applied the name to
what we have been calling 7hiodia crispana Clemens, misunder-
standing, I think, Clemens’ venational terminology.

The only thing I have seen that answers at all to Clemens’ descrip-
tion is énopiosa Heinrich; but this is a western species which prob-
ably does not occur east of the Rockies. Kearfott had a number of
specimens under the name, only one of which is a Laspeyresia, and
it does not fit the description; the head is too pale and the thorax
too heavily dusted with white. Clemens’ description will apply
only to an eastern Laspeyresia with whitish palpi, dark brownish
head, thorax, and wings, a white divided dorsal mark on fore wing,
four pairs of white costal geminations, an ocelloid patch containing
two longitudinal black streaks upon a pale ground somewhat dusted
with ocherous scaling, and with veins 3 and 4 of hind wing stalked.
I have seen no such specimen in any of our collections.

Alar expanse.?—(not given).

Type.—Lost.

Type locality—Pennsylvania.

8. LASPEYRESIA LARIMANA (Walsingham)
(Fig. 316)

Eucelis larimana WALSINGHAM, Trans. Ent. Soc. London, 1895, p. 518.

Enarmonia larimana FERNALD, in Dyar List N. Amer. Lepid., no. 5286, 1903.

Laspeyresia larimana Barnes and McDunnovucH, Check List Lepid. Bor.
Amer., no. 7242, 1917.

The only thing I have seen that matches Walsingham’s descrip-
tion is the specimen Kearfott had under parmatana. It is a male,
originally from the Deitz collection, without locality and labeled
simply, “in timber, 6-14-01.” Its size is about right (13 mm.), and
in pattern and color it agrees in detail with the description of
larimana. I take it to be that species.

Male genitalia figured.

Hind wing with veins 3 and 4 stalked.

Alar expanse.—14.5 mm.

Type.—tiIn British Museum.

Type locality.—Loveland, Colo.

® Memoir 68, Cornell Uniy. Agr. Expt. Sta., 1924, p. 432.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 49

9. LASPEYRESIA GARACANA (Kearfott)
(Figs. 143, 313)

Enarmonia garacana Kerarrort, Trans. Amer. Ent. Soc., vol. 33, 1907, p. 66.
Enarmonia septicola Mryrick, Ent. Mo. Mag., vol. 48, 1912, p. 34.
Laspeyresia garacana BARNES and McDuNNovuGcH, Check List Lepid. Bor.
Amer., no. 7249, 1917.—Forsres, Memoir 68, Cornell Univ. Agr. Exp.
Sta., 1924, p. 394.
Close to larimana Walsingham; differing in genitalia and the ab-
sence of black markings from ocelloid patch.
Veins 3 and 4 of hind wing stalked.
Male and female genitalia figured from paratypes in National
Collection and American Museum from the type localities.
Aedoeagus slender, tapering; cornuti a half dozen very small,
thin attached spines placed in a row.
Distribution.—Texas, Illinois, Ontario.
Alar expanse.—14-15 mm.
Type.—In American Museum.
Type locality.—Chicago, Tl.

10. LASPEYRESIA MEMBROSA, new species
(Figs. 150, 322)

‘Antenna grayish, dusted with black toward base. Palpus sordid
white; third joint rather long, porrected and exposed. Face and
head whitish, latter dusted with gray toward sides. Thorax gray-
ish fuscous dusted with whitish scales. Fore wing with termen
straight; veins 3, 4, and 5 not approximate at termen; color grayish
fuscous with white markings and dusting, making the general color
a rather ashy gray; 14 to 16 paired, evenly spaced, very fine and
obscure white costal geminations; on mid dorsum two obscure gem-
inate white marks; some whitish dusting, continuing from extremi-
ties of these to costa, forms an obscure pale antemedian fascia
defining a dark basal patch, outwardly angulate and somewhat ex-
cavate below middle; post median area grayish fuscous, appearing
to the naked eye as an indistinct dark fascia; ocelloid patch and
apical area dusted with whitish; ocellus defined by two vertical me-
tallic bars enclosing two or three abbreviated black dashes; from
outer white costal geminations two obscure leaden lines extending,
one to inner vertical bar of ocellus, the other to termen below apex;
at apex a round dark spot; cilia leaden gray with a black basal line
peppered with white. Hind wing pale smoky fuscous; cilia paler
with a dark basal band; veins 3 and 4 connate.

Male genitalia figured from paratype in American Museum from
Brownsville, Tex.; female from type in National Collection. The
aedoeagus of the male is very long, slender and sharply bent near

50 BULLETIN 1382, UNITED STATES NATIONAL MUSEUM

middle, scarcely tapering. Signa of female two long slender curved
thorns.

Alar expanse.—12-16 mm.

Type and paratypes.—Cat. No. 28018, USNM; paratypes also
in American Museum and Collection Barnes.

Type locality San Antonio, Tex.

Food plant.—Prosopis.

Described from female type and one paratype from. the type lo-
cality reared from larvae feeding in pods of Mesquite (“ 6-29-17,”
Busck) ; one female paratype from Brownsville, Tex. (“June”) ;
one female paratype from Kerrville, Tex. (“ May, 06”, F. C. Pratt) ;
three male and two female paratypes from Hot Springs, Ariz.
(“from Mesquite, 26-6”) ; three male and one female paratypes from
Baboquivari Mountains, Pima County, Ariz. (“July 15-80, 1903,”
O. G. Poling); one female paratype from Baboquivari Mountains,
Ariz. (O. G. Poling, “15-80 April, 1921”) ; one male paratype from
La Puerta Valley, California (“July, 1911,” Geo. H. Field) ; two
male paratypes from Charlestown Mountains, southern Nev. (“O
G. Poling, 1-15 July, 1921”); and one male paratype from Clark
County, Nev. (“ May 16-23”).

The reared specimens had been in the National Collection for
some time unnamed. Part of the series from the Baboquivari Moun-
tains had been set aside by Kearfott as a new species.

11. LASPEYRESIA MULTILINEANA (Kearfott)
(Fig. 318)
Enarmonia multilineana Kearrotr, Journ. New York Ent. Soe., vol. 16,
1908, p. 178.
Laspeyresia multilineana BARNES and McDunnoucH, Check List Lepid.
Bor. Amer. no. 7250, 1917.—Forsres, Memoir 68, Cornell Univ. Agr.
Expt. Sta., 1924, p. 394.

I have seen no females of this species. The male genitalia are
similar to those of populana Busck; but the two species are quite
different in pattern. Termen of fore wing straight and decidedly
slanting. Hind wing with veins 3 and 4 stalled or connate.

Male genitalia figured from paratype in National Collection from
type locality (“ 26-VI-05,” Criddle).

Distribution—New York, Manitoba.

Alar expanse.—12.5-14 mm.

Type—tIn American Museum.

Type locality —Aweme, Manitoba.

12. LASPEYRESIA INGRATA, new species
(Fig. 315)

Closely resembling multilineana, but with quite different genitalia.
The coastal geminations of fore wing before middle are more dis-

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 51

tinct and, with the white dorsal marks, form an obscure pale fascia
which defines a dark basal patch; the dark postmedian area is also
defined as a rather indistinct fascia; and the apical pair of white
costal geminations are further removed from the others than in
multinineana.

Palpus and face grayish fuscous, slightly dusted with whitish.
Head and thorax fuscous gray; tegula shading to white at tip.
Fore wing fuscous gray; four narrow, rather long, white geminate
marks on middle of dorsum, with a few shorter, fainter white mark-
ings anterior to them on dorsal margin; costa with five pairs of
white dashes, two pair before middle, two beyond, and one near
apex; ocelloid patch consisting of two vertical leaden bars inclosing
two faint, longitudinal black lines; above, a third faint black streak;
termen edged with black: with a white spot below apex and two
or three, more or less fused, white spots at tornus; cilia leaden fus-
cous. Hind wing white toward base, smoky fuscous toward apex;
cilia white with a dark basal band; veins 3 and 4 connate or very
short stalked.

Male genitalia of type figured; aedoeagus stout, bottle necked
toward apex; cornuti, eight short stout fixed spines arranged in a
row.

Alar expanse.—10-13 mm.

Type.—Cat. No. 28019, U.S.N.M.

Paratype.—In collection Barnes.

Type locality. —Aweme, Manitoba.

Described from male type from Aweme, Manitoba (‘‘25-V—05,”
Criddle), and one male paratype from Colfax, Placer County, Calif.
(Vit A, iH. Vachell).

13. LASPEYRESIA ALBIMACULANA (Fernald)
(Fig. 147)

Grapholitha albimaculana FERNALD, Can. Ent., vol. 11, 1879, p. 157.

Enarmonia albimaculana FERNALD, in Dyar List. N. Amer. Lepid., no, 5273,
1903.

Enarmonia articulatana WKerarrorr, Journ. New York Ent. Soc., vol. 16,
1908, p. 177.

Laspeyresia albimaculana BARNES and McDunnovuceH, Check List Lepid.
Bor. Amer., no. 7224, 1917—Forsres, Memoir 68, Cornell Univ. Agr.
Exp. Sta., 1924, p. 395.

Laspeyresia articulatana BaRNEs and McDunnoucH, Check List Lepid.
Bor. Amer., no. 7251, 1917.

An easily recognized species. Fore wing gray with a round, undi-
vided shining white spot on mid dorsum and a conspicuous ae
contrasted black spot on middle of costa. Hind wing with veins
3 and 4 short stalked or connate.

52 BULLETIN 1832, UNITED STATES NATIONAL MUSEUM

Kearfott’s articulatana is an obvious synonym.

Female genitalia figured from specimen in National Collection
from North Evanston, Ill. (A. Kwiat, “ V-18-13”). I have seen no
males.

Distribution.—Maine, Ohio, [linois.

Alar expanse.—11.5-13.5 mm.

Types—Iin National Collection (albtmaculana); in American
Museum (artaculatana).

Type localities—Orono, Me. (albimaculana) ; Cincinnati, Ohio
(articulatana). .

14. LASPEYRESIA POPULANA Busek

(Figs. 145, 319)

Laspeyresia populana BuscKk, Proc. Ent. Soc. Washington, vol. 18, 1916,
p. 151—Barnes and McDunnovucnH, Check List Lepid. Bor. Amer.,
no. 7236-1, 1917.

A somewhat variable but easily recognized species. The genitalia
are similar to those of multilineana; but the pattern is quite different.

Genitalia figured. from paratype (male) and type (female) in
National Collection (reared under Hopk. U.S. no. 12339b, June 15,
1914, from larvae mining cambium of bark of Populus trichocarpa,
J. Brunner, collector).

Distribution.—Montana, Colorado, Manitoba, Alberta. There is
also a male in the Cornell University collection from Ithaca, N. Y.
(“11 July, 1916.”)

Alar expanse.—11.5-14 mm.

Type—tIn National Collection.

Type locality—Missoula, Mont.

Food plant.—Populus trichocarpa (larva in bark).

15. LASPEYRESIA YOUNGANA (Kearfett)

(Figs. 138, 330)

Enormonia youngana KEArRFoTT, Can. Ent., vol. 39, 1907, p. 1.

Laspeyresia youngana Barnes and McDunnoueH, Check List Lepid. Bor.
Amer., no. 7232, 1917.—ForsBes, Memoir 68, Cornell Univ., Agr. Exp.
Sta., 1924, p. 393.

A species of some economic importance as an enemy of spruce.
The larvae feed in the cones upon the seeds and bracts. Rather
widely distributed throughout the northern States and Canada.
There are two generations annually; adults appearing in April-May
and in August-September.

Genitalia figured from paratypes in National Collection from the
type locality.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 53

Distribution.—Ontario, Manitoba, Maine, Colorado, Montana,
Oregon.

Alar expanse.—8-11 mm.

Type.—tin the American Museum.

Type locality.—Ottawa, Canada.

Food plants——Picea alba, P. Sitchensis, Pinus pungens.

16. LASPEYRESIA NIGRICANA (Stephens)
(Figs. 140, 328)

Pseudotomia nigricana STEPHENS, Illus. Brit. Ent., vol. 4, 1884, p. 101.

Semasia nigricana FLtetcHErR, Ontario Exp. Farms Rep. for 1897, 1898, p.
194.—CHITTENDEN, U. S. Dept. Agr. Bull., no. 33, 1902, p. 96; U. S.
Dept. Agr. Buil., no. 66, pt. 7, 1909, p. 95.

Grapholitha nigricana STAUDINGER and REBEL, Cat. Lepid., vol. 2, no.
2160, 1901.

Enarmonia dandana Krarrott, Trans. Amer. Ent. Soc., vol. 33, 1907, p. 65.

Enarmonia ratifera Meyrick, Ent. Mo. Mag., vol. 48, 1912, p. 34.

Laspeyresia dandana Barnes and McDunnovueH, Check List Lepid. Bor.
Amer., no. 7234, 1917.—ForsBes, Memoir 68, Cornell Univ. Agr. Exp.
Sta., 1924, p. 394.

Laspeyresia nigricana BARNES and McDunnouGH, Check List Lepid. Bor.
Amer., no. 7235, 1917.—HeEtnricu, Can. Ent., vol. 55, 1923, p. 13.—
FLuKeE, Bull. 310, Agr. Exp. Sta. Wisconsin, April 1920.—ForsBss,
Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 393.

Laspeyresia novimundi HeEtnricH, Can. Ent., vol. 52, 1920, p. 257.

Endopisa nigricana PirRcE and METCALFE, Genitalia Brit. Tort., 1922, p.
87, pl. 30.

The pea moth of economic literature, an introduced European spe-
cies. Its life history is given in the department and State bulletins
cited above. The type of Kearfott’s dandana is a runted specimen
and smaller than normal examples of nigricana, but in structure and
pattern it agrees. In Europe it has several synonyms. These are
omitted here as they have never appeared in our literature and have
no reference to American localities. The larva feeds in the pods of
garden and field peas and apparently has no other food plant.

Hind wing with veins 3 and 4 stalked.

Genitalia figured from specimens in National Collection from
Sturgeon Bay, Wis., reared July 14, 1920, by C. L. Fluke.

Distribution.—New Jersey, Wisconsin, Washington, Manitoba, Al-
berta, Ontario, Nova Scotia.

Alar expanse.—12-14 mm.

Types.—In British Museum (nigricana); American Museum
(dandana) ; National Collection (novimundz).

Type localities —England (nigricana) ; Essex County Park, N. J.
(dandana) ; Sturgeon Bay, Wis. (novimundi).

Food plant.—Pisum.

54 BULLETIN 132, UNITED STATES NATIONAL MUSEUM
17. LASPEYRESIA CANDANA Forbes
(Figs. 152, 325)

Laspeyresia candana Forbes, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 394.

A brownish fuscous species with a paler brownish semilustrous
median fascia and concolorous brown hind wings; veins 3 and 4 of
hind wing stalked.

Forbes validated the Kearfott manuscript name; but did not desig-
nate types. I therefore do so here, restricting them to a series of 12
males and 2 females from Oak Station, Pa. (May 15 to 21, F. Mar-
loff) which Kearfott had deposited under the name in the three col-
lections.

In addition to these we have in the National Collection a male
from Guy’s Mills, Pa. (May 18, 1915, Heinrich), and a female from
Cincinnati, Ohio. (“ [V—23-04,” A. F. Braun).

Genitalia figured from type in American Museum (male) and
paratype in National Collection (female). Aedoeagus stout; cornuti
two parallel rows of short fixed spines.

Alar expanse —12-16 mm.

Type—tn American Museum.

Paratypes—Cat. No. 28020, U.S.N.M. Also in American Museum
and collection Barnes.

Type locality—Oak Station, Pa.

I refer the species to Laspeyresia with some hesitation. In pattern
it is very close to nigricana, grandicula, and caryana. The male
genitalia are most like those of Carpocapsa pomonella. It also
shares a male character with Carpocapsa and Melissopus in the ap-
pressed pocket inclosing the median pecten on hind wing. The rough
sex scaling on the inner margin of the hind wing also occurs in
caryana and nigricana and in the species under Melissopus. Eventu-
ally it will probably have to have a separate generic designation;
but at present I do not think this is justified.

18. LASPEYRESIA GRANDICULA, new species
(Fig. 320)

Similar to candana in pattern and color of fore wing; but some-
what darker and lacking the fine peppering of whitish scales, and
with very different genitalia.

Antenna of male nearly smooth. Palpus sordid whitish; third
joint dusky. Head brown. Thorax and fore wing semilustrous fus-
cous brown; an obscure angulate median fascia on fore wing, slightly
paler than ground color and broadest toward dorsum; pale costal
geminations indistinct, except four evenly spaced whitish dashes on

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 55

outer half of costa; ocelloid patch as in candana, cilia leaded fuscous
somewhat shaded with paler scaling at tornus and with a black basal
line bordering termen. Hind wing with dorsal margin concave;
blackish brown, much darker than fore wing and (in male) with a
large patch of jet black appressed sex scaling on disk; no sex scaling
on inner margin; hair tuft on lower median vein and at base of vein
1*, snow white; cilia shining snow white, without dark basal line;
veins 3 and 4 connate. Underside of fore and hind wings very dark
brown; a large patch of appressed black scales covering cell of fore
wing in male.

Male genitalia of type figured; aedoeagus (omitted from figure)
moderately stout, tapering; cornuti very minute, scarcely distin-
guishable.

Alar expanse.—16 mm.

Type—Cat. No. 28021, U.S.N.M

Type locality —Moutain Lake, Va.

Described from male type (“June 14-21, 1907,” A. F. Braun).

A striking species easily recognized by its dark brown hind wing,
snow-white hind wing cilia, peculiar male sex scaling and character-
istic genitalia.

19. LASPEYRESIA CARYANA (Fitch)
(Figs. 146, 325)

Ephippiphora caryana Fircu, Third Rep. Ins. New York, 1856, p. 459.

Grapholitha caryae SHIMER, Trans. Amer. Ent. Soc., vol. 2, 1869, p. 394.

Enarmonia caryana FERNALD, in Dyar List N. Amer. Lepid., no. 5268,
1903.—BarRNES and McDonnoueH, Check List Lepid. Bor. Amer., no.
7207, 1917.—ForsBEs, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 392.

A rather common insect in the eastern United States. The larvae
feed in the husks and fruits of hickory and pecan nuts and some-
times do considerable damage. The moth resembles that of candana
but is easily distinguished. The hind wing is whitish on upper
basal half; and in the male there is considerable black scaling in the
area occupied by the anal veins, on the dorsum of the first three
abdominal segments and on the under surface of both fore and hind
wings; the thick white male sex scaling on inner margin of the hind
wing is also pronounced. Hind wing with veins 3 and 4 connate.

Genitalia figured from reared specimens in National Collection
from District of Columbia (“1035 P, 16—July—94,” male) and Cadet,
Mo. (467603, July, 2-90,” female). Aedoeagus slender, tapering
and forked (that is, having a lateral spur, as in some specimens of
the genus Melissopus) ; cornuti not distinguishable.

Distribution—New Jersey, Pennsylvania, District of Columbia,
Maryland, Virginia, Georgia, Florida, Texas, Missouri.

Alar expanse —9-15 mm.

56 BULLETIN 182, UNITED STATES NATIONAL MUSEUM

Types—In National Collection (caryana); location unknown
(caryae).

Type locality—New York (caryana and caryae).

Food plants.—Hickory, pecan.

20. LASPEYRESIA FLETCHERANA (Kearfott)
(Fig. 321)

Enarmonia fletcherana Kearrort, Can. Ent., vol. 39, 1907, p. 127.
Laspeyresia fletcherana BARNES and McDuNNoucuH, Check List Lepid. Bor.
Amer., no. 7246, 1917.—Fornes, Memoir 68, Cornell Univ. Agri. Exp.

Sta., 1924, p. 395.

A strikingly marked species easily identified by pattern and geni-
talia. Hind wing with veins 3 and 4 short stalked. Aedoeagus long,
slender; cornuti three clusters of short fixed spines.

Male genitalia figured from paratype in National Collection from
the type locality (‘“18-VI-1905,” C. H. Young). I have seen no
females.

Specimens in National and Canadian National Collections, Ameri-
can Museum and collection Barnes from Ottawa, Canada.

Alar expanse.—12-14 mm.

Type.—In American Museum.

Type locality —Ottawa, Canada.

21. LASPEYRESIA TANA (Kearfott)
(Figs. 156, 327)
Enarmonia tana Kearrort, Trans. Amer. Ent. Soc., vol. 33, 1907, p. 62.
Enarmonia cirrhas Meyrick, Ent. Mo. Mag., vol. 48, 1912, p. 34.
Laspeyresia tana BaRNES and McDuNNovuGH, Check List Lepid Bor. Amer.,
no. 7248, 1917.

Another striking species. Hind wing with veins 3 and 4 stalked.
Aedoeagus of male stout, nearly straight; cornuti a cluster of 6 to 8
stout, moderately long, fixed spines and a half a dozen slender,
longer, deciduous spines.

Genitalia figured from paratypes in National Collection from the
type locality (A. H. Vachell, “ VII”).

Distribution.—California, Nevada.

Alar expanse.—12-15 mm.

Type.—tIn American Museum.

Type locality —Colfax, Placer County, Calif.

22. LASPEYRESIA CUPRESSANA (Kearfott)
(Figs. 154, 326)

Cydia cupressana Krarrott, Trans. Amer. Ent. Soe., vol. 338, 1907, p. 54.
Carpocapsa cupressana BARNES and McDunnovueH, Check List Lepid. Bor.

Amer., no. 7275, 1917.
Similar to the preceding (¢ana) in genitalia and pattern: but ap-
parently distinct. The genitalia (male and female) are smaller, and
the ocelloid patch and antemedian fascia of fore wing much paler.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 57

Hind wing with veins 3 and 4 connate or short stalked.

Genitalia figured from reared specimens in National Collection
from the type locality (Jan. 11 and 24, 1886).

Specimens in National Museum, American Museum, and collection
Barnes from California.

Alar expanse.—12-15 mm.

Type.—tIn American Museum.

Type locality —Alameda County, Calif.

Food plant—Cupressus macrocarpa (larvae feeding upon the
seeds).

23. LASPEYRESIA PROSPERANA (Kearfott)

(Figs. 144, 324)

Grapholitha succedana WALSINGHAM (not Schiffermiiller), Illus. Lepid.
Heter. Brit. Mus., vol. 4, 1879, p. 97.

Thiodia succedana FERNALD, in Dyar List. N. Amer. Lepid., no. 5169, 1903.

Enarmonia prosperana Kerarrort, Can. Ent., vol. 39, 1907, p. 128.

Eucosma succedana Barnes and McDuNnnoueH, Check List Lepid. Bor.
Amer., no. 7088, 1917.

Laspeyresia prosperana BaRNES and McDuNnNouGH, Check List Lepid. Bor.
Amer., no. 7247, 1917.

What Walsingham determined as the European succedana from
Oregon is undoubtedly what Kearfott described as prosperana. The
two are very similar in pattern and genitalia; but appear to be
distinct. Our American form must at the very least be kept as a
separate race. It has the costa of the harpe much more strongly
curved than has succedana, and the cucullus differently shaped.
Each has a short prong arising from the side of the aedoeagus
near middle, and the organ slender, tapering and decidedly curved.

Genitalia figured from specimens in National Collection from
Field Brook, Calif. (H. S. Barber, “19-05-03,” male), and Easton,
Wash. (female paratype).

Distribution—Colorado, Utah, California, Washington, Alberta,
British Columbia, Alaska.

Alar expanse.—13-18 mm.

Type.—In American Museum,

Type locality —San Luis Obispo, Calif.

24. LASPEYRESIA LEUCOBASIS Busck
(Figs. 157, 332)

Laspeyresia lteucobasis Busck, Proc. Ent. Soc. Washington, vol. 18, 1916,
p. 152.—Barnes and McDunnoueH, Check List Lepid. Bor. Amer.,

no. 7248-1, 1917.
Similar to prosperana but distinguished by genitalia and the char-
acters given in our key., Hind. wing with veins 3 and 4 stalked.

58 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Aedoeagus long, tapering, moderately stout; cornuti a row of a dozen
short slender spines.

Genitalia figured from specimens in National Collection from
Missoula, Mont. (reared June 1, 1915, under Hopk. U. S. no. 11082,
from larvae in bark of Picea engelmanni, B. 'T. Harvey). In nature
leucobasis feeds and flies in company with both rana and laricana.

Specimens in National Collection from Montana.

Alar expanse.—12-14 mm.

Type.—tin National Collection.

Type locality.—Evaro, Mont.

Food plants—Larix occidentalis, Picea engelmanni.

25. LASPEYRESIA GALLAESALICIANA (Riley)
(Figs. 166, 331)

Grapholitha gallaesaliciana RiLey, Trans. St. Louis Acad. Sci., vol. 4, 1881,
p. 320.

Enarmonia gallaesaliciana FERNALD, in Dyar List N. Amer. Lepid., no.
5280, 1903.

Laspeyresia gallaesaliciana BARNES and McDuNNovuGH, Check List of Lepid.
Bor. Amer., no. 7236, 1917.—Forses, Memoir 68, Cornell Univ. Agr.
Exp. Sta., 1924, p. 395.

The palest of the Lespeyresia (except lautiuwscula), and a very
beautiful species. The larvae are gall makers on stems of willow.
Hind wing with veins 3 and 4 short stalked. Aedoeagus slender,
tapering to a narrow band toward apex; cornuti a single cluster
of short rather slender fixed spines.

Genitalia figured from reared specimens in National Collection
from Boston, Mass. (May 6-8, 1910, Miss Clarke).

Distribution.—Massachusetts, New Jersey, Pennsylvania, Illinois,
Missouri.

Alar expanse.—11-13 mm.

Type.—tin National Collection.

Type locality —St. Louis, Mo.

Food plant.—Salizx.

26. LASPEYRESIA LAUTIUSCULA, new species

(Fig. 168)

Antenna dusted with white above. Palpus, face, and head white.
Thorax black marked with white; posterior part of thorax and al-
most all the tegula except anterior margin, white. Fore wing white
with apical area blackish and an incomplete dark basal patch; basal
patch lead colored, rather faint, reaching only to top of cell from
dorsum; apical dark area triangular, inner margin straight and
slanting, extending from outer third of dorsum to apex; costa very
faintly marked beyond middle with outwardly slanting, pale, smoky

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 59

dashes extending as far as top of cell; some blue metallic markings
just below apex and a couple of similarly colored vertical bars above
tornus; cilia dark smoky fuscous with a black basal band. Hind
wing smoky fuscous, white toward base; under surface white with
some fuscous mottling toward apex; cilia whitish with a dark basal
- band from apex to vein 1b; veins 3 and 4 stalked.

Female genitalia of type figured; ductus bursae and neck of bursa
copulatrix strongly chitinized.

Alar expanse.—15 mm.

Type.—In collection Blackmore.

Type locality—Fraser Mills, British Columbia.

‘Described from unique female type (L. E. Marmont, “ 27—V I-22,”
Blackmore No. 58). Similar in color and pattern to gallaesaliciana
Riley and distinguished from that species chiefly by its larger size
and the more heavily chitinized ductus and bursa of its genitalia.

27. LASPEYRESIA? FLEXILOQUA, new species

(Fig. 142)

Antenna sordid white above. Palpus, face, head, and thorax
whitish gray. Fore wing white with blackish markings; an out-
wardly angulate black basal patch, much broken by white but with
the outer margin complete from costa to dorsum; from middle of
costa to cell near upper outer angle, a narrow black band; costa
otherwise finely strigulated with black; a black shade filling termi-
nal area from dorsum at outer third to apex, except for a white
dusting at tornus; the inner margin of this black area is somewhat
irregular and it contains a couple of vertical blue metallic bars above
tornus and two similarly colored streaks below apex; on median
white area, especially toward dorsum, some fine dusting or streaks
of black; cilia leaden fuscous, with a black basal line broken by
a small white spot below apex. Hind wing pale smoky fuscous,
darkest toward apex; cilia paler with a dark basal band; under
surface of hind wing concolorous with cilia; veins 3 and 4 very short
stalked.

Female genitalia of type figured.

Alar expanse.—16 mm.

Type.—tIn Canadian National Collection.

Type locality —Calgary, Alberta.

Described from female type (“6-VII-21”). <A rather striking
form, somewhat similar to dautiuscula but. with different genitalia,
complete basal patch and more black dusting on the white areas.
The generic reference is tentative. It may be that the species should
go in Hemimene (the genitalia seems to suggest it): but, in absence
of a male, this can not be definitely determined.

54346—26——_5

60 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

28. LASPEYRESIA AMERICANA (Walsingham)

(Figs. 158, 336)

Grapholitha americana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol.
4, 1879, p. 67.
Enarmonia americana FERNALD, in Dyar List N. Amer. Lepid., no. 5276,
1903 ; not Kearfott, Proc. U. 8. Nat. Mus., vol. 28, 1905, p. 362.
Laspeyresia americana BARNES and McDunnoucH, Check List Lepid. Bor.
Amer., no. 7244, 1917.

A large Pacific coast species superficially resembling 2’ piblema in-
feléa Heinrich. May be readily recognized by its genitalia and the
large clear white dorsal spot upon an otherwise dark purplish fus-
cous fore wing. Hind wing with veins 3 and 4 short stalked. Aedo-
eagus extremely stout at base and tapering evenly to a very narrow
apex; cornuti a longitudinal row of a half a dozen short stout fixed
spines.

Genitalia figured from specimens in National Collection from
Colfax, Calif. (A. H. Vachell, “V-1”).

Distribution.—California, Washington, British Columbia.

Alar expanse.—13.5-19 mm.

Type.—tn British Museum.

Type locality—Mendocino County, Calif.

29. LASPEYRESIA FLAVICOLLIS (Walsingham)

(Fig. 149)

Cydia? flavicollis WALSINGHAM, Proc. Zool. Soc. London, 1897, p. 130.

There is a perfect female of this beautiful species in the Kearfott
collection at the American Museum. It is labeled “ Everglades,
Florida, April 8-15.” This is our first North American record.

Hind wing with veins 3 and 4 stalked.

Genitalia figured from specimen in American Museum.

Alar expanse—15 mm.

Type——In “Museum Hedemann” (?).

Tupe locality—St. Thomas, British West Indies.

30. LASPEYRESIA NINANA (Dyar)

(Figs. 148, 385)

Carpocapsa ninana RiLry, in Smith’s List Lep. Bor. Amer. no. 5025,
1891.—BarNES and McDunnovucH, Check List Lepid. Bor. Amer., no.
(273, 1917.

Cydia ninana Dyar, List N. Amer. Lepid., no. 5298-1, p. 471.

Although this species is attributed to Riley in the Smith list, the
first description is by Dyar and it must therefore be credited to him.
Like flavicollis, it is probably of tropical origin. Both species may
eventually have to have a different generic designation; but at

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 6]

present we have no characters to justify their separation from
Laspeyresia.

In this and the following five species there is a slight fold along
vein 1° in the male hind wing. On this character we might put
them all in Carpocapsa,; but such an arrangement is not justified by

the genitalia. I am therefore restricting that genus to its type
(pomonella).

Hind wing with veins 3 and 4 stalked.

Genitalia figured from paratypes in National Collection. Aedoe-
agus long, slender, curved, tapering; cornuti a few (3-6) slender,
minute spines.

Specimens in National Collection and American Museum from the
type locality.

Alar expanse—17-19 mm.

Type.—In National Collection.

Type locality.—Arizona.

31. LASPEYRESIA COLORANA (Kearfott)
(Figs. 167, 337)

Cydia colorana Kerarrott, Trans. Amer. Dnt. Soc., vol. 33, 1907, p. 93.
Carpocapsa colorana BARNES and McDuNNovucH, Check List Lepid. Bor.
Amer., no. 7277, 1917.

A striking form with unusual genitalia. In pattern it reminds of
Melissopus, but is considerably paler. Hind wing with veins 3 and
4 stalked.

Aedoeagus stout; cornuti a single cluster of 8 rather short stout
fixed spines.

Genitalia figured from type in Amevyican Museum (male) and
paratype in National Collection (female); latter from Glenwood
Springs, Colo. (“ June-24-30”).

Specimens in National Collection, American Museum, and collec-
tion Barnes from Colorado.

Alar eapanse —22-24 mm.

Type—tIn American Museum.

Type locality.—Salida, Colo.

32. LASPEYRESIA EROTELLA (Heinrich)
(Figs. 141, 338)

Carpocapsa erotella HeInRricH, Proc. Ent. Soc. Washington, vol. 25, 1923,

p. 121.—ForsBes, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 396.

A small easily recognized species. Hind wing with veins 3 and +

stalked. Aedoeagus stout, slightly tapering; conutus a single thin,
moderately long, fixed spine.

62 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Genitalia figured from type (female) and paratype (male) in
National Collection.

In addition to the type and paratypes in the National Collection
there is a paratype from Biloxi, Miss., in the Cornell University col-
lection; also a female of what I take to be a possible western variety
from Patrick’s Creek, Calif. (reared Sept. 14, 1916, under Hopk. U.
S. No. 142894 from Pinus atenwata, J. E. Patterson). This last is
in the National Collection. In pattern and genitalia it agrees with
typical erotella except that the ocelloid patch of fore wing is more
heavily streaked with black. I doubt if it will prove sufficiently dis-
tinct to deserve a varietal name.

Alar expanse.—9-10 mm.

Type.—tn National Collection.

Type locality —Hyattsville, Md.

Food plant—Pinus taeda.

33. LASPEYRESIA TOREUTA (Grote)
(Fig. 159)
Penthina toreuta Grotr, Bull. Buffalo Soe. Nat. Sci., vol. 1, 1873, p. 92.
Cydia toreuta FERNALD, in Dyar List N. Amer. Lepid., no. 5298, 1903.
Carpocapsa toreuta Kearrortt, Proc. U. 8S. Nat. Mus., vol. 28, 1905, p. 362.-—
BaRNES and McDuNnnovucH, Check List Lepid. Bor. Amer., no. 7272,
1917.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 396.

A moderately sized species quite different in pattern and color
from everything in the genus except ingens and piperana. From
these two it is readily separable on genitalia (particularly those of
the female) and distribution. The true ¢oreuta is limited to the
eastern United States above Florida. It is somewhat smaller than
piperana and ingens and has smaller male genitalia and proportion-
ately smaller aedoeagus; the neck of the harpe is more incurvate than
that of péperana and not so long as that of trgens.. Hind wing with
veins 3 and 4 connate or very short stalked.

_Female genitalia figured from specimen in National Collection
from Falls Church, Virginia (reared Aug. 13, 1919 under Hopk. U.
S. No. 12033 ¢ from larva feeding in cones of P. virginiana, Wm. Mid-
dleton).

Distribution —Texas, North Carolina, Virginia, District of Col-
umbia, Pennsylvania.

Alar expansé.—13-15 mm.

Type—tIn National Collection.

Type locality—Pennsylvania.

Food plant—Pinus virginiana (larvae feeding in the cones).

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 63

34. LASPEYRESIA INGENS, new species
(Fig. 161)

Similar to toreuta and piperana but with different genitalia and
distribution; larger than the former.

Palpus sordid whitish; apical joint fuscous beneath. Head sor-
did white shaded with fuscous. Fore wing ashy brown, under
magnification showing the scales beyond base dark brown tipped
with sordid white; basal area glossy; from costa just before middle
to mid dorsum a narrow, slightly angulate metallic bar edged in-
wardly and outwardly with black; a similar, somewhat more angu-
late bar from costa beyond middle to tornus, broken between veins
6 and 8; another similar bar extending along termen to just below
apex, thence curving upward and inward to a white costal spot near
apex; between this and the preceeding bar two obscure white costal
marks terminating in metallic droplets; extreme terminal margin
black; cilia leaden fuscous. Hind wing smoky fuscous; cilia paler
with a dark basal band; veins 3 and 4 short stalked.

Female genitalia of type figured. Male genitalia as in piperana;
but neck of harpe considerably more (nearly twice as much) ex-
tended and with an irregular rounded projection from the arch;
cucullus more irregular in outline, slightly concave on lower margin
and below apex; adeoeagus a third longer.

Alar expanse.—17-20 mm.

Type.—tin collection Barnes.

Paraty pe-——Cat. No. 28022, U.S.N.M. Also in American Museum
and collection Barnes.

Type locality—St. Petersburg, Fla.

Described from female type, one male and two female paratypes
all from the type locality.

35. LASPEYRESIA PIPERANA (Kearfott)
(Figs. 163, 333)

Cydia piperana Kerarrott, Trans. Amer. Ent. Soc., vol. 33. 1907, p. 55.
Carpocapsa piperana BARNES and McDuNNowuGH, Check List Lepid. Bor.
Amer., no 7274, 1917.

Very similar to toreuta Grote; but with different genitalia and
limited to the western (Rocky Mountain and Pacific Coast) States,
where it is a species of considerable economic importance. The larvae
feed in pine cones. The favorite food plant seems to be P. ponderosa.
The species has the same life cycle and feeds and flies with Hedulia
injectiva, with which it also seems to hybridize (see p. 66).

64 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Genitalia figured from specimens in American Museum (male
type) and National Collection (female paratype from the type lo-
cality (C. V. Piper, “11 Mch—78”)).

Distribution.—Idaho, Montana, Washington, Oregon, California.
Represented by large reared series in the National Museum.

Alar expanse.—16-20 mm.

Type.—In American Museum.

Type locality —Pullman, Wash.

Food plant.—Pinus ponderosa, P. jeffrey.

36. LASPEYRESIA MISCITATA, new species
(Fig. 160)

An intermediate between Z. piperana Kearfott and Wedulia in-
jectiva and partaking of the characters of both species. It lacks en-
tirely the hairy vestiture of the latter. Both otherwise resembles it
very closely ; having blackish palpi and similarly colored hind wings.
The head is black as in ¢énjectiva but with a pepering of white at the
scale ends. The fore wing is glossy at base as in piperana, but gen-
erally darker (in most specimens) and colored more as in énjectiva;
at tornus on inner margin of the terminal metallic bar there is more
black scaling than in either of the other two species, this is some-
times extended into irregular streaks and gives the appearance of an
ocelloid patch.

Genitalia as in énjectiva. In the female the genital plate varies
somewhat in size and shape in different specimens; in some it is_
quite like that of znjectiva, in others narrower and smaller; the ex-
treme form is shown in figure 160 (paratype from the type locality).

Alar expanse.—11.5-17 mm.

Type.—Cat. No. 28023, U.S.N.M.

Paratypes.—In National Collection, American Museum, Canadian
National and Barnes collections.

Type locality.—Shasta National Forest, Calif.

Food plants—Pinus ponderosa, P. jeffrey.

Described from male type and 4 female paratypes from the type
locality (reared under Hopk. U.S. No. 11414, May, 1912, from larvae
feeding in cones of P. jeffreyi, J. M. Miller); 2 male and 2 female
paratypes from Ashland, Oreg. (reared May 11, June 24, and Aug.
26, 1914, under Hopk. U. S. Nos. 10876 and 12539 A, from P. pon-
derosa, P. D. Sargent); 2 male and 1 female paratypes from Fel-
ton, Calif. (reared Dec. 30, 1916 under Hopk. U. S. No. 11312a
from P. ponderosa, P. D. Sargent) ; one female paratype from Bray,
Calif. (reared September 18, 1915, under Hopk. U.S. No. 14298a from
P. ponderosa, J. D. Riggs); and one female paratype from Verdi,
Nev. (“June 1-10,” A. H. Vachell); this last had been included

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 65

by Kearfott as one of the paratypes of his manuscript species,
“Enarmoma bandana” (see remarks under Grapholitha caeruleana,
p. 31). The above-reared specimens selected from a large series from
cones also infested with either LZ. piperana or H. injectiva.

I dislike very much to name this form, as I am strongly of the
opinion that it represents a hybrid of piperana and injectiva; but as
this has not as yet been demonstrated by rearings from known par-
ents, there is nothing else to do.

12. HEDULIA, new genus
(Figs. 162, 334)

Genotype—Hedulia injectiva, new species.

Characters as in Grapholitha except:

Head, palpi, legs, and underside of thorax and abdomen covered
with fine rather long hair.

Fore wing with some fine hair-like scales toward base; termen
convex.

Hind wing with veins 3 and 4 stalked; inner margin hairy.

Abdomen of male simple.

A derivative of the toreuta group of Laspeyresia. The hairy
vestiture is unique in the family and reminds of Symmnona in the
Tortricidae. Contains only the one North American species.

HEDULIA INJECTIVA, new species
(Figs. 162, 334)

Similar in pattern to Laspeyresia piperana, but darker, with white
hind wing cilia and bicolored hind wings. Distinguished at once by
the hairy vestiture.

Antenna, palpus, face, head, and thorax black; inner side of pal-
pus sordid whitish ocherous. Fore wing blackish fuscous, the ends
of the scales tipped with sordid ocherous, giving a dark wood brown
color to the naked eye; basal area not appreciably glossy; a some-
what irregular and rather obscure metallic band from costa before
middle to mid dorsum and widening out a trifle toward dorsum; a
similar, but more regular and angulate band from costa beyond
middle to tornus, in some specimens broken near middle; another
similar bar (or band) along termen to below apex; extreme costal
margin whitish ocherous, this pale scaling produced beyond middle
into from 4 to 6 geminate marks, from the apical one of which a
leaden band continues toward termen terminating close to upper end
of terminal metallic bar; extreme terminal margin black; cilia
leaden fuscous. Hind wing smoky blackish fuscous, shading to sor-
did whitish toward base; cilia white with a dark basal band. Under
side of fore wing with a slight dusting of white near termen, and

66 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

with pale costal markings somewhat more distinct than on upper
surface; underside of hind wing blackish, dusted with whitish on
disk and toward apex. Abdomen black above. Hairy vestiture of
abdomen and legs mixed black and grayish ocherous.

Genitalia figured from type and paratype from the type locality.

Alar expanse.—16-20 mm.

Type and paraty pes.—Cat. No. 28024, U. S.N.M. Paratypes also in
American Museum, Canadian National and Barnes collections.

Type locality —Reno, Nev.

Food plant.—Pinus.

Described from male type; 4 male and 1 female paratypes from
the type locality (labeled: “In pine cones, issued 1-24-11, J. B.
Smith”); 1 male and 2 female paratypes from Ureka, Calif. (reared
under Hopk. U. S. No. 11413 and 10889a, February 10-16, 1912,
and March 2, 1914, from larvae in cones of Pinus jeffreyi, J. M.
Miller) ; 2 male paratypes from Pine Valley, Calif. (reared under
Hopk. U. S. No. 13276, August 27, 1915, from cones of P. jeffrey?,
F. P. Keen) ; 1 female paratype from Mona National Forest, Calif.
(Hopk. U. S. No. 12557a?, issued February 22, 1915, from cones of
P. jeffreyz) ; and 1 male paratype from Ashland, Oreg. (Hopk. U. S.
No. 12539 hi, issued August 26, 1914, from cones of P. ponderosa,
P. D. Sergent). These are from a large series reared by the Forest
Insect Division of the Bureau of Entomology. In addition to the
above, we have reared specimens from several other California
localities and two examples (a male and female) labeled “ From
pine cones, North Carolina.”

This species occurs in the same localities and has the same life
history and habits as piperana Kearfott, and seems to be even more
abundant and destructive than the latter. Its larvae feed in pine
cones. The favorite food plant appears to be P. jeffreyi, though
cones of P. ponderosa and other pines are occasionally attacked. It
seems to hybridize with L. piperana (see p. 65).

13. Genus MELISSOPUS Riley
(Figs. 2, 9, 36, 113)
Melissopus Riitry, Trans. St. Louis Acad. Sci., vol. 4, 1881, p. 322.
Genotype.—Carpocapsa latiferreana Walsingham (North America).
Thorax smooth.
Fore wing smooth; termen slightly concave; 12 veins all separate;
7 to termen; 11 from cell just before middle; 10 remote from 9;
upper internal vein of cell from between 10-11; 3, 4, and 5 somewhat
approximate at termen; 2 from cell before 24, straight; no costal
fold in male.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 67

Hind wing with pecten from lower median vein, in male this is
developed as a narrow, long hair pencil concealed in a deep, semi-
elliptical, closely appressed pocket lying along basal half of vein
2; 8 veins; 6 and 7 approximate toward base; 3 and 4 stalked; 2 from
_ cell before middle in male, normal in female; inner margin (in male)

developed into a shallow pocket filled with broadly spatulate white
scales above, and rough scaling beneath.

Hind tibia and basal joint of hind tarsi of male dilated and clothed
with dense, latterally appressed scale tufts above and below.

Male genitalia with harpe simple; outer surface unspined; cucul-
lus well defined, evenly and heavily spined; neck incurvation slight;
neck smooth except at base of cucullus; sacculus simple, rather
densely clothed with fine, short, hairlike spines. Tegmen elongate;
inner posterior margins scobinate; posterior lateral extremities some-
times produced into hornlike projections resembling a_ widely
bifurecate uncus. Uncus absent. Socii absent. (Gnathos scarcely
defined, very weakly chitinized. Aedoeagus straight, tapering
sharply before middle and continued as a very slender tube; simple
or with a lateral spur from near middle; cornuti absent.

Abdomen of male simple.

Female genitalha with two thornlike signa. Ductus bursae short,
unchitinized. Bursa copulatrix with neck evenly and _ strongly
granulate.

A monotypic genus derived from the toreuta group of Laspeyresia.

MELISSOPUS LATIFERREANUS (Walsingham)
(Figs. 2, 9, 31, 32, 33, 34, 35, 36, 112, 113)

Carpocapsa latiferreana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol.
4, 1879, p. 70.

Melissopus aurichalceana Ritry, Trans. St. Louis Acad. Sci., vol. 4, 1881,
p: 323.

Melissopus latiferreanus FERNALD, in Dyar List N. Amer. Lepid., no. 5295,
1903. Barnes and McDunnovueH, Check List Lepid. Bor. Amer., no.
7269, 1917—Forsrs, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 397.

Cydia inquilina KEarrort, Trans. Amer. Ent. Soc., vol. 33, 1907, p. 55.

Carpocapsa inquilinad BaRNES and McDuNnnoueH, Check List Lepid. Bor.
Amer., no. 7276, 1917.

An extremely variable species in color, size, and structure, and
one which seems to be in the process of breaking up into several
races or even species. In extreme forms (A and G) the male geni-
talia differences are very striking, but they do not correspond with
either size or color differences and in large series from different
localities there are so many intergrades that division into clearly
definable races is impossible. I list below the forms that I have seen.

54346—26—_6

68 BULLETIN 1382, UNITED STATES NATIONAL MUSEUM

Further material will undoubtedly show others. It is possible that
we had originally two species, an Eastern (var. G) and a Western
(var. A) which have interbred to produce the intermediate forms;
but this is a mere guess. Careful and extensive rearings from known
parents will be necessary before we can determine the exact status
of any of the varieties. Until we have such, it would be very unwise
to make any new names or attempt any splitting of the species.

Var. A (fig. 36)—Tegumen of male genitalia without posterior
projecting horns; aedoeagus with a long lateral spur reaching from
below middle almost to apex; apex of spur bent over, broadened and
serrate. Color ranging from pale reddish brown to rosy gray brown.
Expanse: 13-20 mm. Distribution: California, Oregon, and Wash-
ington. Reared specimens from acorns.

This is the typical latiferreanus. Kearfott’s inquilina also goes
here.

Var. B (fig. 32) —Tegumen without posterior horns; aedoeagus
with rather long spur, latter, however, not reaching to apex of
aedoeagus, smooth and pointed. Color as in variety A, but most
specimens rather pale. Expanse: 18-20 mm. Distribution: Utah.
None of specimens in collections reared.

Var. C (fig. 33)—Tegumen developed posteriorly into two stout,
well separated, rather long, slightly roughened, partially curved
hornlike projections; aedoeagus simple. Color as in varieties A and
B. Expanse: 13-20 mm. Distribution: California, Arizona, New
Mexico. Reared specimens from acorns.

Var. D (fig. 34) —Tegumen developed posteriorly into two very
short hornlike projections; aedoeagus with very short, pointed,
lateral spur. Color, very dark red-brown; hind wing Bede brown
with strongly contrasted white cilia; a rather strong black dusting
on forewing between the metallic cross bars. Expanse: 15-20 mm.
Distribution : Pennsylvania, Virginia, Connecticut. The only reared
specimens are a couple in the National Collection (Hopk. U. S.
No. 12106 Falls Church, Va., July 10 and 25, 1914, Heinrich) bred
from chestnut husks.

The most distinct form so far as color is concerned. There is a
pale Nevada form, however, with similar genitalia.

Var. .—Tegumen as in variety A; aedoeagus as in variety B.
Color, reddish brown. Expanse: 11-13 mm. Distribution: West
Virginia, Pennsylvania, Illinois. Reared specimens from beech nuts
(Quaintance No. 7590, French Creek, West Virginia, F. E. Brooks).
Collected adults from Pennsylvania and Illinois intergrade between
this and the following variety and are difficult to place.

Var. F (fig. 35).—Genitalia as in variety / except that posterior
projections of tegumen are straight and longer. Color yellow or

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 69

yellowish red. Expanse: 16-17 mm. Distribution: Missouri, Texas,
North Carolina, Pennsylvania. Reared specimens from acorns.

Riley’s type of aurichalceana goes here.

Var. G (fig. 31)—Tegumen with long, straight, roughened, horn-
like posterior projections; aedoeagus simple. Color reddish brown
or reddish ocherous. Expanse: 15-17 mm. Distribution: Penn-
sylvania, Missouri, Florida, Mexico. Reared specimens from acorns.

In all these forms the female genitalia also show some slight vari-
ations. Varieties A (fig. 113), and # (fig. 112) exhibit the extreme
differences. Between these there is a gradual intergradation in the
other varieties.

Alar expanse.—11-20 mm.

Types.—In British Museum (latiferreanus) ; National Collection
(aurichalceana) ; American Museum (inquilina).

Tipe localities —Medocino County, Calif. (latiferreanus) ; Kirk-
wood, Mo. (aurichalceana) ; San Francisco, Calif. (inguilina).

Food plants—Oak acorns, beech nuts, chestnut burs.

14. Genus CARPOCAPSA Treitschke
(Figs. 37, 169)

Carpocapsa TREITSCHKE, Schmet. Eur., vol. 8, 1830, p. 160 (=Cydia Wal-
singham not Hiibner).
Genotype.—Phalaena Tinea pomonella Linnaeus (Europe).

Thorax smooth.

Fore wing smooth; termen straight; 12 veins, all separate; 7 to
termen; 11 from cell slightly beyond middle; 9 and 10 well sepa-
rated; upper internal vein of cell from between 10-11; 3, 4, and 5
nearly parallel beyond cell, not approximate at termen; 2 from cell
before 24, straight; no costal fold in male.

Hind wing with pecten from lower median vein, in male de-
veloped as a long slender black hair pencile inclosed in a shallow
pocket along vein ic; 8 veins; 6 and 7 approximate toward base,
3 and 4 stalked; 2 form cell at outer 44 in male; inner margin in
male simple.

Hind tibia of male smooth scaled.

Male genitalia with outer surface of harpe unspined; cucullus
well defined, finely and evenly spined; neck incurvation shght; a
short, thornlike projection from neck near cucullus (a similar de-
velopment is also found in a few species of Laspeyresia), otherwise
smooth; sacullus weakly spined. Tegumen a rather narrow chiti-
nized band. Uncus absent. Socii absent. Gnathos a simple weakly
chitinized band. Aedoeagus moderately long, stout, slightly taper-
ing; cornuti a cluster of short, stout fixed spines (6-8).

Abdomen of male simple.

70 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Female genitalia with two thorn-like signa. Ductus bursae short,
strongly chitinized and scobinate.

As it is now defined this is not a good genus. The only two dis-
tinguishing characters (the thorn-like projections from harpe near
cucullus and the fold inclosing the pecten in hind wing of male)
are shared in greater or less degree by some species of Laspeyresia
which on genitalia and habitus go better with other and more nor-
mal Laspeyresia species than they do with pomonella. Indeed the
hind wing fold is not a good character.at all for in the toreuta
group of Laspeyresia it disappears so gradually as to leave one
often in doubt whether it is or is not present. I am convinced,
however, that some day when we know more of the larvae, Laspey-
resia will be further divided and that Carpocapsa will then apply
legitimately to one of the divisions. In the meantime nothing is
gained by suppressing a widely used name that eventually will
be restored. I am therefore keeping Carpocapsa and restricting
the genus to the type. None of the other species now included
goes as well with pomonella as with some typical Laspeyresia.
The two in out North American lists (saltitans Westwood and
sabastianniae Riley) I omit altogether from this paper. They are
Mexican insects, of whose occurrence in the United States we have
no authentic records.

KEY TO THE SPECIES OF CARPOCAPSA

1. General color of fore wing and thorax, grayish fuscous; hind wing smoky

FUSCOUS et = Sees Fae PE be 2 eee Spe get sd a in ad gee (1) pomonella.
General color of fore wing and thorax, golden ocherous; hind wing pale
OCT CHOU See se ee ene eer ee ah ee me LN ge pete (2) variety simpsoni.

1. CARPOCAPSA POMONELLA (Linnaeus)

(Figs. 37, 169, 329)

Phalaena Tinea pomonella LInNAEvS, Syst. Nat., ed. 10, vol. 1, 1758,
p. 538.

Carpocapsa pomonana TREITSCHKE, Schmet. Eur., vol. 8, 1830, p. 161.

Carpocapsa pomonella Harris, Inj. Ins., 1862, p. 484.—StTravupiIncer and
REBEL, Cat. Lepid., vol. 2, no. 2257, 1901.—Busck, Proc. Ent. Soc.
Washington, vol. 5, 1908, p. 235.—Simpson, Ent. Bull. U. S. Dept. Agr.,
no. 41, 1903.—Fostrr, Ent. Bull. U. S. Dept. Agr., no. 80, pt. 5, 1910;
Ent. Bull. U. S. Dept. Agr., no. 97, 1911.—HaAmm™emr, Ent. Bull. U. S.
Dept. Agr., no. 115, pt. 1, 1912—QUAINTANCE and GEYER, Ent. Bull.
U. S. Dept. Agr., no. 429, 1917.—Barnes and McDunnouGH, Check
List Lepid. Bor. Amer., no. 7270, 1917.—ForBres, Memoir 68, Cornell
Univ. Agr. Expt. Sta., 1924, p. 396.

Cydia pomonella WALSINGHAM, Proc. Zool. Soe. London, 1897, p. 1380;
Biol. Cent. Amer. Lepid. Heter., vol. 4, 1914, p. 259.—FERNALD, in
Dyar List, N. Amer. Lepid., no 5296, 1903.—Pterce and METCALFE,
Genitalia Brit. Tort., 1921, p. 82.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 71

The notorious codling moth has a very extensive literature.
Simpson’s bulletin, cited above, gives a bibliography to 1903 and the
Biologia all the systematic references to 1914. Only a few of the
more important references are given here. Detailed accounts of the
life history will be found in the Government bulletins.

Genitalia figured from specimens in National Collection from
Hyattsville Md. (male), and unknown locality (female).

Distribution.—The species has a wide distribution, corresponding
practically to the range of the cultivated apple. The following are
only the North American records from specimens in the four col-
lections studied: New Hampshire, New York, New Jersey, Penn-
sylvania, Maryland, Virginia, Georgia, Ohio, Iowa, Missouri, Ar-
kansas, Utah, California, Washington, British Columbia, Ontario,
Quebec.

Alar expanse. —15-22 mm.

Type.—Location unknown.

Type locality—KEurope.

Food plants—Apple, pear, walnut, quince.

2. CARPOCAPSA POMONELLA SIMPSONI (Busck)

Cydia pomonella simpsoni Buscx, Proe. Ent. Soe. Washington, vol. 5, 1903,
Dee2sab:
Carpocapsa pomonella sinypsoni BARNES and McDunnoueH, Check List
Lepid. Bor. Amer., no 7270a, 1917.
A rare and striking color variety. It is not in any strict sense
a race and shows no structural differences from typical pomonella;
but the varietal designation is convenient and should be retained,
particularly gs there are no intermediate, intergrading color forms.
Distribution—New Mexico, Colorado, Idaho, California.
Alar expanse —16-18 mm.
Type—tIn National Collection.
Type locality —Boise, Idaho.
Food plant—Apple.

15. Genus GYMNANDROSOMA Dyar
(Figs. 1, 3, 5, 38, 121)
Gymnandrosoma Dyar, Proc. Ent. Soc. Washington, vol. 6, 1904, p. 60.
Genotype.—Gymnandrosoma punctidiscanum Dyar (North America).

Thorax with posterior tuft.

Fore wing smooth; termen convex; 12 veins, all separate; 7 to
termen; 11 from cell at middle; 10 remote from 9; upper internal
vein of cell from between 9 and 10; 8, 4, and 5 remote at termen; 2
from cell just beyond middle, straight; no costal fold in male.

79 BULLETIN 1382, UNITED STATES NATIONAL MUSEUM

Hind wing with normal pecten from lower median vein; 8 veins; 6

and 7 approximate toward base; 5 parallel with 4 or (in desotanum)
. bent at base; 3 and 4 connate; inner margin in male roughly scaled
above and excavated below into a broad shallow pocket.

Hind tibia of male dilated, heavily rough sealed and with a heavy
dorsal hair pencile from base above.

Male genitalia as in H'cdytolopha except: spining of cucullus en-
croaching on neck of harpe; sacculus more weakly spined than neck.

Abdomen of male with a pair of dorso-lateral yellow hair penciles
from caudal edge of second segment.

Female genitalia with two strong thornlike signa. Ductus
bursae rather short; weakly chitinized at genital opening. Bursa
copulatrix with one or two strong chitinized patches on neck.

Closely related to E'cdytolopha.

KEY TO THE SPECIES OF GYMNANDROSOMA

1. Pale areas on outer third of fore wing, sordid whitish; hind wing with vein 5

SUT es a TE ee a (1) punctidiscanum.
Pale areas on outer third of fore wing brownish ocherous; hind wing with
Velnr ovbDentwaAt bases. 1s. se ee ee. se Se ee (2) desotanum.

1. GYMNANDROSOMA PUNCTIDISCANUM Dyar
(Figs. 1, 3, 5, 38, 121, 341)

Gymnandrosoma punctidiscanum Dyar, Proc. Ent. Soc. Washington, vol. 6,
1904, p. 60.—Knrarrorr, Proc. U. S. Nat. Mus., vol. 28, 1905, p. 362.—
Barnes and McDunnovuenH, Check List Lepid. Bor. Amer., no. 7252,
1917.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 397.

Resembles “/'cdytolopha insiticiana, differing chiefly in the second-
ary sexual characters of the male and in having a strong white dot
on fore wing at end of cell.

Genitalia figured from specimens in National Collection from
Quincy, Ill. (Poling, male) and Washington, D. C. (Busck, “Aug.
1903,” female).

Distribution.—Massachusetts, New York, New Jersey, Pennsyl-
vania, Maryland, District of Columbia, North Carolina, South Caro-
lina, Florida, Texas, Missouri, Illinois, Indiana.

Alar expanse.—16.5-25 mm.

Type.—In National Collection.

Type locality Washington, D. C.

2. GYMNANDROSOMA DESOTANUM, new species
(Fig. 123)

Antenna blackish fuscous; basal joint black. Palpus extending
a trifle more than the length of the head beyond it; blackish fuscous
slightly dusted with ocherous; upper edge sordid whitish. Face,

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 7S

head, and thorax blackish fuscous with a faint sprinkling of dark,
sordid ocherous scales. Fore wing with basal two-thirds blackish
fuscous; outer margin of this dark patch angulate, extending from
costa beyond middle outwardly to lower outer angle of cell and
thence inward to mid dorsum; a small round white discal dot at
upper outer angle of cell; a dark angulate subtornal spot with apex
touching the apex of the dark basal patch; area between lower outer
margin of basal dark patch and subtornal spot, sordid brownish
ocherous, distinguishable as a broad geminate dash; apical area sor-
did brownish ocherous with an irregular, blackish fuscous inwardly
curved, subapical band extending from apex to tornus; costa faintly
strigulated with blackish and brownish ocherous; in some speci-
mens an obscure pale blotch on costa before middle; cilia dark
fuscous with some sprinkling of dull ocherous. Hind wing dark
smoky fuscous; cilia but slightly paler with an obscure dark basal
band.

Female genitalia of type figured.

Alar expanse.—21-22 mm.

Type.——In American Museum.

Paraty pe-—Cat. No. 28025, U.S.N.M.; also in collection Barnes.

Type locality—Everglades, Fla.

Described from female type and four female paratypes from the
type locality, without collector label and dated “Apr. 19-1912” and
“April 8-15.” These had been set aside by Kearfott as a new species.

I have not seen the male and am therefore unable to place the
species with absolute certainty; particularly as vein 5 of hind wing
is bent at base as in the Olethreutinae. This, however, is probably
only a specific aberration. In wing pattern and female genitalia it
is quite similar to punctidiscanum.

16. Genus ECDYTOLOPHA Zeller
(Figs. 10, 115)
Ecdytolopha ZELurr, Verh. Zool.—bot. Ges. Wien, vol. 25, 1875, p. 266.
Genotype.—Eecdytolopha insiticiana Zeller (North America).

Thorax with posterior tuft.

Fore wing smooth; termen convex; 12 veins, all separate; 7 to
termen; 11 from cell at middle; 10 remote from 9; upper internal
vein of cell from between 9 and 10; 3, 4, and 5 remote at termen;
2 from cell before 24 (near middle), straight; no costal fold in
male.

Hind wing with normal pecten on lower median vein; 8 veins;
6 and 7 approximate toward base; 3 and 4 connate or very short
stalked; in male an appressed pocket containing a strong yellow
hair pencile at base of vein 1a.

Hind tibia of male loose scaled but not appreciably dilated or
tufted.

74 BULLETIN 182, UNITED STATES NATIONAL MUSEUM

Male genitalia with outer surface of harpe unspined; cucullus
large, spoon-shaped, finely and evenly spined; neck incurvation
slight; neck and sacculus evenly and finely clothed with hair-like
spines. Uncas absent. Socii absent. Gnathos a weakly chitinized
band. Aedoeagus long, slender, slightly curved and scarcely taper-
ing; cornuti a small cluster of moderately long, slender, deciduous
spines.

Abdomen of male simple.

Female genitalia with two thorn-like signa. Ductus bursae
rather long, unchitinized except at genital opening. Bursa coupla-
trix large, finely granulate at neck.

A small North American genus closely related to Gymnandrosoma
and with affinities to the H'ndothenia group of the Olethrentinae.
The genitalia are typically Laspeyresiin; but otherwise the genus
would go better with Hndothenia than with Laspeyresia. Prob-
ably a primitive form and (with Gymnandrosoma) linking the
Laspeyresiinae and Olethreutinae.

KbY TO THE SPECIES OF ECDYTOLOPHA

1. Fore wing with subapical bar distinct_____________=_______ (1) insiticiana
Fore wing with subapical bar nearly obsolete (very faintly indicated)___ 2.
2. Fore wing with subtornal spot on dorsum, distinet___-_________-__~_ (2) mana.
Fore wing with subtornal spot obsolete or very faint_________ (3) islandana.

1. ECDYTOLOPHA INSITICIANA Zeller
(Figs. 10, 115, 340)

Redytolopha insiticiana ZELLER, Verh. Zool.-bot. Ges. Wien., vol. 25, 1875,
p. 266.—PacKkarp, Fifth Report U. S. Ent. Com., 1890, p. 359.—FerEr-
NALD, in Dyar List N. Amer. Lepid., no. 5287, 1903——Barnes and Mc-
DunnoueH, Check List Lepid. Bor. Amer., no. 72538, 1917.—¥orses,
Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 397.

An insect of considerable economic importance. The larva is a
stem borer in Robinia. It attacks only new growth and forms a
large elongate gall which cracks open with age and disfigures the
tree. In the vicinity of Washington there are two generations a
year; the species overwintering as larvae under the debris on the
surface of the ground in flattened bean-shaped cocoons made of
pieces of fallen leaves, evenly cut, sewed together, and lined with
silk. Pupation takes place in April, and moths from overwintering
cocoons issue from early May until the end of June. These lay eggs
which hatch in from 5 to 6 days; the larvae feeding up, pupating
and again producing moths from early July to early September.
The larvae feed up in about 20 days during summer; but in the fall
take considerably longer, and those that hatch last from the eggs
often die before they have completed their growth. The entire life

- NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 15

cycle from adult to adult is completed in a little over a month (35
to 40 days) in early and middle summer. Larvae are to be found,
therefore, in nearly all stages from late May to early November (our
last field date for the vicinity of Washington is November 10).

A few specimens have been reared by W. G. Boyd at Weir,
Miss. (“7-14-1923”) from larvae boring in stems of Wisteria. As
this is an introduced plant, the infestation was probably accidental or
is evidence of a new, acquired habit. I know of no records other
than this upon any host but Robinia.

Genitalia figured from reared specimens in National Collection
from Falls Church, Virginia (“ Hopk. U. 8. No. 12103 7, issued 31
July 1914,” male; and “ Hopk. U. S. No. 121038 n, issued 26 June
1916,” female).

Distribution—New York, New Jersey, Pennsylvania, Maryland,
District of Columbia, Virginia, North Carolina, Florida, Mississippi,
Ohio, New Mexico, Colorado, Arizona, California, Ontario, Mani-
toba.

Alar expanse.—17-26 mm.

Type.—tIn British Museum.

Type locality —Massachusetts.

Food plants.—Roninia, Wisteria.

2. ECDYTOLOPHA MANA (Kearfott)
(Fig. 117)

Olethreutes mana KearrotTT, Trans. Amer. Ent. Soc., vol. 33, 1907, p. 14.—
Barnes and McDunnovucH, Check List. Lepid. Bor. Amer., No. 6822,
1917.

Olethreutes thaliastis Meyrick, Ent. Mo. Mag., vol. 48, 1912, p. 34.

Represented only by the female type in New York and a female
paratype in the Barnes Collection. Hardly to be distinguished from
insiticiana (of which it may be only an aberration) except by the
genitalia. The differences in structure are shown in figures 115, 117.

Genitalia figured from type.

Alar expanse-—18 mm.

Type.——tIn American Museum.

Lype locality —Black Jack Springs, Tex.

3. ECDYTOLOPHA ISLANDANA (Kearfott)
(Fig. 339)

Olethreutes islandana KearFort, Trans. Amer. Ent. Soc., vol. 33, 1907, p.
80.—BaRNES and McDunnoven, Check List Lepid. Bor. Amer., no.
6876, 1917.

Olethreutes insulicola Mryricx, Ent. Mo. Mag., vol. 48, 1912, p. 35.

Ecdytolopha islandana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 398.

76 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Known only from the male type. Kearfott described from two

specimens, but I am unable to locate the one that is supposed to be in
the National Museum. The type is in very poor condition, much
rubbed, mouldy, and lacking a head. It looks like a runted specimen
of insiticiana. The genitalia are similar except for size, being con-
siderably smaller. The name had better be kept until other speci-
mens can be obtained.

10.

able

13.

14.

Male genitalia figured from type.
Alar expanse —14 mm.

Type.—tin American Museum.

Type locality —Plummer Island, Md.

Subfamily OLETHREUTINAE

KEY TO THE GENERA OF OLETHREUTINAD

. Hind wing with veins 3-4 separate (or approximate) —----------_--_~_ 2.
Hind wing with 3-4 connate (rarely stalked) --_-----------__--_--_--- 4,

. Hind wing with 3, 4, and 5 well separated at base; 6 and 7 approximate or
slightly anastomosing beyond cell; thorax with posterior tuft_______ By
Hind wing with 3, 4, and 5 equidistant but closely approximate at base; 6
angaGostalked,: ‘thorax SMOOthe 2082. = soe en ae (2) Bactra.

. Gnathos heavily chitinized and solidly fused with anellus____ (4) Ahmosia.
CETTE TT OSS TN OTT BaD ota a ee (3) Polychrosis.

. Fore wing with termen concave, and veins 3, 4, and 5 approximate at
LeEMen” THOraAx SMOOth ADOVE2 = 282 oe eae (1) Episimus.
Fore wing with termen convex or straight and veins 3, 4, and 5 not ap-
proximate at termen; thorax with posterior tuft__________-_________~- 5.
Seeing nwin ai witha G=tuStaliked. ete seth sees le ee a ee ee 6.
Hind wing with 6-7 anastomosing beyond cell___________________-_ (7) Tia.
Hind wing with 6—7 approximate toward base________--_____-_-_-_------ 8

| NOCIi ‘SMOOLDMrigid=. 20u_. Bit ak A OTA EO Pe oy Sat (8) Hulda.
SOGE Un aity eG See ee 2 erate ri eld ol la oe ce f

PU NCUSETECOUCEH mwa Kev ihl Qatar a ee ge (6) Taniva.
WNEHIS HONEA SELON Spat ULAbe! ee ee (5) Endothenia.

. Fore wing with upper internal vein of cell from between veins 9-10.

(11) Zomaria.

Fore wing with upper internal vein of cell from between veins 10-11___ 9.
. Fore wing with vein 2 from cell beyond 34____________________ (19) Evora.
Hore wine with vein’) from cell before 342. 2 ee ee 10.
Hind tibia of male with hair pencile from base________________________ 13.
Hind tibia of male without hair pencile from base____________________ ie
Gnathos produced into a strongly chitinized projecting tongue____ (9) Esia.
Gnathos norm gl Sse Tepe ES Pa ee OAS See Se ire
. Uncus strongly spined beneath; socii nearly obsolete_______ (13) Sciaphila.

Uncus unspined beneath (much reduced) ; socii well developed.
(10) Eumarozia.

GATOS (SCLONS DY SCOMMACO ee ten ee ee eee (14) Badebecia.
Gmathosssmootny bet Ska ce Oe ed Lee Le EAE OS a eee ay ee ee ee Se 14.
Subanal plate of gnathos strongly chitinized; semitubular___ (12) Aphania.

Subanal plate of gnathos weakly chitinized, flat__.____________________ 15.

Uncus

Tt 7
eqern itn iat Hottest Hed
jade Aypeoq pre kavoy dam | iii ix os Ser ie ape et
pourds Apyevom diy wm | x< ix x x ix xX
sourds 300138 Disc tie tat Wale otal th ot
qua diy ye poursivor ATpreMuy Mcgee: ENS Hon emeili eaten Hey

----|----|--

[os eee | see eee tie

----|----|----| 7

S255) 2S

-25-|----]--|

—_
al
a
>

’

COMPARATIVE TABLE OF STRUCTURAL CHARACATERS: OLETHREUTINAE

Wings

2
.
E

bet Ri

(war a}) eden OY oa] |

Wap eried
= oe 1-0 SOP,

enoayrordde 1-0 eapy |

——

XK XK XXX XRX

fen ae cee

enwouoo nowy, |X! |
AqAyuNs 79 r9AG00 wom, | |XX XK
H s19;0q 1199 mo EOPA| XX | x

eu0je 10 at

; % paoteg 19 moe ae, ¢

xX XK

fs ‘ama
aH owinpronddy 400 o-h¢ Haye A
| oma 1» orecrpxouddy oe eCTDA

pene ee

XXKXKK XKK

§ m0y prremmdos tram ome, |

01-6 wows yo waren soddy | |

11-01 m0 9% [wouNITT ATEN,

XXX Xx

yma sope180d INOUE,

. 907 oywtapROUdde 46 OA,
|
|

Tio souMeod aA)

Genus

‘54S48—25. (Face p 76.)

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE

GYNIZAGUHLIAIO THL JO AWAY, OMANGTDOIAHG—Z “Ol

GIWYLYOL ZAILIWIYd

JVNILNIGHLI 10

SPLNISHLIT) v93g avg
(F/)
(9) (eyo
VYOHAOISV IFUL ViesHOZ vie
(Si) vizourwny jwamny (4) VINYL
(01) (9) VINSHLOONJ
(SisowHy
(%) | vaLovg
VYOAJ (2)
61) WSF oe ‘d SAWISId
(G) (1)
IVNIWSOONA
FVOIDIYLYO_L SIVNUS FLA FAS V7

78 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

15. Hind tibia of male swollen with scale tuftings________ (15) Phaecasiophora.
Enmdstibiaoteamnale smoothuscaled= = - = oo sseeh ne 16.
iG bursa of female with twoysiena- 2202s he ee ee (17) Hedia,
iBicsa,or female with single;signum or none=_ 222225". Sse eee ale
17. Hind wing of male with projecting basal lobe__-_----__--__ (16) Exartema.
Hind wing of male without projecting basal lobe_-_-___- (18) Olethreutes.

1. Genus EPISIMUS Walsingham
(Figs. 21, 22, 39)

Episimus WALSINGHAM, Proc. Zool. Soc. London, 1891, p. 501, 1897, p. 122.
Genotype.—Carpocapsa transferrana Walker (Brazil).

Thorax smooth above; with expansible tuft from near head below
base of fore wing.

Fore wing smooth; termen concave; 12 veins, all separate; 7 to
termen; 7, 8, and 9 approximate at base; upper internal vein of
cell from between 10-11; 3, 4, and 5 approximate at termen; 2 from
cell near 14, straight or but very slightly bent.

Hind wing with 8 veins; 6 and 7 approximate towards base; 3 and
4 connate (stalked in tyrius); 5 closely approximate to 4 at base;
male without chitinots ridge at inner margin.

Hind tibia of male without hair pencile from base.

Male genitalia with harpe narrowly elongate; outer surface un-
spined; cucullus long and narrow, finely and evenly spined, apex
evenly rounded; sacculus with a few long, flat, hairlike spines
(ScSp) from base; spine cluster Spc upon a slight projection from
lower margin of harpe beyond base; spine cluster Spc? absent.
Uncus developed, rather long, slender, simple. Socii broad, flexible,
heavily haired. Gnathos moderately chitinized, somewhat reduced
and terminating in a narrow, tapering, finely pointed, flat subanal
plate. Aedoeagus moderately long, straight; cornuti a dense cluster
of long slender deciduous spines.

Female genitalia with two thornlike signa. Ductus bursae rather
short, with a chitinized collar at juncture of bursa.

A tropical American genus represented in North America by a
few species of West Indian or Central American origin. In addi-
tion to the three here treated several others common to the West
Indies probably inhabit lower Florida. I have before me a badly
mutilated specimen of what looks like nestotes Walsingham, and
three specimens of an apparently undescribed species. They are
all in too poor condition, however, for definite determination.

The genus is a very compact, clearly defined one. The genitaha
throughout are of a uniform type with very slight specific differ-
ences. The pattern (fig. 22) also is remarkably consistent and
typical. Its striking feature is an ocelloid patch on fore wing, con-

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 719

sisting of a pale area at tornus with a more or less pronounced
dark central spot, a partially encircling dark band and, above this,
two or more black dots or streaks, the whole reaching well up toward
apex at termen, bounded inwardly by more or less metallic scaling
and above by a narrow dark curved band extending to termen.
Specific differences in color and markings are often slight but ap-
pear to be consistent.

On several characters (the smooth thorax, distinctly concave ter-
men, approximate condition of veins 3, 4, and 5 at termen, and the
strongly marked ocelloid patch of fore wing) EH’ pistmus would go
as well in EKucosminae as in Olethreutinae. The male genitalia are,
‘however, more olethreutine than otherwise, and in all the species that
I have seen except one (tyriws) veins 3 and 4 of hind wing are dis-
tinctly connate. In tyrius they are stalked.

The genus, therefore, while it lies close to the Eucosminae and
shows certain affinities to the ’pinotia branch of that subfamily
should be regarded as part of the Olethreutinae.

I should perhaps mention that the tuft on thorax under fore wing
and which Walsingham gives as a distinguishing character for
EH’ pisimus occurs in practically all Olethreutinae as a simple scale-
tuft. In H'pistmus, however, it has more or less of a mixture of hair
and scales and in some species the hairs are very thick and long
(transferranus and nesiotes for example).

KEY TO THE SPECIES OF BPISIMUS

1. Fore wing with a conspicuous, rounded, dark spot on dorsum just before

PID (1 1 Gitees hed ene an, Serre ne a Ne ee a (2) augmentanus.

HOLE RWAN Sv MOULy SUC Masa eer ne ee eee ee oe ee as

2. Fore wing with a large reddish purple costal patch covering most of basal

half of wing; dorsal area bordering this patch, whitish; hind wing with

weins). 3 jand4 stalkediocte lk | oo eo ieee ey FS yee. eA eto (3) tyrius.

Fore wing with costal patch an obscure dark smudge on midcosta; dorsal
margin at base not whitish; hind wing with 3 and 4 connate.

(1) argutanus.

1. EPISIMUS ARGUTANUS (Clemens)
(Figs. 183, 392)

Bactra? argutana CLEMENS, Proc. Acad. Nat. Sci. Philadelphia, 1860, p.
358.—PAaAcKARD, Fifth Rep. U. S. Ent. Comm., 1890, p. 282.

Catastega hamameliella CLEMENS, Proc. Ent. Soe. Philadelphia, vol. 1,
1861, p. 87; Tin. N. Amer., 1872, p. 178.—Dyar, Proc. Ent. Soc. Wash-
ington, vol. 5, 1903, p. 128.—FErNatp, in Dyar List N. Amer. Lepid.,
no. 5807, 1903.—BarneEs and McDuNnnoueuH, Check List Lepid. Bor.
Amer., no. 7279, 1917.

Grapholitha (Hedya) allutana ZeLtitErR, Verh. Zool.-bot. Ges. Wien, 1879,
p. 295.

80 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Episimus argutanus Dyar, Proc. Ent. Soc. Washington, vol. 4, 1901, p.
469.— FERNALD, in Dyar List N. Amer. Lepid., no. 5207, 1903.—KrEar-
Fort, Ins. New Jersey, 1910, p. 543.—Barnes and McDuNNovuecH, Check
List Lepid. Bor. Amer., no. 7110, 1917.—Forpres, Memoir 68, Cornell
Univ. Agr. Exp. Sta., 1924, p. 447.

Enarmonia argutana WALSINGHAM, Biol. Cent. Amer. Lepid. Heter., vol. 4,
1914, p. 238.

A rather common species and probably the most widely dis-
tributed of all, occurring throughout North and Central America
and the West Indies. Dyar has suggested the synonymy of hama-
meliella and argutana and in this is without doubt correct. Several
of the moths in the National Collection were reared from Hama-
melis. The favorite food plant, however, seems to be sumac (Rhus
glabra and R. copalina, less frequently 2. toxicodendron). 'There is
some little color variation in different specimens due to the amount
of dark dusting upon fore wing; but the pattern markings other-
wise are uniform.

Male genitalia figured from specimen in collection Barnes from
Shasta Retreat, Siskiyou County, Calif. (“June 16-23”); female
from reared specimen in National Collection from Glencarlyn, Va.
(on Rhus copalina, Hopk. U.S. no. 12133, June 10, 1914, Heinrich).

Distribution—F lorida, North Carolina, Virginia, Maryland, Dis-
trict of Columbia, Pennsylvania, New Jersey, New York, Maine,
Illinois, Missouri, Texas, New Mexico, Colorado, California, Mani-
toba, Ontario.

Alar expanse.—11-15 mm.

Types.—In Academy Natural Science, Philadelphia (argutanus) ;
British Museum (allutana).

Type localities—Pennsylvania (argutanus, hamameliella) ; Texas
(allutana).

Food plants—Rhus, Hamamelis, Crataegus, Ulmus, Solidago,
EKuphorbia heterophylla.

2. EPISIMUS AUGMENTANUS (Zeller)
(Fig. 393)
Grapholitha (Hedya) augmentana ZELLER, Hor. Soc. Ent. Ross., vol. 18,
1877, p. 160.
Episimus augmentanus Dyar, Proce. Ent. Soc. Washington, vol. 4, 1901, p.
468.—FERNALD, in Dyar List N. Amer. Lepid., no. 5206, 1903.—BaARNES
and McDunnouau, Check List Lepid. Bor. Amer., no. 7109, 1917.
Represented in the National Collection by two reared males from
Palm Beach, Fla. (Dyar, no. 843, iss. Feb. 25 and 27, 1900). ‘These
are the only North American specimens I have seen. Male genitalia
figured from one of the above.
The species is very similar in pattern to the tropical transferranus
Walker, but is apparently distinct.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 81

Alar expanse.—14-15 mm.

Type.—tin collection Staudinger.

Type locality.—Cuba.

Food plant.—Metopium toxiferum (“ Rhus metopium”).

3. EPISIMUS TYRIUS Heinrich
(Fig. 187)

Episimus tyrius HetnricH, Proc. Ent. Soe. Washington, vol. 25, 1923, p.
107.—Forses, Memoir 68, Cornell Uniy. Agr. Exp. Sta., 1924, p. 447.

So far represented by the type in the Cornell Collection, a female
paratype from St. Petersburg, Fla., in the Barnes Collection, a male
from Biloxi, Miss. (June 13, 1917), also in the Cornell Collection,
and a male and female from the Fernald Collection in the National
Museum (these last are part of the same reared series as the type.
They have been partially eaten by a Dermestid but wings and abdo-
mens are intact). In all specimens veins 3 and 4 of hind wing are
stalked, a very exceptional character in this subfamily and here
apparently a specific rather than an individual aberration.

Female genitalia figured from type. Male genitalia similar to
that of transferranus (fig. 39), but with slightly heavier spine-tuft,
Spe.1

Alar expanse.—14-15 mm.

Type.—tin collection Cornell University.

Type locality—Westbury Station, Queens County, N. Y.

Food plant.—A cer dasycarpum.

2. Genus BACTRA Stephens
(Figs. 44, 45, 46, 47, 49, 342)

Bactra STEPHENS, Illus. Brit. Ent., vol. 4, 1834, p. 124.
Genotype.—Tortriz lanceolana Hiibner (Europe).

Thorax smooth.

Fore wing smooth; termen straight; 12 veins, all separate; 7 to
termen; 8 and 9 approximate (but not closely so) ; 10 well separated
from 9 but closer to 9 than to 11; upper internal vein of cell from
between 10-11; 3, 4, and 5 not approximate at termen; 2 from cell
slightly before 24, straight.

Hind wing with 8 veins; 6 and 7 stalked; 3 and 4 separate; 3, 4,
and 5 approximate at base and equidistant; male without chitinous
ridge at inner margin.

Hind tibia of male without hair pencile from base.

Male genitalia with harpe short and stout; outer surface unspined ;
cucullus broad, strongly spined along lower margin; sacculus broad,

So BULLETIN 132, UNITED STATES NATIONAL MUSEUM

weakly haired at base; spine cluster Spc! rarely absent (in type and
furfurana), when present developed as a strong ridge of spines upon
a free arm projecting over cucullus; spine cluster Spc? present.
Tegumen short, broad, roundly arched. Uncus short, stout, with a
row of strong spines on inner side at margin. Socii small, flexible,
weakly haired. Gnathos simple, weakly chitinized. Aedoeagus
moderately long to very long, stout, more or less curved; cornuti a
eluster of two or more short spines, rarely absent.

Female genitalia with signum a small flat.scobinate patch. Duc-
tus bursae moderately long, simple.

A compact easily distinguished genus of rather wide distribution,
showing, in genitalia at least, rather marked affinities to Lndothenia.
The male genitalia are quite characteristic. There appear to be two
types of harpe: the lanceolana type with spine group Spc’ absent
and the cucullus spines dense and slender; and the verutana type
with Spc! strongly developed and upon a free arm, and with the
cucullus spines fewer but extremely stout (compare figs. 44, 46).
Otherwise the genitalia are of the same general type, with slight but
clear-cut specific differences.

In pattern and markings the species are variable, and grade into
each other. This has caused considerable misidentification and con-
fusion. Except for the more strongly and characteristicly marked
specimens, accurate and certain determination is only possible by
means of the genitalia. The following specific key is drawn for such
typically marked examples, and will help to identify them, but will
not serve for all specimens of any given species.

KEY TO THE SPECIES OF BACTRA

1. Fore wing with outwardly angulate dark basal patch; if incumplete, at least

indicated by outer dark margin on costal half_____________~_ (2) furfurana.
Fore wing without such basal patch; dark basal markings not reaching
EO COS ease rar Cae a oe ee a See TA RR se ae We ate Pee

2. Fore wing with a strong, straight, continuous, central longitudinal fuscous
shadelifrom: base. tol fapexct. cbs See i ee ey tds Peg OT Ce 3.
Fore wing without such; where there is a continuous dark streak from
base to apex the latter is sharply angulate at end of cell_____________ 4.

3. Dark costal geminations on fore wing very short_____-_-----__ (6) maiorina.
Darkecostal Seminavions, (ONS == s— ew eee (7) priapeia (part).

(8) sinistra.

4, Fore wing with a conspicuous white dot at end of cell.
(4) verutana albipuncta.
HORS syn Sav OWE, SUC hy oe 2 ee Le
5. Dark markings on disk forming a more or less interrupted longitudinal
streak, or a longitudinal streak at base and a short transvere streak across
end of cell, hooked at its lower extremity________________ (1) lanceolana.
Dark markings on disk two round or elongate blackish spots, one near base,
the:other at end ofcelkieet task ae See Ee ee ee eee 6.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 83

6. General color of fore wing pale cinereous ocherous; dark dusting blackish
TATRC Ty Se OS pea ae. ee eee (3) verutana (typical).

(7) priapeia (part).

General color of fore wing pale brownish ocherous; dark dusting more brown

RAE LEU TAU PDL Gh CK See oP BEA ee 2S eC es a ae et SEE (5) verutana chrysea.

1. BACTRA LANCEOLANA (Hiibner)
(Figs. 44, 342)

Tortriz lanceolana HUBNER, Samm. Hur. Schmet., Tort., 1800, fig. 80.
Bactra lanceolana STAUDINGER and REBEL, Cat. Lepid., vol. 2, no. 2017,
1901.—FERNALD, in Dyar List N. Amer. Lepid., no. 5006, 1903.—PIERcE
and MeEtTcALreE, Genitalia Brit. Tort., 1922, p. 40, pl. 14.
Pierce’s figures of the genitalia fix the concept of this species to
a form with broad, roundly arched incurvation on lower margin of
harpe between sacculus and cucullus. In furfurana, which is very
close in genitalia, pattern, and general habitus, the lower margin of
harpe is angulate and there is no incurvate or definable neck area
(comp. figs. 44, 45). European and American workers have fre-
quently confused the two species, identifying suffused and weakly
marked furfurana as lanceolana. The latter occurs here, but is quite
rare. All the American specimens I have seen under the name are
referable elsewhere, some few to mazorina, but the greater number to
furfurana or verutana (var. albipuncta). The majority of the
European specimens in our collections under Janceolana (and so
labeled by European workers) are also furfurana.
Male genitalia figured from specimens in National Collection from
British Columbia (E. H. Blackmore, “60, F-13-VITI-20”).
This is the only authentic American specimen I have seen.
Alar expanse.—15 mm.
Type.—Location unknown.
Type locality. Europe.
Food plant.—Juncus.

2. BACTRA FURFURANA (Haworth)
(Figs. 45, 170, 343)

Tortriz furfurana HAawortH, Lepid. Brit., 1811, p. 466.

Bactra furfurana SravupIncer and Reset, Cat. Lepid., vol. 2, no. 2020,
1901.—FERNALD, in Dyar List N. Amer. Lepid., no. 5007, 1903.—PIERCcE
and MetTcaLFe, Genitalia Brit. Tort., 1922, p. 40, pl. 14.—ForBgEs,
Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 470.

A common species throughout the United States. The characters
given in our key will distinguish the more strongly marked speci-
mens (that is, those with complete basal patch, and median bar
extending to costa) ; but the species is quite variable and specimens
marked like typical /anceolana are not uncommon. In fact these are

84 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

usually wrongly identified as lanceolana. For accurate and certain
determination of any Bactra a genitalia preparation is necessary.
Fortunately these organs (both male and female) show good specific
differences. .

Male and female genitalia figured from specimens in National
Collection from Europe (male) and Washington, D. C. (August
Busck, July, female).

Distribution.—Ontario, District of Columbia, South Dakota, Ilh-
nois, Tennessee, Missouri, Louisiana, Alabama, Texas, Montana,
California. .

Alar expanse.—10-16 mm.

Type—tIn British Museum.

Type locality —England.

Food plant.—Juncus.

3. BACTRA VERUTANA Zeller

(Figs. 47, 171, 346)

Bactra lanceolana verutana ZELLER, Verh. Zool.—bot. Ges. Wien, vol. 25,
1875, p. 247.— FERNALD, in Dyar List N. Amer. Lepid., no. 5006, 1903.—
BaRNES and McDunnoucH, Check List Lepid. Bor. Amer., no. 6789,
1917.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 470.

A common species in Florida and the other Gulf States. The
typical form occurs rarely in northern localities. Described as a
variety of lanceolana and so listed in our catalogues. It is, how-
ever, quite distinct.

Male and female genitalia figured from specimens in National
Collection from Orlando, Fla. (G. G. Ainsle, March, 1914, male,
and “2-2-18,” female).

Distribution —F lorida, Texas, Mississippi, Louisiana, North Caro-
lina, Indiana, Missouri, Ontario, Alberta.

Alar expanse.—11-17 mm.

Type—tIn British Museum.

Type locality.—Dallas, 'Tex.

Food plant—Cyperus (U.S. Bureau of Entomology rearing).

4. BACTRA VERUTANA ALBIPUNCTA, new variety
(Figs. 46, 347)

A Rocky Mountain race of verutana, with darker head (pale
brownish ocherous) and with brownish fuscous rather than blackish
markings on fore wing. In well-marked specimens the outer discal
dark spot is continued in a dark shade to mid costa and there is a
distinct white dot at end of cell. The chief differences, however,
are in the male genitalia. In albipuncta spine cluster Spec? is
grouped on the outer margin of the raised area of sacculus while in

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 85

typical verutana it is on or very near the basal margin (comp. figs.
47,46). The spining of cucullus is also stouter in albipuncta. There
are no significant differences in female genitalia.

Male genitalia figured from paratype in National Collection
(“ Colo. 2527”).

Alar expanse.—14-18 mm.

Type and paratypes.—Cat. No. 28026, U.S.N.M. Paratype also in
American Museum, collection Barnes, and Canadian National Col-
lection.

Type locality—Denver, Colo.

Described from male type and 8 male and 1 female paratypes
from the type locality (Oslar, collector) ; 1 male paratype labeled,
“Colo. 2527;” 3 female paratypes from Vineyard, Utah; 2 male
paratypes from Eureka, Utah; 5 male paratypes from Deer Creek,
Provo Canon, Utah; and 1 male paratype from Stockton, Utah
(all Utah specimens collected by Tom Spalding and bearing various
July dates).

The above out of a large series which had been in the collec-
tions under either /anceolana or furfurana. I have also before me
a few specimens from Arizona.

Several genitalia slides were made from specimens from the dif-
ferent localities. In all, the distinguishing male characters were
found to be constant.

5. BACTRA VERUTANA CHRYSEA, new variety
(Figs. 49, 348)

A California race of verutana.

Like albipuncta but paler, more uniformly yellowish, and with
white dot on outer margin of cell obsolete or very faint. Male geni-
talia with lower margin of cucullus of harpe right angulate (in albz-
puncta and typical verutana it is more evenly rounded) ; spine clus-
ter Spc* also more scattered in middle of raised area of sacculus.

Male genitalia figured from type.

Alar expanse.—16-18 mm.

Type.—tn collection Barnes.

Paratypes.—Cat. No. 28027, U.S.N.M. Also in American Museum
and collection Barnes.

Type locality —Loma Linda, Calif.

Described from male type and 9 female paratypes from the type
locality dated “ March 24-30” (type), “Aug. 24-31” (1 paratype),
“Sept. 1-7” (2 paratypes), “Sept. 16-23” (3 paratypes), “Sept.
24-30” (1 paratype), “Oct. 8-15” (1 paratype), and “ Oct. 16-23”
(1 paratype) ; and one female paratype from Laguna Beach, Calif.
(Baker, no date).

86 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

6. BACTRA MAIORINA Heinrich
(Figs. 173, 344)

Bactra maiorina HeEtnricH, Proc. Ent. Soc. Washington, vol. 25, 1923, p.
105.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 470.

A distinct species, as the genitalia show. In the male the aedoeagus
is longer and more curved, spine cluster Spc? is stronger and the
heavy spining of cucullus more closely crowded toward lower margin
than in any of the varieties of verutana. The differences in female
genitalia are shown in figures 171, 173.

Apparently confined to the single food plant (Scirpus) and pos-
sibly the same as the European scirpana Herrich-Schaefer, now listed
as a synonym of furfurana. A genitalia study of European speci-
mens reared from Scirpus would be necessary to establish this.

Male genitalia figured from paratype in collection Barnes from
Vineyard, Utah (Tom Spalding, “ VI-2-12”); female from para-
type in collection Barnes from Arlington, Va.

Distribution.—Virginia, Indiana, Illinois, Missouri, Utah. There
is also a specimen in the Cornell collection from Ithaca, N. Y.

Alar expanse.—13-20 mm.

Type.—tIn National Collection.

Type locality Arlington, Va.

Food plant.—Scirpus fluviatilis.

7. BACTRA PRIAPEIA Heinrich
(Figs. 172, 345)

Bactra priapeia Hrtnricu, Proc. Ent. Soc. Washington, vol. 25, 1923, p.
105.—ForBEs, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 470.

Since the publication of this species a large number of Bactra col-
lected at San Benito, and Brownsville, Tex., have been received from
Dr. W. Barnes. Most of the specimens were verutana or furfurana
(dated May to September). The females of priapeto in this lot
showed quite different genitalia from those of the female paratypes
of the original type series. Inasmuch as the Texas females are in
large series (equal in number to the males), I take it that they must
be the true females of priapeia, and that the paratype from the type
locality is something else, possibly a hybrid of priapeza and verutana
or of verutana and furfurana. JI am designating it as a new species
under the name sinéstra (see following).

In pattern priapeia is somewhat variable, several specimens hav-
ing the median dark shade of forewing broken into two discal dots
or blotches as in verutana. The abnormally long aedoeagus of the
male and the strongly chitinized ductus bursae of the female, how-
ever, are diagnostic and readily identify the species.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 87

Genitalia figured from specimens in National Coilection from
Sabine River Ferry, La. (June 20, 1917, male paratype), and San
Benito, Tex. (“July 16-23,” female).

Distribution.—Texas, Louisiana, Florida. In the National Col-
lection there is also a well-matched pair (male and female) from
Corazal, Canal Zone, Panama (August Busck, collector).

Alar expanse.—16-24 mm.

Type.—tIn collection Cornell University.

Type locality —Sabine River Ferry, La.

8. BACTRA SINISTRA, new species
(Fig. 174)

Similar to male of B, priapeta and only to be distinguished from
that species by the female genitalia. These are quite characteristic,
however. The peculiar structures are shown in our figure. All
pattern markings like those of male of priapeia; fore wing with a
median longitudinal dark fuscous streak from base to apex.

Alar expanse —16-17 mm.

Type and paratype—Cat. No. 28028, U.S.N.M. Paratype also in
Cornell University and Barnes collection.

Type locality—Sabine River Ferry, La.

Described from female type (Cornell University Lot 542, Sub 20) ;
one female paratype from Biloxi, Miss. (Cornell University Lot
542, Sub 11); and two female paratypes from Gulfport, Miss.
(F. H. Benjamin).

I should not name this form were it not for the fact that the type
had been wrongly referred as a paratype of priapeia Heinrich, and
that two other specimens had been sent out under that name. I be-
lieve it is only a hybrid of priapeia and verutana (or verutana and
furfurana) ; but as this is by no means certain, a separate specific
designation must be given.

3. Genus POLYCHROSIS Ragonot
(Figs. 12,'15, 19, 20, 41, 358)
Polychrosis RAGonot, Ann. Ent. Soe. France, vol. 63, 1894, p. 209.
Genotype.—Tortriz botrana Schiffermtiller (Europe).

Thorax with posterior crest.

Fore wing smooth; termen convex; 12 yeins, all separate (rarely
8 and 9 connate) ; 7 to termen; 7, 8, and 9 approximate; 10 remote
from 9; upper internal vein of cell from between 10-11; 3, 4, and 5
not approximate at termen; 2 from cell slightly beyond 34, some-
what bent.

Hind wing with 8 veins; 6 and 7 approximate toward base
(rarely anastomosing beyond cell); 3 and 4 separate; 3, 4, and 5

88 BULLETIN 182, UNITED STATES NATIONAL MUSEUM

equidistant and well separated; male without chitinous ridge at
inner margin.

Hind tibia of male with short hair pencile from base.

Male genitalia with harpe moderately long; outer surface un-
spined; cucullus well spined throughout, apex evenly rounded; sac-
culus normally with a strong tuft of hairlike spines from base;
spine clusters Spc! and Spc? strongly developed, produced from
neck (Spc? rarely absent). Uncus reduced, weakly chitinized, finely
spined beneath. Socii absent. Gnathos simple. Aedoeagus long,
slender, slightly curved; cornuti absent.

Female genitalia without signum. Ductus bursae moderately
long, simple.

Abdomen (of male) with a narrow elongate pocket of papilliform
hairs on each side of basal segment (fig. 15).

The above description is drawn for the American species, which
differ rather markedly from the type (botrana) in both fore-wing
venation and genitalia and probably should have separate generic
rank. The European species have similar genitalia to botrana; but
nothing has quite the same fore-wing venation (10 approximate to
9). Neither do they agree any better with the type of Lobesia
(permiatana Hiibner) ; artemesiana Zeller and kreithneriana Hornig
have similar fore-wing venation (10 and 11 approximate), but lack
the peculiar trigonate hind wing of permixtana. Eventually we
shall probably have to restrict the two genera (Polychrosis and
Lobesia) to their respective types and find new designations for the
species now referred to them. For the present or until such time as
the exotic forms can be carefully studied and generic limits exactly
defined, I think it better to keep our American species under Poly-
chrosis.

Meyrick in his Australian Revision *° gives, as one of the charac-
ters of Polychrosis, 6 and 7 of hind-wing connate or stalked. I do
not know how this applies to eastern species. It does not for the
European or American. In the type and most of the other species
6 and 7 are clearly tortriciform (approximate toward base). Rarely
(in specimens, and possibly holding for individual species) they
anastomose slightly beyond the cell.

KEY TO THE SPECIES OF POLYCHROSIS

1. Basal two-thirds of fore wing ashy gray; without dark median band.
(14) ecyclopiana.
Basal two-thirds of fore wing not ashy gray; dark median band present_ 2.
2. Basal patch and antemedian band of fore wing concolorous and coal-

CYS 0 Up Ie ga a Ce cL ee al) Ne er ee 3.
Basal patch and antemedian band not concolorous; both defined to the
foie Seta it (eh Coy as es RE SS Se WEAR MR inne Rt Wits RR east SL ene Se ES aN I ie

270 Proc. Linn. Soc. N. S. W., vol. 36, 1911, p. 256.

10.

ra Ip

12.

13.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 89

. Head and thorax a uniform ferruginous ocherous_______ (1) liriodendrana.

Head and thorax pale ocherous or purplish fuscous dusted with ocher-
OUR See ea te ee ae ge 4.

. Median band of fore wing well separated from pretornal spot by a metallic

band as broad as dorsum of median band_______________ (2) viteana,
Median band more or less coalescing with pretornal spot, the intervening
metallic band narrow and dusted with ocherous and brownish scales_ 5.

. Fore wing with costa markedly rounded toward apex; veins 8 and 9 very

closely approximate at base, practically connate; 7 and 8 more separated ;
median band slightly broader on costa than dorsum, Hind wing blackish
EUS COU See eee BRA Sh a ee ASRS) . EEE EEE AON RAS (3) monotropana.
Fore wing with costa only slightly rounded (nearly straight toward apex;
veins 8 and 9 separate at base, further apart than 7 and 8; median band
narrower on costa than dorsum. Hind wing brownish (smoky) fuscous,
decidediyipaler: toward-ibaseses< 2 ee Se ee ke ee 6.

. Subapical spot much dusted with blackish, concolorous with median band.

(5) rhoifructana.
Subapical spot with little or no blackish dusting, paler than median band.
(4) cypripediana.

. Hind wing smoky only toward apex; distinctly white toward base________ 8.

Hind wing smoky throughout; not appreciably whitish toward base____ 9.

. Fore wing with dark pattern markings distinctly bordered with white; alar

PXPAMSCyUMG ET? Dy ag es ah ra eee ee epee (6) yaracana.
Fore wing with no white borders to dark pattern markings; alar expanse
PMN Gs OP OVER == eta: eee wane ew aap a 20 NE et (13) blandula.

. No white scaling whatever upon fore wing; pale dashes bordering outer

COStal SHOTS OCHETOUS! Ss Loe Rene | SSE A (11) slingerlandana.
Some white scaling upon fore wing; pale dashes (those bordering outer
costal spots at least) white; sometimes median and basal dark mark-

IN CS Sea thyR CULE: waitin Nitec a es RAE Ue Be ee ee 10.
Subapical spot uniformly blackish brown______----__--____ (12) carduana.
Subapical spot paler; sometimes with shading or dotting of black scales,

but not uniformly blackish or blackish - brown ---___-..-_.________..- ae
Dorsal half of median band bright ocher yellow____________ (10) aruncana.
Dorsal half of median band not bright ocher yellow__________________ 12;
Subapical spot showing considerable black scaling, as dark as median

eS SAE ee pee ens eile wk ee, asd SE es eee 13
Subapical spot without (or with but the faintest trace of) black sealing,

paler; than median) band te. 324% ot fe sspi ek ey (7) spiraeifoliana.
Antemedian band paler on dorsum than costa, appearing to the naked eye

as a rounds: whitish dorsal’ spot] = = 22) 2 gs (8) aemulana.

Antemedian band no paler on dorsum than costa, lead colored throughout.
(9) vernoniana.

1. POLYCHROSIS LIRIODENDRANA Kearfott
(Figs. 175, 361)

Polychrosis liriodendrana Krarrott, Trans. Amer. Ent. So0c., vol. 30, 1904,
p. 293.—Barnes and McDuNnNovueH, Check List Lepid. Bor. Amer., no.
6780, 1917.—ForBes, Memoir 68 Cornell Univ. Agr. Exp. Sta., 1924, p.
473.

Polychrosis magnoliana KrarrottT, Trans. Amer. Ent. Soc., vol. 33, 1907,
p. 6.—Barnes and McDunnowuGH, Check List. Lepid. Bor. Amer., no.
6781, 1917.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 473.

90 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

I have examined the types of Kearfott’s two species carefully and
can see no real difference between them, genitalic or otherwise, except
size. P. liriodendrana is somewhat larger; but in reared series from
Liriodendron we have specimens quite as small as any from Magnolia.
The larvae feed commonly upon the leaves, rarely in the seed pods.

Male genitalia with long tuft from base of sacculus of harpe;
arch of neck wide; spine cluster Spc? absent; aedoeagus smooth.

Genitalia figured from reared specimens in National Collection
from Falls Church, Virginia (Hopk. U. S. no. 11149, Heinrich, June
28, 1913, male) and Montclair, New Jersey (‘“ K-672, iss. VIII-
10,” Kearfott, female).

Distribution—New Jersey, Pennsylvania, District of Columbia,
North Carolina.

Alar expanse —10-14 mm.

Types——In American Museum.

Type locality—District of Columbia (liriodendrana and mag-
noliana) .

Food plants.—Liriodendron tulipifera, Magnolia virginiana.

2. POLYCHROSIS VITEANA (Clemens)
(Figs. 12, 182, 362)

Endopiza? viteanad CLEMENS, Proc. Acad. Nat. Sci. Philadelphia, 1860, p.
359.

Penthina vitivorana Packarp, Guide Study Ins., 1869, p. 336.—RiLey, Rep.
Ins. Missouri, vol. 1, 1869, p. 183—WatsH and Ritey, Amer. Ent.,
vol. 1, 1869, p. 177.

Polychrosis botrana FernaLp (and Authors, not Schiffermtiller), in Dyar
List N. Amer. Lepid., no. 5005, 1903.

Polychrosis viteana Krarrott, Trans. Amer. Ent. Soc., vol. 30, 1904, pp.
287-293.--BARNES and McDunnovueH, Check List Lepid. Bor. Amer.,
no. 6779, 1917.—ForBes, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 473.

The common American grape berry moth of economic literature.
For a number of years, until Kearfott pointed out its distinctness, it
had been confused with the European botrana. The two species
differ radically i. both genitalia (compare figs. 358, 362) and vena-
tion; botrana having vein 10 of fore wing rather close to 9 at base,
while viteana has it well separated, if anything nearer 11.

Male genitalia with weak spining on base of sacculus of harpe;
spine clusters Spc! and Spc? strongly developed, arch between them
narrow; a third cluster consisting of two heavy, flat, sword-like,
closely grouped spines arising from harpe at base of cucullus;
xedoeagus with small projecting tooth on upper edge near apex.

Male and female genitalia figured from reared specimens in Na-
tional Collection from Northeast Pennsylvania (“ X—16,” Cushman).

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 91

Distribution —Massachusetts, New York, New Jersey, Pennsyl-
vania, Maryland, District of Columbia, Michigan, Missouri.

Alar expanse.—10-14 mm.

Types——In Academy Natural Science (viteana); Museum Com-
parative Zoology (vitivorana).

Type localities—Pennsylvania (viteana) ; Hudson, Ohio (vitivor-
ana).

Food plant.—V itis.

3. POLYCHROSIS MONOTROPANA, new species
(Fig. 366)

Antenna, palpus, and face brownish ocherous; basal joint of an-
tenna blackish above. Head and thorax brownish ocherous dusted
with purplish scales. Fore wing with costa markedly rounded near
apex; basal and antemedian areas metallic blue; a faint, narrow,
scarcely angulate, black band indicating outer margin only of usual
basal patch, disappearing toward dorsum; antemedian area with
central black gemination and a couple of faint white geminations on
costa, otherwise unmarked; median band wider on costa than dor-
sum, brown heavily dusted with black; outer costai spots broad,
brown dusted with black; subapical spot large, broadly oval, con-
nected at middle with termen and joining first and second outed
costal spots above, brown heavily dusted with black; triangular
pretornal spot brown, well dusted with black; cilia dark metallic
purple. Hind wings blackish fuscous; cilia white with very dark
basal band.

Male genitalia of type figured. Harpe with short tuft from base
of sacculus; spine clusters Spct and Spe? well developed, arch be-
tween them wide, aedoeagus smooth.

Female genitalia as in spiraeifoliana except genital plate some-
what larger and stouter.

Alar expanse.—9.5-10 mm.

Type—tIn American Museum.

Paratypes.—Cat. No. 28029 U.S.N.M.

Type locality —Cincinnati, Ohio.

Food plant.—Monotropa uniflora (larvae in seed capsules).

Described from male type from the type locality (A. F. Braun,
“ VITI-24-07”); and two female paratypes from Cabin John
Bridge, Md. (R. M. Fouts, Aug. 22, 1923), all reared.

In genitalia not to be distinguished from the following species,
but with different wing shape, darker hind wings, and slightly dif-
ferent fore wing venation (8 and 9 more closely approximated at
base).

54346—26—_7

92 BULLETIN 182, UNITED STATES NATIONAL MUSEUM

4. POLYCHROSIS CYPRIPEDIANA Forbes
(Figs. 15, 367)

Polychrosis cypripediana Forses, Memoir 68, Cornell Uniy. Agr. Exp. Sta.,
1924, p. 473.

Like the preceding (monotropana) in color, pattern, and genitalia,
but with paler hind wings and less black dusting on dark areas of
fore wing. There is little or no black scaling on subapical spot,
and the latter is considerably paler than the median band. Veins
8 and 9 of fore wing while not far apart at base are still distinctly
separate and farther apart than 7 and 8. The termen is also
straighter near apex. In pattern, color, and wing structure it is
still closer to rhoifructana, but is easily distinguished from the latter
by the male genitalia having a short tuft on base of sacculus.

The female genitalia are as in monotropana.

Male genitalia of type figured

Alar expanse.—9-10.5 mm.

Type.—tIn American Museum.

Paratypes.—Cat. No. 28030, U.S.N.M. Also in American Museum
collection Barnes and Canadian National Collection.

Type locality —Aweme, Manitoba.

Food plant—Cy pripedium (larvae feeding on seeds).

Described from male type, 2 male and 4 female paratypes all
reared and from the type locality and dated as follows: type and 1
male and 1 female paratypes, “Jan. 14-09”; 1 male paratype, “Jan.
1-09”; 2 female paratypes, “14-IV-07”; 1 female paratype, “Jan.
1906, from larva collected 25 Aug. 1905”, (N. Criddle).

These specimens Kearfoot had set aside as a new species under the
manuscript named cypripediana, validated by Forbes and therefore
credited to him. Inasmuch as Forbes designated no types, I do so
here. Several of the specimens are in poor condition. The mid-
winter issuing dates are probably due to indoor rearing.

5. POLYCHROSIS RHOIFRUCTANA Kearfott

(Wigs. 176, 372)

Polychrosis rhoifructana Kearrott, Trans. Amer. Ent. Soc., vol. 30, 1904,
p. 296.—Barnes and McDunnovueH, Check List Lepid. Bor. Amer.,
no. 6786, 1917.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 473.

Larvae feed in the fruits of Sumac and occasionally upon seeds of
Cornus and Kalmia. Reared specimens from the last two Kearfott
had set aside as new species; but there is nothing upon which to
separate them from typical rhoifructana.

NORTH AMERICAN LASPEYRESIINAE AND OLERHREUTINAE 98

Male genitalia with long tuft from base of sacculus of harpe;
spine clusters Spct and Spc? strongly developed; arch between them
wide; aedoeagus smooth.

Male and female genitalia figured from reared specimens in Na-
tional Collection from Washington, D. C. (“3257” male type) and
Falls Church, Va. (Rhus, Hopk. U. S. No. 12196, Heinrich, female).

Distribution.—District of Columbia, Virginia, Pennsylvania, New
York, New Jersey, Rhode Island, Connecticut, Maine, Ohio.

Alar expanse—9-12 mm.

Type.—tn National Collection.

Type locality —W ashington, D. C.

- Food plants —Rhus, Kalmia, Cornus.

6. POLYCHROSIS YARACANA Kearfoit
(Fig. 370)

Polychrosis yaracana KerarrotTt, Trans. Amer. Ent. Soc., vol. 33, 1907;
p. 5.—BarneEs and McDunnoucH, Check List Lepid. Bor. Amer., no.

6784, 1917.—Forsres, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 472.

Polychrosis signifera Mryrick, Ent. Mo. Mag., vol. 48, 1912, p. 34.

Easily recognized by white hind wing with smoky shade at apex
and white scaling on fore wing. Male genitalia with long tuft from
base of sacculus of harpe; spine cluster Spct and Spc? well de-
veloped, arch between them narrow; aedoeagus smooth.

Male genitalia figured from specimen in National Collection from
Oak Station, Pennsylvania (F. Marloff, “ VI-2-12”). Female geni-
talia like those of spiraezfoliana.

Distribution.—Ohio, Pennsylvania, New York, Ontario.

Alar expanse.—9-11 mm.

Type—aIn American Museum.

Type locality. — Cincinnati, Ohio.

7. POLYCHROSIS SPIRAEIFOLIANA Heinrich
(Figs. 178, 360)

Polychrosis spiraeifoliana Hrtnricu, Proce. Ent. Soe. Washington, vol. 25,
1923, p. 106—ForBres, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,

p. 472.

Male genitalia with moderately long tuft (half the length of that
of rhotfructana) from base of sacculus of harpe; spine clusters Spc'
and Spc? well developed; arch between them angulate but wide;
sacculus but slightly rounded; aedoeagus with slight tooth on side.

Male genitalia figured from type; female from paratype in Na-
tional Collection from Hampton, N. H. (S. A. Shaw, “ 8-9-1905.”).

Q4 BULLETIN 1382, UNITED STATES NATIONAL MUSEUM

Distribution.—Pennsylvania, New Jersey, New Hampshire.
Alar expanse.—8-10 mm.

Type.—tn collection Barnes.

Type locality.—Hazelton, Pa.

Food plant.—Spiraea salicifolia.

8. POLYCHROSIS AEMULANA, new species
(Figs. 180, 369)

Superficially like vernoniana and spiraeifoliana and hardly to be
distinguished except by genitalia. The latter, however, easily iden-
tify it. The diagnostic pattern characters (such as they are) are
given in the key.

Male genitalia with sacculus of harpe but slightly rounded, a mod-
erately long tuft from base; spine clusters Spc’ and Spe? set close
together, arch between them narrowly angulate; aedoagus with a
round spine projecting downward from lower margin at apex.

Male genitalia of type figured. Female genitalia figured from
paratype in collection Barnes from Essex County Park, N. J.

Alar expanse.—9-10 min.

Type—tIn American Museum.

Paratype-——Cat. No. 28031, U.S.N.M. Also in collection Barnes.

Type locality—Hazelton, Pa.

Described from male type and one female paratype from the type
locality (Dietz, “7-3-05”) and one female paratype from Essex
County Park, N. J. (Kearfott, June 3).

9. POLYCHROSIS VERNONIANA Kearfott

(Figs. 177, 371)

Polychrosis vernoniana Krarrott, Trans. Amer. Ent. Soc., vol. 33, 1907, p.
7.—BARNES and McDunnouGH, Check List Lepid. Bor. Amer., no. 6787,
1917.

Polychrosis ambrosiana KrarrortT, Trans. Amer. Ent. Soc., vol. 33, 1907,
p. 8—BaRnes and McDunnovuaH, Check List Lepid. Bor. Amer., no.
6788, 1917.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 472,

There are no differences in genitalia or pattern between Kearfott’s
two supposed species.

Male genitalia figured from paratype (of ambrosiana) in National
Collection from the type locality; as in spiraeifoliana except with
sacculus more angulate, basal tuft longer, and arch between spine
clusters Spc! and Spc? round rather than angulate.

Female genitalia figured from paratype (of ambrosiana) in the
American Museum.

Disiribution—New Jersey, Pennsylvania, District of Columbia,
Ohio, Missouri.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 95

Alar expanse.—8-11.

Types.—In American Museum.

Type localities —Caldwell, New Jersey (vernoniana) ; Cincinnati,
Ohio (ambrosiana.)

Food plants.—Veronia noveboracensis, Ambrosia trifida (larvae
feeding upon seeds).

10. POLYCHROSIS ARUNCANA Kearfott
(Fig. 365)

Polychrosis aruncana Kearrort, Trans. Amer. Ent. Soe., vol. 33, 1907, p.
5.—BarNneEs and McDunnoucGH, Check List Lepid. Bor. Amer., no. 6783,
1917.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 472.

Distinguished by the bright ocher yellow dusting on median band
of fore wing. Male genitalia as in spiracifoliana except basal tuft on
sacculus very short and spine clusters Spc' and Spc? set closer to-
gether; figured from paratype in National Collection.

Female genitalia as in spiraeifoliana.

Paratypes from type locality in National Collection, American
Museum, and collection Barnes.

Alar expanse.—8.5-11 mm.

Type.—tn American Museum.

Type locality—Cabin John Bridge, Md.

Food plant.—Aruncus aruncus (larvae in seeds).

11. POLYCHROSIS SLINGERLANDANA Kearfott
(Figs. 179, 359)

Polychrosis slingerlandana Krarrort, Trans. Amer. Ent. Soc., vol. 30, 1904,
p. 295.—BarNEs and McDunnoueH, Check List Lepid. Bor. Amer., no.
6782, 1917.—ForBEs, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 472.

Among Kearfott’s cotypes is one specimen (in the National Mu-
seum) collected by August Busck at Plummer Island, Md., which
does not belong here. It is a runted male of Endothenia hebesana
Walker. The rest of his material is reared and authentic.

Male genitalia figured from specimens in American Museum from
Essex County Park, N. J. (“ K-257, iss. VIII-5-02”); with very
short tuft from base of sacculus; spine clusters Spc! and Spc? well
developed; arch between them broad and rounded; aedoeagus with a
prominent emarginate projection on upper edge at apex.

Female genitalia figured from paratype in National Collection
from the type locality (“ K—257, VIII-9”’).

Specimens in the National Collection, American Museum, and
collection Barnes from New Jersey.

Alar expanse.—8-10 mm.

96 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Lype-—Iitn American Museum.
Type locality —Montclair, N. J.
_ Food plant—Eupatorium perfoliatum. (larvae in seeds and on
young leaves).
12. POLYCHROSIS CARDUANA Busck

(Figs. 181, 363)

Polychrosis carduana Buscx, Journ. New York Ent. Soc., vol. 15, 1907,
p. 1384.—Barnes and McDunnovucH, Check List Lepid. Bor. Amer., no.
6785, 1917—Torees, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 472.

Male genitalia without spine tuft from base of sacculus; spine
cluster Spe? large and heavy, arch between it and Spc* broad and
rounded; aedoeagus with short spine on upper edge near apex.

Male and female genitalia figured from reared specimens in Na-
tional Collection from Decatur, Ill. (“on thistle, July 24-31,” male),
and Hyattsville, Md. (Busck, “Aug. 06,” female paratype).

Distribution —Maryland, New Jersey, Illinois.

Alar expanse.—9-12 mm.

Type.—tn National Collection.

Type locality —Hyattsville, Md.

Food plant.—Cirsium (Carduus) (larvae on top leaves and in
flowers).

13. POLYCHROSIS BLANDULA, new species

(Fig. 368)

Resembling yarkana in its white hind wings, but otherwise quite
different.

Palpus and face sordid ocherous fuscous shaded with black. Fore
wing with a blackish basal patch; antemedian band beginning on
costa as a pair of dull metallic bars separated by a line of black
scales and continued on dorsum as a large white spot; median band
a black bar from costa to top of cell, below this ocherous, slightly
dusted at middle with blackish scales, obscure and more or less fused
with the faint ocherous subtornal spot; subapical spot large, filling
most of the terminal area, pale ocherous fuscous faintly margined
inwardly by paler ocherous shading; dorsum finely spotted with
blackish; costal spots faint, blackish to beyond middle, toward apex
brownish ocherous somewhat dusted with black; a blackish apical
spot and some slight dusting of black on termen below apex; cilia
leaden with a blackish basal band broken with white scaling at
tornus. To the naked eye the fore wing shows a blackish triangular
shade on basal half extending from dorsum at basal third to outer
third of costa and including the basal patch, the costal half of the
antemedian and median bands and the first of the outer costal spots;

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 97

the rest of the wing, except for the strongly contrasted white spot
forming the dorsal half of the antemedian band, is a rather pale
ocherous fuscous. Hind wing white with a dark, smoky shade
toward apex and termen; cilia white with a dark basal band.

Male genitalia with a short tuft-from base of sacculus of harpe;
spine clusters Spe and Spe? strongly developed; arch between
them wide; aedoeagus with a rather prominent tooth from lower
surface at middle; genitalia of type figured.

Alar expanse.—15.5 mm.

Type.—tn Canadian National Collection.

Type locality —Aweme, Manitoba.

_ Described from unique type (N. Griddle, “9-VI-1921”). A
striking form easily recognized by its large size, white hind wings,
and peculiar markings.

14. POLYCHROSIS CYCLOPIANA, new species
(Figs. 185, 364)

Palpus, face, head, and thorax ashy gray. Fore wing with basal
two-thirds ashy gray very faintly cross marked with fuscous scaling
and with outer margin of the gray area inwardly concave; outer
third whitish ocherous with a large, round, black subapical spot
occupying the center of the field; this spot nowhere touching termen ;
from costa near apex, and lying between subapical spot and termen,
two fine blackish lines running to termen; on dorsum near tornus a
small triangular blackish spot; cilia ashy gray with very slight
fuscous dusting. Hind wing smoky fuscous; cilia leaden fuscous
with slightly darker basal band.

Male genitalia figured from paratype in Canadian Collection;
female from type.

Alar expanse.—12.5-14 mm.

Type.—Cat. No. 28032, U.S.N.M.

Paratype.—In Canadian National Collection.

Type locality —Brown’s Mills, N. J.

Food planis—Magnolia virginiana, M. glauca.

Described from female type reared July, 1920, by Harry B. Weiss
from larva feeding in seed pod of Swamp Magnolia and male para-
type from the Canadian National Collection labeled: “20-VII-24.
Larva on imported Magnolia glauca Linnaeus presumably from
vicinity of Philadelphia.”

A striking species. In pattern like no other American Polychrosis.

4. AHMOSIA, new genus
(Figs. 58, 186)

Genotype.—Ahmosia galbinea, new species (North America).
Thorax with posterior tuft.

98 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Fore wing smooth; termen straight; 12 veins, all separate; 7 to
termen; 8, 9, and 10 approximate; upper internal vein of cell from
between 10-11; 3, 4, and 5 not approximate at termen; 2 from cell
at 24, straight.

Hind wing with 8 veins; 6 and 7 approximate toward base; 3 and
4 separate; 5 approximate to 4; termen very slightly concave below
apex; male without chitinous ridge at inner margin.

Hind tibia of male with short hair pencile from base.

Male genitalia with harpe narrow, moderately elongate; outer sur-
face unspined; cucullus narrow, tapering, strongly spined; sacculus
narrow, weakly spined at base; spine cluster Spc produced slightly
from base of cucullus; spine cluster Spc? absent. Uncus strongly
chitinized, stout, tapering, curved, apex bluntly pointed. Socii ab-
sent. Gnathos heavily chitinized and solidly fused with anellus.
Aedoeacus long, slender, straight; cornuti absent.

Female genitalia without signum. Ductus bursae rather short,
simple.

In venation (except that vein 5 of hind wing is further separated
from 4 than 4 is from 3) this genus agrees with Polychrosis. The
latter, however, has very different genitalia. I note a similar sepa-
rate condition of veins 3 and 4 of hind wing in some specimens of
the European euphorbiana Freyer and lacunana Duponchel; but
these two have rather typical Olethreutes genitalia. Such discrepen-
cies but show the instability of structure in this family and the utter
impossibility of properly grouping the species upon any one set of
characters.

KEY TO THD SPECIBS OF AHMOSIA

1. Median dark band of fore wing broad on dorsum; alar expanse 14 mm. and

CBS ee se Mm aa Hie eRe pp a eC ae (1) galbinea.
Median dark band narrow on dorsum; alar expanse under 13 mm.

(2) aspasiana.
1. AHMOSIA GALBINEA, new species

(Figs. 58, 186, 386)

Palpus whitish ocherous; second joint with two fuscous spots on
outer side and a very slight fuscous shading toward apex; terminal
joint pale. Face and head whitish ocherous or very pale yellow.
Thorax brownish ocherous with very little paler scaling. Fore wing
sordid whitish ocherous, with the usual pattern markings pale brown
or ocherous fuscous somewhat dusted with blackish; basal patch
reaching to costa, outwardly angulate and deeply excavate below
middle; antemedian pale area broad and faintly streaked vertically
with pale ocherous fuscous; merdian band broad, irregular and rather
poorly defined, strongly dusted with black on costa and at middle;

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAB 99

subtornal spot triangular, rather pale and partially fused with
median band but distinguishable; subapical bar pale fuscous, con-
stricted at middle, swollen toward apex and base and not reaching to
costa; outer costal spots rather large but faint; some faint black
scaling on subapical bar and a few (3 or 4) blackish spots along
termen; cilia brownish ocherous, whitish at tornus. Hind wing
pale smoky fuscous; cilia whitish with dark basal band.

Male genitalia of type figured. Female genitalia figured from
paratype in National Collection.

Alar expanse.—14-20 mm.

Type—In American Museum.
| Paratype.—Cat. No. 28033, U.S.N.M. Also in American Museum,
Canadian National, and Barnes collections.

Type locality —Eureka, Utah.

Described from male type, 10 male and 1 female paratypes from
the type locality (May to August, Tom Spalding); 1 male and
1 female paratypes from Vineyard, Utah (‘“ VIII-7-18” and
“VII--8-12”) ; 3 male paratypes from Deer Creek, Provo Canyon,
Utah (“ VII-16-18” and “IX-—11-18,” Spalding) ; 1 male paratype
from Denver, Colo. (Oslar); 2 female paratypes from Chimney
Gulch, Golden, Colo. (Oslar, May) ; 1 female paratype from Almota,
Washington (C. V. Piper); 2 male and 1 female paratypes from
Jemez Springs, N. Mex. (“June 8-15”); 1 female paratype from
Mesilla, N. Mex. (C. N. Ainslie); 1 male paratype from Clark
County, Nev. (“June 24-30”); and 1 male paratype from Saska:
toon, Saskatchewan (Kenneth M. King, “ 21-IX-1922”).

This has been for some time in the collections as an undescribed
species and part of the above series had been set aside by Kearfott
under a manuscript name. The description, however, was never
published. Superficially it looks like a large, rather pale Poly-
chrosis.

2, AHMOSIA ASPASIANA (McDunnough)
(Figs. 184, 385)
Argyroploce aspasiana McDunNovuGH, Can. Ent., vol. 54, 1922, p. 44.

' Olethreutes aspasiana Forses, Cornell Univ. Agr. Exp. Sta., 1924, p. 453.

Smaller and darker than the foregoing, with smaller genitalia
and somewhat slenderer uncus.

I have seen only the type material in the Canadian National
Collection.

Genitalia (male and female) figured from paratypes from the type
locality.

Alar expanse.—11 mm.

Type.—tn Canadian National Collection.

Type locality—Mer Bleue, Ottawa, Ontario.

54346—26—_8

100 BULLETIN 182, UNITED STATES NATIONAL MUSEUM

5. Genus ENDOTHENIA Stephens
(Fig. 48)
Endothenia SterHENS, List Brit. Animals, pt. 10, Lepid., 1852, p. 28.
Genotype.—Tortrix gentianana Hiibner (Europe).
Orthotaenia STAINTON (not Stephens), Man. Brit. Butterflies, and Moths,
vol. 2, 1859, pp. 260, 262.
Genotype.—Tortriz antiquana Hiibner (Europe).

Thorax with posterior tuft.

Fore wing smooth; termen straight or slightly convex; 12 veins;
all separate; 7 to termen; 8 and 9 approximate at base; upper in-
ternal vein of cell from between 10-11; 3, 4, and 5 not approximate
at termen; 2 from cell before 24, straight.

Hind wing with 8 veins; 6 and 7 stalked; 3 and 4 connate; 5
remote from 4 at base; in male with or without chitinous ridge on
inner margin.

Hind tibia of male with or without hair pencile.

Male genitalia with harpe slender and broadening considerably
toward base; outer surface unspined; cucullus narrow, elongate;
saccullus weakly spined toward base; spine cluster Spe’ upon a
raised prominence projecting toward costa; spine cluster Spc? not
developed. Uncus long, stout, curved; tip spatulate and strongly
spined above and beneath. Socii reduced (fusing with tegumen
and defined chiefly by hairy tufting); scarcely flexible; strongly
haired. Gnathos absent, or represented only by a pair of weakly
chitinized lateral arms branching from well back on tegumen and
connecting with anellus. Aedoeagus short, stout, usually as broad
as long; penis with or without cornuti, latter when present an irreg-
ular cluster of several stout, moderately long spines.

Female genitalia with single signum; latter developed as a squa-
mous sack. Ductus bursae moderately long; chitinized only toward
genital opening.

A genus easily recognized by its characteristic male genitalia.
The hind wing venation (5 remote from and nearly parallel with 4)
would seem to place it in the Laspeyresiinae; but the genitalia for-
bid this. In pattern and general habitus some of the species are
strikingly similar to those of Polychrosis. The genitalia in some
respects resemble very much those of Bactra, with which, however,
it does not appear to have any very close connection. It is most
closely related to Zaniva, Tia, and Hulda, from which it differs
chiefly in male genitalia and the remoteness of veins 5 from 4 in hind
wing.

KEY TO THE SPECIES OF ENDOTHENIA

1. Fore wing with a distinctly whitish post median area____----------_--- 2.
Fore wing sometimes with pale post median area, but latter not white___ 4.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE lO]

2. Outer margin of dark area of fore wing nearly vertical (slightly angulate

outwardly at middle) ; extending no farther out on dorsum than on costa.

(4) melanosticta.

Outer margin of dark area decidedly irregular, extending farther out on

GLOTHSTEU EN G8 OREN SACO) OWE) Sy oe NER es Se a ee ee: Se 3.

3. Outer costal and subapical dark markings nearly obsolete_-_ (1) montanana.
Dark costal spots, subapical bar, and dark terminal shading distinct.

(2) rubipunctana.

4. Dark markings of fore wing conspicuous as a blackish, more or less triangu-

lar subapical bar, a blackish subtornal spot, and a broadly triangular or

hooklike black patch at end of cell reaching to costa but darkest below

middle (representing remains of a median band)__(9) antiquana nubilana.

Darkemnarkings, otherwise: <2 ea el Fh 5.

5. Ground color of fore wing pale sordid ocherous; conspicuous dark markings

confined to a blackish smudge on mid costa, a black apical spot and a fine

Dlaekesubapicalebart 22 es wees See a ere en (3) sordulenta.

Ground color Sometimes ocherous, but much suffused with dark scaling;

conspicuous dark markings otherwise____-_______--__+______________ 6.

6. Median and basal dark area of fore wing mottled or shaded with deep black

seales; dark sealing of palpi semiiridescent bluish-black_____________ Gs
Median and basal dark areas with little or no appreciable black scaling ;
EDT SCAUITT ROL, YOUU EUS COTE a we ee ern er ne ee 8.

7. Costa of fore wing arched at middle; apex evenly rounded___ (6) daeckeana.
Costa of fore wing straight beyond base; apex slightly pointed.

(5) hebesana.

8. Antemedian area of fore wing no paler than basal area; alar expanse over

BS Asami S20 e IEE). PDR) a ePL ts EM POE 2 Rents fs) 3 (8) infuscata.

Antemedian area slightly paler than basal area; alar expanse under 15 mm.

(7) conditana.

1. ENDOTHENIA MONTANANA (Kearfott)

(Figs. 190, 349)

Olethreutes nimbatana montanana KeErarrott, Bull. Amer. Mus. Nat. Hist.,
vol. 23, 1907, p. 157.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 456.

Argyroploce nimbatana montanana BARNES and McDunnoucH, Check List
Lepid. Bor. Amer., no. 6821a, 1917.

Is only superficially like némbatana.

Male genitalia figured from specimens in collection Cornell Uni-
versity from Ithaca, N. Y. (Cornell Lot no. 450, sub 434, W. T. M.
Forbes). Female genitalia figured from type. .

Specimens in National Collection and American Museum from
New York and North Carolina.

Alar expanse.—17-20 mm.

Type—In American Museum.

Type locality —Mount Graybeard, N. C.

102 BULLETIN 132, UNITED STATES NATIONAL MUSEUM
2. ENDOTHENIA RUBIPUNCTANA (Kearfott)
(Figs. 197, 356)
Olethreutes rubipunctana KEARFOTT, Trans. Amer. Ent. Soe., vol. 33, 1907,
p. 14: 5
Argyroploce rubipunctana Barnes and McDunnoucH, Check List Lepid.
Bor. Amer., no. 68238, 1917.

Similar to the European gentianana Hiibner, but with uncus much
narrower at apex.

Hind tibia of male without hair pencile.

Male and female genitalia figured from specimens in National
Collection from Shasta Retreat, California (“June 16-23,” male)
and Colorado (female paratype).

Distribution—Washington, California, Nevada, Colorado, Ari-
zona.

Alar expense.—17-22 mm.

Type.—tn American Museum.

Type locality.—Colorado.

Food plant.—Seeds of Iris.

3. ENDOTHENIA SORDULENTA, new species

(Figs. 196, 352)

Antenna with basal joint bluish black. Palpus with outer side
and end of second joint and entire third joint blue black, otherwise
whitish ocherous. Face whitish; overhanging scale tuft above, bluish
black. Head and thorax brownish ocherous with a scattering of
black scales. Iore wing pale sordid ocherous marked with brown and
black; basal patch very faintly indicated by fine cross lines of brown
or blackish scales; a black smudge on midcosta continuing as an ir-
regular fine blackish line nearly to dorsum and representing the re-
mains of the usual median bar; outer costal spots pale brown; sub-
apical bar black, narrow, evenly curved, touching neither termen nor
costa, connecting above with a straight, narrow, faint, dark band
which extends to dorsum before tornus; a black spot at apex; cilia
whitish ocherous somewhat spotted with bluish black toward ends.
Hind wing sordid whitish shading to pale brownish fuscous toward
outer margin; cilia whitish with faint dark basal band.

Hind tibia of male with yellow hair pencile from base.

Male and female genitalia figured from paratypes in National Col-
lection from Clear Creek and Chimney Gulch, Colo.

Alar expanse.—18-22 mm.

Type.—tIn American Museum.

Paratypes—Cat. No. 28034, U.S.N.M. Also in American Museum,
Canadian National, and Barnes collections.

Type locality —Denver, Colo.

“NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 103

Described from male type from Denver (Oslar “6—-7—7”’) ; 1 male
and 1 female paratypes from Clear Creek (Oslar); 1 male and 3
female paratypes from Chimney Gulch near Golden (Oslar, July) ;
and 1 female paratype from Berkeley (Oslar) ; all Colorado local-
ities.

This series Kearfott had set aside under a manuscript name as a
new species. It 1s possibly only a local race of rubipuncta.

4. ENDOTHENIA MELANOSTICTA (Walsingham)

(Figs. 191, 353)

Penthina melanosticta WALSINGHAM, Trans. Ent. Soc. London, 1895, p. 500.

Olethreutes melanosticta FERNALD, in Dyar List N. Amer. Lepid., no. 5048,
1903.

Epinotia flavillana DyAr, Proc. Ent. Soe. Washington, vol. 5, 1903, p. 230.

Argyroploce melanosticta BARNES and McDuNNovuaeH, Check List Lepid.
Bor. Amer., no. 6858, 1917.

Enarmonia flavillana BARNES and McDunnoucH, Check List Lepid. Bor.
Amer., no. 7165, 1917.

The largest American specics in the genus. The male has a yellow
hair pencile on the hind tibia, the female a dusky border along
termen on underside of hind wing. In the male the underside of the
hind wing is pale throughout.

Male and female genitalia figured from specimens in National Col-
lection from Yellowstone National Park, Wyo. (male), and Wil-
liams, Ariz. (female, type of flavillana).

Distribution.—Colorado, Arizona, Utah, Wyoming, California.

Alar expanse.—25-30 mm.

Types.—In British Museum (melanosticta) ; National Collection
(flavillana).

Type locatities—Larima County, Colo. (melanosticta) ; Williams,
Ariz. (flavillana).

5. ENDOTHENIA HEBESANA (Walker)
(Figs. 188, 350)

Sciaphila hebesana WALKER, Cat. Lepid. Heter. Brit. Mus., vol. 28, 1863,
p. 342.

Carpocapsa inexpertana WALKER, Cat. Lepid. Heter. Brit. Mus., vol. 28,
1863, p. 394.

Penthina fullerea Rivey, Amer. Ent., vol. 2, 1870, pp. 204, 371.

Penthina hebcsana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol. 4,
1879, p. 31.

Olethreutes hebesana FERNALD, in Dyar List N. Amer. Lepid. no. 5038,
1903.—Kerarrort, Proc. U. S. Nat. Mus., vol. 28, 1905, p. 850; Can. Ent.
vol. 37, 1905, p. 207.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 458.

Arguroploce hebesana Barnes and McDunNovuGH, Check List Lepid. Bor.
Amer., no. 6827, 1917.

104 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

A very common species throughout the United States. Male with
yellow hair pencile on hind tibia.

Male and female genitalia figured from specimens in National
Collection from New Brunswick, N. J. (H. B. Weiss, Oct., 1921,
male) and Norfolk, Va. (female).

Distribution —Massachusetts, New York, New Jersey, Pennsyl-
yania, Maryland, District of Columbia, Virginia, North Carolina,
Florida, Texas, Ohio, Indiana, Missouri, California, British Colum-
bia, Alberta, Manitoba, Ontario.

Alar expanse —11-17 mm.

Types—In British Museum (hebesana, inexpertana); National
Collection (fullerea).

Type localities—‘ North America” (hebesana, inexpertana) ;
Missouri? (fullerea).

Food plants—Antirrhinum, Gerardia, Iris, Orthocarpus, Pen-
stemon, Physostegia, Solidago, Stachys, Tigridia, Verbascum, Ver-
bena (larva feeding on seeds).

6. ENDOTHENIA DAECKEANA (Kearfott)
(Fig. 351)

Olethreutes daeckeana Kearrott, Trans. Amer. Ent. Soc., vol. 33, 1907,
p. 12—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 458.

Argyroploce daeckeana Barnes and McDunnoueH, Check List Lepid. Bor.
Amer., no. 6829, 1917.

Very similar to hebesana; but somewhat larger, with fore wing
broader and more rounded at apex, and with slightly larger geni-
talia. The female genitalia is hardly different from that of hebesana.
The male has a yellow hair pencile on the hind tibia and like hebe-
sana has the underside of hind wing rough scaled along vein 1b.

- Male genitalia figured from paratype in National Collection.

I have seen only specimens of the type series. Paratypes are in the
American and National Museums and the Barnes collection.

Alar expanse.—17-19 mm.

Type—Iin American Museum.

Type locality —Toms River, N. J.

Food plant.—Sarracenia purpurea (larvae boring in stalks and

flowers).
7. ENDOTHENIA CONDITANA (Walsingham)

(Fig. 354)
Penthina conditana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol. 4,
1879, p. 31.
Olethreutes conditana FERNALD, In Dyar List N. Amer. Lepid., no. 5037,
1903.

Argyroploce conditana BaRNEs and McDuNNouGH, Check List Lepid. Bor.
Amer., no. 6826, 1917.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 105

There are a couple of cotypes of this species in the National Col-
lection. Nothing else that I have seen exactly matches it. Of the
specimens in the Kearfott collection under the name, two females
from New Jersey and Arkansas are possibly it. They have the hind
wing cilia much paler than those of the cotypes. There is also a male
in the National Collection from Hessville, Indiana (A. Kwiat,
“'V-8—14”) which may be a variety. The genitalia are very like
those of conditana, but the pattern is more that of infuscata. These
differences may not be significant as conditana is probably as vari-
able as antiquana. Both conditana and infuscata are without cor-
nuti and have the aedoeagus extended at apex into a thin chitinous
tongue. Both also lack the hair pencile on male hind tibia.

Male genitalia figured from cotype.

Alar expanse.—10-12 mm.

Type—tIn British Museum.

Type locality Mendocino County, Calif.

8. ENDOTHENIA INFUSCATA Heinrich
(Fig. 355)

Endothenia infuscata HEINRICH, Proce. Ent. Soc. Washington, vol. 25, 1923,

p. 109.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 459.

-Closest to conditana Walsingham, but with termen of fore wing

less slanting and with stouter genitalia. The pattern is also more
like that of a suffused, brownish antiqguana.

There are two male paratypes from Ithaca, N. Y., in the collec-
tion at Cornell University. I do not know the female. In the Ca-
nadian National Collection there are also a couple of specimens that
I take to be this species. They differ somewhat in pattern, being less
entirely suffused and having more black dusting on median band of
fore wing; but the genitalia agree with those of infuscata. They
may represent a distinct race but at this time I do not feel justified in
naming them as such.

Male genitalia is figured from type.

Alar expanse.—17-20 mm.

Type.—In National Collection.

Type locality— Forest Glen, Md.

9. ENDOTHENIA ANTIQUANA NUBILANA (Clemens)
(Figs. 192, 357)

Siderea? nubilana CLEMENS, Proc. Ent. Soc. Philadelphia, vol. 5, 1865, p.
140.—Guide Study Ins. 1869, p. 333.

Sericoris vetulana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol. 4,
1879, p. 32.

106 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Olethreutes nubilana FrrNauLp, in Dyar List N. Amer. Lepid., no. 5050,
1903.—KeEaRFoTT, Can. Ent., vol. 37, 1905, p. 207.—Forsrs, Memoir 68,
Cornell Univ. Agr. Exp. Sta., 1924, p. 459.

Olethreutes vetulana FERNALD, in Dyar List, N. Amer. Lepid., no. 5051,
1903. -

Argyroploce nubilanad BARNES and McDunnovueH, Check List Lepid. Bor.
Amer., no. 6861, 1917.

Argyroploce nubilana vetulana Barnes and McDunNnoucH, Check List
Lepid. Bor. Amer., no. 6861a@. 1917.

There is only one difference between our form and the European
antiquana: a few more short spines (8-12) on the inner side of uncus
at apical margin. In the European specimens before me they num-
ber 4 to 6; but I doubt very much if the character is constant. None
of our American specimens, however, show so few. For this reason
Iam keeping nubilana as a racial designation. There is no difference
in female genitalia. Walsingham’s vetulana is smaller than most
western nubilana, but the same size as typical eastern specimens. A
cotype of the former is in the National Collection. Our variety, as
well as the European, shows considerable variation in color, and dif-
ferent American specimens from any given locality vary consider-
ably more in size. I have seen runted specimens as small as 13 mm.
expanse.

Male without hair pencile on hind tibia; penis with several (8-14)
stout cornuti.

Male and female genitalia figured from specimens in National Col-
lection from Oak Station, Pennsylvania (F. Marloff, “June 14-06,”
male) and Aweme, Manitoba (Criddle, female).

Distribution—Maryland, Pennsylvania, Illinois, lowa, Wisconsin,
New Mexico, Colorado, California, British Columbia, Alberta, Mani-
toba, Saskatchewan, Quebec.

Alar expanse.—15-22 mm.

Types.—In Academy Natural Science (nubélana) ; In British Mu-
seum (vetulana).

Type localities—Pennsylvania? (nubiélana) ; northern California
(vetulana).

Food plants.—Stachys, Mentha, Symphytum (In roots and lower
parts of stems. European records. No American rearings).

6. TANIVA, new genus
(Figs. 50, 189)

Genoty pe.—Lipoptycha albolineana Kearfott (North America).

Characters as in /'ndothenia except:

Hind wing with vein 5 bent at base and less remote from 4; in
male with a slight chitinous ridge on inner margin.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 107

Male genitalia with harpe not markedly wider at base than else-
where; spine cluster Spc? present as a row of fine spines upon a rigid,
triangular projection from neck. Uncus reduced; bifid; apex un-
spined. Socii free, weak, short, flexible, drooping, finely haired.
Gnathos normal; represented by a pair of lateral arms supporting
the usual ribbon-like, weakly chitinized subanal plate. Aedoeagus
moderately long, considerably longer than broad; cornutus a single
short, weak spine.

Closely related to E'ndothenia.

TANIVA ALBOLINEANA (Kearfott)
(Figs. 50, 189, 389)

Lipoptycha albolineana Kearrott, Bull. Amer. Mus. Nat. Hist., vol. 23,
1907, p. 160.

Argyroploce abietana FERNALD, Can. Ent., vol. 40, 1908, pp. 349, 432.—
Barnes and McDunnoueH, Check List Lepid. Bor. Amer., no. 6853,
1917.

Olethreutes piceae Buscx, Proc. Ent. Soc. Washington, vol. 18, 1916, p. 151.

Hemimene albolineana BARNES and McDuNNouGH, Check List Lepid. Bor.
Amer., no. 7265, 1917.

Argyroploce piceae BARNES and McDuNnnoueH, Check List. Lepid. Bor.
Amer., no. 6864-1, 1917.

Olethreutes abietana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 453.

I have compared the types of all the above, and there is no doubt
of the synonymy. The species is locally important as an enemy
of spruce, often doing considerable damage. The larvae eat into
the leaves near their base, cut them off and web them together in a
net of silk and frass.

Male and female genitalia figured from specimens in National
Collection from Colorado Springs (male, reared May 24, 1915, under
“ Hopk. U. S. No. 13900” from Picea engelmanni) and Monument
Park, Colo. (female, reared July 15, 1915 under “ Hopk. U. S. No.
139002 from same food plant).

Distribution —North Carolina, New York, Massachusetts, Maine,
Minnesota, Michigan, Iowa, Colorado, British Columbia, Alberta,
Manitoba, Ontario.

Alar expanse.—11-15 mm.

T'ypes.—In American Museum (albolineana) ; National Collection
(abietana, piceae).

Type localities —Black Mountain, North Carolina (albolineana) ;
Amherst, Massachusetts (abietania); Monument Park, Colorado,
(piceae).

Food plants.—Picea and Abies.

108 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

7. TIA, new genus
(Figs. 53, 195)

Genotype.—Argyroploce vulgana McDunnough (North America).

Characters as in E’ndothenia except:

Hind wing with veins 6 and 7 anastomosing just beyond cell; 5
approximate to 4 at base; no chitinous thickening on inner margin
in male.

Male genitalia with a squamous pad arising from membrane over
basal opening of harpe; spine cluster Spc! absent; spine cluster
Spe? strongly developed. Uncus slender; very slightly bent; apex
pointed. Socii fusing with tegumen but very broad and defined by
heavy hair tufting. Gnathos represented only by a thinly chitinized
flat subanal plate attached to alimentary tube; lateral arms ab-
sent. Aedoeagus swollen at apex; penis squamous; cornuti absent.

Female genitalia with signum a single very weak scobinate patch:
ductus bursae short. ~

Closely related to Hndothenia.

TIA VULGANA (McDunnough)
(Figs. 538, 195, 412)
Argyroploce vulgana McDuNNouGcH, Can Ent., vol. 54, 1922, p. 46.

Paratypes of this have been deposited by Doctor McDunnough
in the Barnes and National Collections.
Male and female genitalia figured from paratypes in collection
Barnes from the type locality.
Alar expanse.—14-16 mm.
Type.—tIn Canadian National Collection.
Type locality —Nordegg, Alberta.
Food plant—Unknown (probably birch, according to McDun-
nough).
8. HULDA, new genus
(Figs. 52, 193)

Genotype—Penthina impudens Walsingham (North America).

Characters as in E'ndothenia except:

Hind wing with vein 5 approximate to 4 at base.

Male genitalia with spine cluster Spc? on harpe strongly devel-
oped. Tegument abnormally developed, folded over beneath and
hooding the anal tube. Uncus reduced to a mere vestige. Soci
long, narrow, strongly chitinized and rigid; projecting at right
angles from tegumen; unhaired. Gnathos a narrow, arched band
arising from near base of tegumen. .Aedoeagus slender, moderately
long; cornuti absent.

Female genitalia with signum a weak scobinate patch.

“NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAB 109
HULDA IMPUDENS (Walsingham)

(Figs. 52, 193, 388)

Penthina impudens WALSINGHAM, Trans. Ent. Soc. London, 1884, p. 135.
Olethreutes impudens FERNALD, in Dyar List N. Amer. Lepid., no. 5073,
1903.—ForBEs, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 456.
Argyroploce impudens BARNES and McDunnouGH, Check List Lepid. Bor.

Amer., no. 6871, 1917.

A variable species, some specimens having the whitish areas of
fore wing much more suffused with fuscous, especially toward apex,
a complete dark median band and well-marked subapical bar. Be-
tween this and the typical pale form with broken median band and
an almost obsolete subapical bar, the intergradations are very grad-
ual, so no varietal lines can be drawn.

Male and female genitalia figured from specimens in National
collection from Oak Station, Pennsylvania (“June 4-05,” male) and
Plummer Island, Maryland (Busck, May 30, 1916, female).

Distribution.—North Carolina, Virginia, Maryland, Pennsylvania,
New Jersey, Massachusetts, New Hampshire, Ontario, Manitoba.

Alar expanse.—10.5-14 mm.

Type.—tiIn British Museum.

Type locality—North Carolina.

9. ESIA, new genus
(Figs. 56, 202)

Genotype.—Olethreutes approximana Heinrich (North America).

Thorax with posterior tuft.

Fore wing smooth; termen convex; 12 veins, all separate; 7 to ter-
men; 8 and 9 closely approximate; upper internal vein of cell
from between 10-11; 3, 4, and 5 not approximate at termen; 2 from
cell at 24, straight.

Hind wing with 8 veins; 6 and 7 approximate toward base; 3 and
4 connate; 5 bent at base, but rather well separated from 4; termen
nowhere appreciably notched; inner margin simple in male.

Hind tibia of male without basal hair pencile.

Male genitalia with harpe moderately long, broadened beyond base,
with a row of long flat spines on outer surface; cucullus stout, densely
and strongly spined, apex evenly rounded; spine cluster Spc’ strong ;
spine cluster Spc? fused with spining on outer surface; sacculus
weakly spined toward base, not extended in an arch over neck.
Uncus reduced, rounded, very weakly spined. Socii absent. Gnathos
constricted and closely encircling alimentary tube toward extremity
of tegumen, well chitinized, produced into a free chitinized blade be-

110 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

neath; subanal plate well chitinized, ribbon like, fusing with anellus.
Aedoeagus moderately long, slender, curved; cornuti absent.

Female genitalia with single signum developed as an impressed
scobinate patch. Ductus bursae moderately long; strongly chitinized
at middle; straight.

Directly derived from Olethreutes. Contains, besides the Ameri-
can type, two European species, charpentierana Hiibner and spuriana
Herrich-Schaefer.

ESIA APPROXIMANA (Heinrich)

(Fig. 414)

Olethreutes approximana HEINRICH, Ins. Ins. Mens., vol. 7, 1919, p. 65.

Superficially like Hedia cyanana Murtfeldt; but structurally quite
different.

Male and female genitalia figured from type and paratype in Na-
tional Collection from the type locality (June, 1916).

Specimens in National Collection, American Museum, and New
York State Collection from New York and New Jersey. There is
also a female in the Canadian National Collection from Aweme,
Manitoba (Norman Criddle, “27—V1I-1922”) labeled; “bred from
Stiranama.”

Alar expanse.—12-13 mm.

Type.—tIn National Collection.

Type locality.—Rensselaer, N. Y.

Food plant—Lythrum (“ Loosestrife,” larvae rolling the terminal
leaves).

10. EUMAROZIA, new genus

(Figs. 60, 194)

Genotype—Grapholitha (Poecilochroma) malachitana Zeller
(North America).

Thorax with posterior tuft.

Fore wing smooth; termen convex; 12 veins, all separate; 8 and
9 closely approximate; upper internal vein of cell from between
10-11; 8, 4 and 5 not approximate at termen; 2 from cell at 34,
very slightly bent (almost straight).

Hind wing with 8 veins; 6 and 7 approximate toward base; 3 and
4 connate; 5 approximate to 4; termen slightly sinuate below apex;
in male a chitinous ridge on inner margin.

Hind tibia of male without hair pencile.

Male genitalia with harpe eucosmaform; outer surface unspined;
cucullus strongly spined toward neck; spine cluster Spc? a single
long, strong spine; spine cluster Spe? a very small short tuft; sac-
culus weakly haired toward base. Uncus nearly obsolete. Socii

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 1]]

produced, small, hairy, flexible. Gnathos normal, a simple band
with weakly chitinized subanal plate. Aedoeagus moderately long
and slender, straight; cornuti absent.

Female genitalia with two signa, developed as stout triangular,
thorn-like disks. Ductus bursae long, looped, chitinized through-
out its length.

A monotypic North American genus.

EUMAROZIA MALACHITANA (Zeller)
(Figs. 60, 194, 413)
Grapholitha (Poecilochroma) malachitana ZELLER, Verh. Zool.-bot. Ges.
Wien, vol. 25, 1875, p. 292.

Olethreutes malachitana FERNALD, in Dyar List N. Amer. Lepid., no. 5044,
1903.—WALSINGHAM, Biol. Cent. Amer. Lepid. Heter., vol. 4, 1914,
p. 252.—Forses, Memoir 68, Cornell Uniy. Agr. Exp. Sta., 1924, p. 457.

Argyroploce malachitana BARNES and McDuNnNnoucH, Check List Lepid.
Bor. Amer., no. 6854, 1917.

The well-known leaf tier on persimon. A beautiful species with
olive green and rose lavender fore wings, not to be confused with
anything else.

Male and female genitalia figured from reared specimens in Na-
tional Collection from Falls Church, Va. (Hopk. U. S. No. 12155h,
Aug. 2, 1914, male) and Smith Point, Tex. (F. H. B. “ Bottimer
No. 40,” female).

Nistribution.—District of Columbia, Maryland, Virginia, Ohio,
Illinois, Missouri, Arkansas, Lousiana, Texas, Florida, Alabama,
North Carolina.

Also recorded from Central and South America.

Alar expanse.—11.5-16 mm.

Type——tIin British Museum.

Type locality.—Missouri.

Food plant.—Diospyros virginiana.

11. ZOMARIA, new genus
(Figs. 59, 199)

Genotype.—Penthina interruptolineana Fernald (North America).

Thorax with posterior tuft.

Forewing smooth; termen convex (evenly rounded) ; 12 veins, all
separate; 8 and 9 well separated; uper internal vein of cell from
between 9-10; 3, 4, and 5 not approximate at termen; 2 from cell
at 24, slightly bent.

Hind wing with 8 veins; 6 and 7 approximately toward base; 3 and
4 connate; 5 approximate to 4; termen evenly rounded; secondary
hair tuftings in male (costal tuft, or hair pencile on base of vein 1a)
but no thickening at inner margin.

112 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Hind tibia of male without basal hair pencile.

Male genitalia with harpe moderately long; outer surface un-
spined; cucullus finely and evenly spined, apex evenly rounded;
spine cluster Spc’, stout; spine cluster Spc? a long tuft upon a more
or less produced digitus; sacculus arched over fe ok weakly spined
at base. Uncus elongate, rounded, weakly chimed Soci elon-
gate, narrow, finely haired, attached along underside of uncus.
Gnathos well developed, very broad; produced beneath into a free,
sharp, strongly chitinized blade; subanal plate not differentiated.
Aedoeagus moderately long, nearly straight; apex pointed; cornuti
absent.

Female genitalia with signum a scobinate patch (in type with
two ear-like projections). Ductus bursae moderately long, simple,
straight.

A distinct North American genus. It genitalia characters closest
to Hsia; but in pattern and general habits more like H'wmarozia.
The three species have the same color scheme; old rose with darker
purplish pattern markings, some faintly silvery scaling and more
or less ocherous dusting.

KEY TO THE SPECIES OF ZOMARIA

1. Subapical bar of forewing not continued to costa; male with strong hair
tuft on hind wing at base of costa, without strong hair pencile from base

DIP. SEIU goal Wr Peete ss eS SE a ie I Ee NE (1) interruptolineana.
Subapical bar continued as a curved band to midcosta; male without hair
tuft on hind wing at base of costa, but with a strong yellow hair pencile
frFOmMeHaAsesOL vein VakA) RIMS ARAN RIE PERRO Pep

2. Dorsum of forewing strongly suffused with yellowish from base to beyond
POUT 1 Ce Mee a a cee UST es SO sign Wy dbl Ser ay BIE ren. (2) rosaochreana.
Dorsum somewhat yellowish at base, but yellow tint abruptly terminating
SLO Te TINT Ch UN eye a ae Te ge (3) andromedana.

1. ZOMARIA INTERRUPTOLINEANA (Fernald)
(Figs. 59, 199, 408)

Penthina interruptolineana FERNALD, Trans. Amer. Ent. Soc., vol. 10, 1882,
p. 70.

Olethreutes interruptolineana FERNALD, in Dyar List N. Amer. Lepid., no.
5040, 1903.—KeEarrorTt, Ins. New Jersey, 1910, p. 540.—Forses, Memoir
68, Cornell Uniy. Agr. Exp. Sta., 1924, p. 457.

Argyroploce interruptolineana BARNES and McDunnoueH, Check List
Lepid. Bor. Amer., no. 6849, 1917.

Male and female genitalia figured from specimens in National Col-
lection from East River, Conn. (C. R. Ely, “No. 253,” “ VII-27,”
male), and Hampton, N. H. (S. A. Shaw, “ VIII-9-1905,” female).

Distribution—New Hampshire, Connecticut, New Jersey, Mary-
land, District of Columbia, Ontario.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 113

Alar expanse.—12-15 mm.
Type.—tin National Collection.
Type locality—New Hampshire.
Food plants.—Gaylussacia, Vaccinium (larva a leaf tier).
2. ZOMARIA ROSAOCHREANA (Kearfott)
(Figs. 200, 409)
Olethreutes rosaochreana KEarroTt, Trans. Amer. Wnt. Soc., vol. 33, 1907,
p. 11.
Argyroploce rosaochreana BARNES and McDuNnnoucH, Check List Lepid.

Bor. Amer., no. 6852, 1917.

-Male and female genitalia figured from paratypes in National Col-
lection and American Museum.

Specimens in National Collection, American Museum and collec-
tion Barnes from Florida.

Alar expanse.—11.5-12 mm.

Type—tIn American Museum.

Type locality.—F lorida.

3. ZOMARIA ANDROMEDANA (Barnes and McDunnough)
(Figs. 198, 410)

Olethreutes andromedana Barnes and McDunnovueH, Cont. Nat. Hist.
Lepid. N. Amer., vol. 3, 1917, p. 228.

Very close to rosaochreana, but distinguished from the latter by
the character given in the key and by differences in male and female
genitalia. The latter are shown in the figures.

Male and female genitalia figured from paratypes in National
Collection.

Specimens in National Collection, American Museum, and collec-
tion Barnes from Florida.

Alar expanse.—12-15 mm.

Type.—tin collection Barnes.

Type locality—F¥ ort Meyers, Fla.

Food plant—Andromeda.

12. Genus APHANIA Hiibner
(Fig. 54)

Aphania Hutsner, Verz. Schmet., 1826, p. 386.
Genotype.—Tortri« scriptana Hiibner (Hurope).

Brachytaenia STePHENS, List Brit. Animals, pt. 10, Lepid., 1852, p. 25.
Genotype.—Tortriz semifasciana Haworth (Europe).

Thorax with posterior tuft.

Fore wing smooth; termen straight or slightly convex; 12 veins,
all separate; 7 to termen; 8 and 9 well separated at base; upper

114 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

internal vein of cell from between 10-11; 3, 4, and 5 not approximate
at termen; 2 from cell at or beyond 24 but before 34, straight.

Hind wing with 8 veins; 6 and 7 approximate toward base; 3 and
4 connate; .termen concave below apex but nowhere appreciably
notched; in male inner margin bearing a chitinous ridge.

Hind tibia of male with strong hair pencile from base.

Male genitalia with harpe slender, elongate; outer surface un-
spined; cucullus broad toward apex, densely spined; neck long and
slender; sacculus not extended in an arch pocketing neck; spine
cluster Spc! upon a produced lobe from sacculus, strong; spine clus-
ter Spc? not developed; sacculus weakly spined toward base. Uncus
developed, simple, slightly tapering, moderately chitinized; tip
weakly spined. Socii large, oval, flexible, strongly haired. Gnathos
with strongly chitinized, semitubular subanal plate. Aedoeagus mod-
erately long; slightly curved; with posterior blind sack; cornutus a
single stout spine with a swollen base.

Female genitalia with two signa; latter developed as deeply im-
pressed scobinate patches. Ductus bursae moderately long; wrinkled
behind genital opening; strongly chitinized and bent (or looped) at
middle.

A compact genus equally represented in Europe and North
America. The genitalia, both male and female, are quite character-
istic. Specific differences are slight, but apparently constant. In
pattern most of our species are paralleled in Europe. Similar forms
on the two continents, however, are separable on structural charac-
ters. Only one species, capreana Hiibner, is common to both.

KEY TO THE SPECIES OF APHANIA

ivNo white or whitishsareas on fore; wines se. ee ee ee Ze

Fore wing with one or more white or whitish areas___-______--_______~- 3h

. Fore wing with a fine black longitudinal dash on center of disk; median

band obsolete, at least below middle; subapical band represented only

by two fine black lines not reaching costa____________ (11) infida (part).

Fore wing without black dash on disk; median band faint but complete to
dorsum; apical band faint but distinguishable and reaching to costa.

(12) removana.

3. A longitudinal white streak from extreme base through center of wing to

COPD 2Y GAR) OFLU CSSA eS r TSS hee A cata ty Sco AR A (7) albeolana.
(8) apateticana.
No such longitudinal white streak from extreme base__________-________ 4.

4, Fore wing with a pale antemedian area extending from costa to dorsum
and completely separating a dark basal patch from median dark
MAT KINGS Ss METI LAS ANE RAMS, ER A pte ES Ens Se a es eae 2 5.

Fore wing with basal two-thirds of wing very dark greyish fuscous; no
antemedian pale area except occasionally a faint whitish patch
CTD S Ga es ge ia Ee i Meg a oe 6.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 115

5. Antemedian pale area broad; median dark band complete and as well de-
fined) ony dorsum: 28) On! Costas 2 ee ee (9) deceptana.
(10) dextrana,
Antemedian pale area narrow; median dark band more or less obsolete
below cell, much stronger toward costa__-----------_ (11) infida (part).

6. A strong ferruginous dusting along terminal margin of fore wing.
(4) tertiana.
No such ferruginous dusting along termen (terminal margin dusted with
DLO WRIShe tUSCOUS)) sew ea ea ele eee ee ee te
7. Outer margin of basal dark area distinctly angulate_________________-_ 8.
Outer margin of basal dark area slanting, scarcely irregular, almost straight.
(6) strigosa.
8. Fore wing broad; slightly less than 214 times as long as broad________~_ 9.
_ Fore wing narrow; somewhat more than 2% times as long as broad___ 10.
9. Outer margin of dark basal area with a commalike white mark jutting in
near center, and slightly indented near dorsum____________ (1) capreana.
Outer margin of dark area without such white comma mark, and not in-
MEN teGanesi sOTS Ue eee ee ee ree ee EE er (2) youngana.
10. Apical white area of fore wing with dark markings (except for costal
dashes), very) faint. = 2 a ne Ce re A eee (3) frigidana.
Apical white area much restricted by dark fuscous markings_ (5) afficticia.

1. APHANIA CAPREANA (Hiibner)
(Figs. 265, 375)

Tortrix capreana HUsnNer, Samm. Eur. Schmet. Tort., 1818, fig. 250.

Antithesia capreana WILKINSON, Brit. Tort., 1859, p. 23.

Olethreutes capreana STAUDINGER and REBEL, Cat. Lepid., vol. 2, no. 1864,
1901.— FERNALD, in Dyar List N. Amer. Lepid., no. 5033, 1903.—Krar-
FoTT, Can. Ent., vol. 37, 1905, p. 207.—Forsres, Memoir 68, Cornell
Univ. Agr. Exp. Sta., 1924, p. 455.

Argyroploce capreana KENNEL, Palaeark. Tort., Lfg. 3, Zoologica, vol. 21,
Heft 54, 1913, p. 371.—Barnes and McDunnovueH, Check List Lepid.
Bor. Amer., no. 6818, 1917—McDunnovueH, Can. Ent., vol. 54, 1922,
p. 41.

Argyroploce funerea MryRIck, Exot. Microlepid., vol. 2, pt. 11, 1920, p. 350.

Apotomis capreana PiERcE and MeEtTcALFre, Genitalia of Brit. Tort., 1923,
p. 42, pl. 15.

Meyrick says of his funerea that it is probably what we have been
calling capreana in America, but that it is quite distinct. From his
description, however, it can hardly be anything else. We have two
very similar species in North America, the true capreana and youn-
gana McDunnough, distinguished by genitalia and the shape of the
outer margin of the dark area of fore wing. Both have been identi-
fied as capreana; but most of the specimens in our collection under
the name are the true capreana. The character that Meyrick gives
for funerea (that is, outer edge of dark area “obtusely angulate
inwardly slightly above middle and slightly indented near dorsum,
otherwise hardly irregular”) fits absolutely with capreana; for that
reason I am listing his species as a synonym. Our lists also give

116 BULLETIN 182, UNITED STATES NATIONAL MUSEUM

maestana Wocke as a synonym; but as this is purely European
synonymy I am omitting the name.

Male and female genitalia figured from specimens in National Col-
lection from Ottawa, Ontario (C. H. Young, “13—VII—1906,”
male), and Wellington, British Columbia (Taylor, female).

Distribution—New Hampshire, Ontario, Saskatchewan, Manitoba,
Alberta, British Columbia, Washington.

Alar expanse—17-22 mm.

Types.—Location unknown (capreana); in collection Meyrick
(funerea).

Type localities—Europe (capreana) ; Toronto, Ontario (funerea).

Food plants.—Salix, Populus.

2. APHANIA YOUNGANA (McDunnough)
(Figs. 270, 374)

Argyroploce youngana McDunNNoueH, Can. Ent., vol 54, 1822, p. 41.
Olethreutes youngana Forsres, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 455.

In general appearance very similar to capreana but distinct.
Readily separable on genitalia and the characters given in the key.

Male genitalia figured from type; female from specimen in Na-
tional Collection from Glen House, White Mountains, N. H. (“ Aug.
1-7, 1600 ft.’’).

Distribution —Minnesota, New Hampshire, Maine, Ontario, Que-
bec.

Alar expanse.—19-22 mm.

Type.—tn Canadian National Collection.

Type locality—Meach Lake, Quebec.

3. APHANIA FRIGIDANA (Packard)
(Figs. 267, 376)

Penthina frigidana Packarp, Proc. Boston Soc. Nat. Hist., vol. 11, 1867,
p. 57.

Olethreutes frigidana FERNALD, in Dyar List N. Amer. Lepid., no. 5030,
1903.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta. 1924, p. 454.

Penthina moeschleri KENNEL, Iris, vol. 138, 1900, p. 249.

Olethreutes moeschleri STAUDINGER and REBEL, Cat. Lepid., vol. 2, Add., no.
1871 ter, 1901.

Argyroploce moeschleri KENNEL, Palaeark. Tort., Lfg. 3, Zoologica, vol. 21,
Heft 54, 1913, p. 377.

Argyroploce frigidana Barnes and McDunNnovucGH, Check List Lepid. Bor.
Amer., no. 6817, 1917.

Genitalia figures from specimens in National Collection (male)
and collection Barnes (female) from Hopedale, Labrador (‘July
24-31”), These specimens agree with Packard’s type and Kennel’s
ficure of moeschleri.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 117

The specimen in the American Museum from Franconia, N. H.,
which Kearfott had under frigidana is not that species. It is in
too poor condition to be identified with certainty; but appears to be
a rather small capreana. Kearfott did not know frigidana and what
he determined as such from British Columbia and elsewhere is
capreana.**

There is a mutilated paratype of Packard’s species in the Academy
of National Science, and a cotype from the Fernald Collection in the
National Museum. In the Barnes collection there is also a male
that I think.is this species, or a race of it, from Como, Park County,
Colo. (Oslar, June).

_ Alar expanse —16.5 mm.

Types.—In Museum Comparative Zoology (frigidana) ; collection
Staudinger (moeschleri.).

Type localities—Labrador (frigidana and moeschleri).

4. APHANIA TERTIANA (McDunnouzgh)
(Fig. 377)

Argyroploce tertiana McDuNNovGH, Can. Ent., vol. 54, 1922, p. 42.
Olethreutes tertiana ForBes, Memoir 68, Cornell Uniy. Agr. Exp. Sta.,
1924, p. 455.

Male genitalia of type figured.

Represented as far as I know only by the type.

This species, afficticia, and frigidana Packard are very similar in
appearance but show enough difference in genitalia to prevent their
being lumped together. Each has a strong curved cornutus, shorter
in frigadana than in the other two species. A. afficticia has a much
stouter aedoeagus than either frigidana or tertiana; and both frigi-
dana and afficticta have a broader spine cluster (Spe') projecting
from harpe than tertiana.

Superficially ¢ertzana can be separated by the ferruginous scaling
along termen and toward apex of fore wing. There seems to be no
trace of it in the others. Otherwise the pattern is somewhat con-
fusing. The outer margin of the basal dark area of fore wing is
distinctly angulate on the left wing of the type; but on the right
wing it is slanting and straight (as in strigosa).

The name should be restricted to the male type, as the female
allotype is a different species (Olethreutes buckellana McDun-
nough).

Alar expanse—16 mm.

Type.—tIn Canadian National Collection.

Type locality —Ottawa, Canada.

11 See Dyar, Proc. U. S. Nat. Mus., vol. 27, 1904, p. 927.

TS BULLETIN 182, UNITED STATES NATIONAL MUSEUM
5. APHANIA AFFICTICIA, new species
(Fig. 378)

Palpus dark fuscous, whitish toward base and on inner side.
Lower face white. Upper face, head, and thorax brownish fuscous.
Basal two-thirds of fore wing brownish fuscous; outer margin of
this dark area vertical from costa to middle, where it bulges out to
form a blackish-brown dot on cell, and thence in a rather convex
curve to dorsum before tornus; outer area bordering basal area, white ;
apex clouded with fuscous, this dark shade merging with an ob-
scure subapical dark bar longitudinally streaked by three black
lines; between subapical bar and outer margin of basal dark area,
a broad fuscous shade extends from tornus to a point opposite cell;
toward its apex this latter is marked by two fine longitudinal black
lines; cilia fuscous with a white shading at tornus. Hind wing
rather smoky brown; cilia whitish with a dark basal band.

Male genitalia of type figured. Aedoeagus very heavy; cornutus
a stout curved moderately long spine.

Alar expanse.—17 mm.

Type.—In American Museum.

Type locality—Mount Washington, N. H.

Described from unique male type. Closest to tertiana McDun-
nough and very like that species in pattern; but with much stouter
aedoeagus and broader projecting spine cluster (Spc') on harpe.

6. APHANIA STRIGOSA, new species
(Fig. 382)

Palpus whitish, fuscous toward apex of second joint; third joint
fuscous. Face white with a transverse bar of fuscous above. Head
fuscous with some whitish dusting toward front. Thorax fuscous
with a few sordid white scales on tegula and tuft. Fore wing with
basal two-thirds dark brownish fuscous, and outer third white
much clouded with dark fuscous; outer margin of basal dark area
slanting from just beyond middle of costa to dorsum near tornus
with a slight projection upward from middle, otherwise scarcely
irregular; on costa beyond base and before middle two obscure
whitish dashes; some blackish dusting in basal dark area, most con-
spicuous as a short median streak toward outer margin and includ-
ing the projection into white area; white area mostly suffused with
dark fuscous except for a rather narrow margin bordering the basal
dark area and broadening out toward costa; four dark dashes on
outer half of costa; no definable subapical bar in dark terminal
suffusion, but near termen just above middle two short, longitudinal,

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 119

black dashes; cilia fuscous with some white scaling at tornus. Hind
wing dark smoky fuscous; cilia sordid whitish with a dark basal
band.

Male genitalia of type figured. Cornutus a straight, moderately
long, stout spine.

Alar expanse.—15 mm.

Type—Cat. No. 28035, U.S.N.M.

Type locality—Dawson, Alaska.

Described from single male received through B. Preston Clark
and dated “6-16-16.” In genitalia and pattern very similar to the
IXuropean sororculana Zetterstedt but with appreciably narrower
fore wings.

The male genitalia are similar to those of dewtrana McDunnough
but considerably smaller.

7. APHANIA ALBEOLANA (Zeller)
(Figs. 272, 383)

Penthina albeolana ZELLER, Verh. Zool.-bot. Ges. Wien, vol. 25, 1875, p.
262.

Olethreutes hartmanniana albeolana FERNALD, in Dyar List N. Amer.
Lepid., no. 5085, 1903.

Olethreutes albeolana KEARFoTT, Can. Ent., vol. 37, 1905, p. 48; Ins. New
Jersey, 1910, p. 540.——-Forspes, Memoir 68, Cornell Univ. Agr. Exp.
Sta., 1924, p. 456.

Argyroploce albeolana BARNES and McDuNnNnouGH, Check List Lepid. Bor.
Amer., no. 6816, 1917.

In this species there is a white suffusion through center of wing
fusing with the whitish apical and antemedian costal area. The
dorsum is clouded with fuscous from base to near tornus; there is
a small quadrate dark spot on costa near base, a larger dark patch
on mid costa and a distinct black dot at end of cell.

Male genitalia figured from specimen in National Collection from
Essex County Park, N. J. (“ V-21-04,” W. D. Kearfott) ; female
from specimen in American Museum from Hampton, N. H. (Shaw).

Cornutus of male a stout curved spine. In the European scrip-
tana Hibner (hartmanniana) with simile: pattern, the cornutus is
a long, thin, straight spine.

Distribution.—New York, New Jersey, Rhode Island, New Hamp-
shire, Massachusetts, Maine, Nova Scotia.

Alar expanse.—17-19 mm.

Type.—tIn British Museum.

Type locality—Massachusetts.

Food plant.—Betula (larva crumpling the leaves).

120 BULLETIN 132, UNITED STATES NATIONAL MUSEUM
8. APHANIA APATETICANA (McDunnough)
(Figs. 269, 379)
Argyroploce deceptana McDuUNNOUGH, Can. Ent., vol. 54, 1922, p. 42 (name
preoccupied). ~ .
Argyroploce apateticana McDuNNovwvGH, Can. Ent., vol. 54, 1922, p. 168.
Olethreutes apateticana Fornres, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 455.

A distinct species with the pattern of albeolana; but with a trifle
more dark dusting and with different genitalia. The latter are
similar to those of infida but smaller. Cornutus a single, long,
stout, straight spine.

Genitalia figured from type (male) and paratype (female).

Represented by the type and two paratypes in the Canadian
National Collection from Ontario and Quebec. There is also a
specimen (male) from Vavenby, British Columbia in Mr. Black-
more’s collection which I take to be this species or a western variety.
It’s genitalia are a trifle larger than those of the type, but in color
and pattern it agrees very well.

Alar expanse.—15 mm.

Type.—tIn Canadian National Collection.

Type locality—Ottawa, Canada.

9. APHANIA DECEPTANA (Kearfott)
(Figs. 268, 373)

Olethreutes deceptana Kearrott, Can. Ent., vol. 37, 1905, pp. 41, 207.

Argyroploce deceptana BarNEsS and McDuNNoueH, Check List Lepid. Bor.
Amer., no. 6819, 1917.—Forsres, Memoir 68, Cornell Univ. Agr. Exp.
Sta., 1924, p. 453.

A distinct species, somewhat variable in color. Resembles the
European inundana Schiffermiiller; but with more white dusting in
antemedian and postmedian areas of fore wing.

Male and female genitalia figured from specimens in National
Collection from Regina, Saskatchewan (male paratype) and Dick-
inson County, Minnesota (female).

Cornutus of male rather short, stout, straight.

Distribution—Minnesota, Washington, Manitoba, Saskatchewan,
Alberta.

Alar expanse.—19.5—23 mm.

Type.—tIn American Museum.

Type locality —Aweme, Manitoba.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 121
10. APHANIA DEXTRANA (McDunnough)
(Fig. 384)

Argyroploce dextrana McDuNNouGH, Can. Ent., vol. 55, 1923, p. 165.

Represented by the type material in the Canadian Collection, one
specimen (male) from London, Ontario, in the National Collection,
and a doubtful male from Aweme, Manitoba, in the American Mu-
seum. The genitalia are similar to those of albeolana Zeller, but the
cornutus is shorter. In pattern like deceptana Kearfott and super-
ficially not to be distinguished from it.

Male genitalia of type figured.

Specimens in Canadian National Collection from Ontario and
Alberta.

Alar expanse.—17-19 mm.

Type.—tin Canadian National Collection.

Type locality —Ottawa, Ontario.

11. APHANIA INFIDA, new species
(Figs. 271, 380)

A very variable species in color. Close to both removana Kear-
fott and dextrana McDonnough.

In all but two of the specimens there is a heavy dusting of white
in antemedian and postmedian areas of fore wing; an outwardly
angulate dark blackish fuscous basal patch and the costal half of
a median dark band rather well contrasted, the latter a rather broad
roughly triangular blotch including first outer costal dark spot; be-
yond it two distinct costal spots and a fine apical dash; dorsal part
of median band and subtornal spot very faint, almost obsolete.

In two males from Quebec there is no white scaling and entire fore
wing is a smoky fuscous with a faint ochreous tint on outer half;
basal patch, median bands and costal markings obsolete except for a
very faint narrow rhomboid patch on midcosta, representing the
costal half of the median band.

In all the males there is a very distinct, short, median longitu-
dinally black streak in middle of disk (cutting what would be the
middle of the median band) and two fine black lines arising from a
point on termen above tornus and curving upward and apart from
each other in the direction of, but not reaching more than half way
to costa (these correspond to the subapical bar present in many Ole-
threutinae) ; underside of fore wing semi-iridescent, smoky. Hind
wing whitish somewhat smoky toward apex and termen; underside
whitish; cilia whitish with dark basal band.

122 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Male genitalia of type figured; cornutus stout, moderately long,
hardly curved (in one specimen with a short spur from near base) ;
outer angle of sacculus a sharp right angle; spined projection (Spc')
from sacculus, truncate, as broad as long and with outer edge straight.
Female genitalia figured from paratype in Canadian National Col-
lection from Aweme, Manitoba; genital opening large; ductus bur-
sae strongly chitinized near genital opening.

Alar expanse.—18-19.5 mm.

Type and paratypes.—Cat. No. 28036, U.S.N.M. Paratypes also in
Canadian National Collection, American Museum, and collections
Barnes and Blackmore.

Type locality —St. Johns, Quebec.

Described from male type and paratype from the type locality
(“9-VI-15” and “ 12-VI-15,” W. Chagnon, nos. 41 and 53); 1 male
paratype from Toronto, Canada (Evans “9-VI-11”); 1 male para-
type from Sebec Lake, Me. (“July 1-7”); 1 male paratype from
New Hampshire (August Busck); I male paratype from Wisconsin
(Buchholz) ; 1 female paratype labeled “ Brandon, Aug.07,, J. F.”;
1 male paratype from Calgary, Alberta (“ 15-VII—06,” F. H. Wolley
Dod) ; 1 male paratype from Waterton, Alberta (July 12-1923, H. L.
Seamans); 1 female paratype from Aweme, Manitoba (“14-VII-
1922,” Norman Criddle) ; 10 male paratypes from Nordegg, Alberta
(J. McDunnough, various June and July dates, 1921) ; 1 male para-
type from Victoria, British Columbia (EK. H. Blackmore, “no. 561,”
“8 VIT-i7”); and 1 male paratype from Franconia, N. H.

12. APHANIA REMOVANA (Kearfott)
(Figs. 266, 381)

Olethreutes removana Kerarrott, Trans. Amer. Ent. Soc., vol. 33, 1907, p.
15.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 453.
Argyroploce removana BARNES and McDunnoueGH, Check List Lepid. Bor.

Amer., no. 6860, 1917.

An almost uniform slate-gray species with pattern markings very
obscure and with no black dusting or markings on fore wing.

Male and female genitalia figured from specimens in National Col-
lection from Greenwood Lake, N. J. (reared from Willow, “ VII-
24,” male), and Hessville, Ind. (A. Kwiat, “ VI-13-08,” female).

Cornutus of male a rather short and thin spine, hardly curved at
tip.

Distribution Pennsylvania, New Jersey, Indiana, Illinois, Mani-
toba, Ontario, Quebec.

Alar expanse.—16-19 mm.

Type—tIn American Museum.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 123

Type locality—New Brighton, Pa.
Food plant —Salix.

13. Genus SCIAPHILA Treitschke
(Figs. 17, 51)

Sciaphila TreitscHKE, Schmet. Eur., vol. 7, 1829, p. 233. (=Peribrosca
Gistel).
Genotype.—(Phalaena Tortriz wahlbomiana Linnaeus) =Phalaena
Tortrix branderiana Linnaeus (Europe).

Thorax with posterior tuft.

Fore wing smooth; termen convex; 12 veins, all separate; 7 to
termen; 8 and 9 approximate; upper internal vein of cell from be-
tween 10-11; 3, 4, and 5 not approximate at termen; 2 from cell
before 34, straight.

Hind wing with 8 veins; 6 and 7 approximate toward base; 3 and
4 connate; termen evenly rounded; male without chitinous ridge at
amner margin.

Hind tibia of male without hair pencile.

Male genitalia with harpe elongate, slender, broadened beyond
base; outer surface spined; cucullus broadened toward apex; neck
long and slender; sacculus not extended in an arch pocketing neck;
spine clusters Spc' and Spe? strongly developed and closely ap-
proximate; sacculus weakly spined toward base. Uncus short, broad,
lobed, rather densely spined. Tegumen narrowly elongate. Socii
almost obsolete. Gnathos with prominent, moderately chitinized,
spatulate, flattened subanal plate. Aedoeagus short, stout, expand-
ing toward apex and more or less scobinate; cornuti absent.

Female genitalia with two signa; latter developed as sharp,
elongate, narrow, strongly chitinized blade-like projections. Ductus
bursae moderately long; simple (chitinized only toward genital
opening). :

A genus closely related to Aphania. Contains one North Ameri-
can species.

Pierce and Metcalfe’? make wahlbomiana a synonym of branderi-
ana. Under the former name European authors have hitherto identi-
fied quite a different species. In consequence of which Sciaphila has
been sunk to Cnephasia. It seems strange that all should have made
such a mistake; and I am still somewhat sceptical about the cor-
rectness of the new synonymy. However, as Pierce and Metcalfe
state that they have compared the Linnaean types there is nothing
to do but to follow them and remove Treitschke’s generic name from
the Tortricidae and apply it here.

12 Genitalia Brit. Tort., 1922, pp. 15, 48.

54046—26——9

124 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

SCIAPHILA DUPLEX (Walsingham)
(Figs. 17, 51, 257, 387)

Penthina duplez WALSINGHAM, Proc. Ent. Soe. London, 1905, p. 501.

Olethreutes duplex FERNALD, in Dyar List N. Amer. Lepid., no. 5049, 1903.—
Dyar, Proc. U. 8. Nat. Mus., vol. 27, 1904, p. 925.—Kerarrortt, Can. Ent.,
vol. 37, 1905, p. 207.—ForsBes, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 456.

Argyroploce duplex BaRNES and McDuNnnouGH, Check List. Lepid. Bor.
Amer., no. 6859, 1917.— BLAcKMorE, Rep. Prov. Mus. Nat. Hist. British
Columbia, 1921, p. M-383.

Argyroploce thallasana McDuNNouGH, Can. Ent., vol. 54, 1922, p. 48.

The American form is probably only a race of the European
branderiana Linnaeus. Our specimens, however, usually have the
antemedian and postmedian areas of fore wing distinctly white.
There are also slight genitalia differences. In duplex the cucullus is
narrower toward apex, more weakly spined along the lower margin
and the aedoeagus is more strongly scobinate. From the differences
shown in genitalia among European specimens it would seem that
there are two very distinct varieties under the one name in Europe.
In order to avoid any possible confusion, I am for the present keep-
ing duplex as a separate species. Both it and branderiana are varia-
ble in pattern. Both have the same food plant and like female
genitalia.

McDunnough’s thallasana is only a dark suffused form of duplew.

Male and female genitalia figured from specimens in National Col-
lection from Kaslo, British Columbia (H. G. Dyar, “ No. 19140, on
Aspen,” male) and Scranton, Pa. (E. A. Lister, “ VI-4-1905,”
female).

Distribution —Pennsylvania, New York, New Hampshire, Ver-
mont, Wisconsin, Colorado, Utah, Nevada, California, British Co-
lumbia, Alberta, Manitoba, and Ontario.

Alar expanse.—21-27, mm.

Types.—tIn British Museum (duplex) ; Canadian National Collec-
tion (thallasana).

Type localities—Loveland, Colo. (duplex); Aweme, Manitoba
(thallasana).

Food plant.—Populus tremuloides.

14. BADEBECIA, new genus
(Figs. 48, 256)

Genotype.—Tortrix urticana Hiibner (Kurope and North
America).

Characters as in A phania except:

Forewing with vein 2 from cell before 24.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 125

Male genitalia with uncus much reduced, narrow, pointed.
Gnathos a strongly chitinized pointed arch, scobinate (set with long
spines) beneath and supporting a very thinly chitinized, flattened
subanal plate. Aedoeagus produced at apex into a long, narrow
introvertable chitinized ribbon bearing at its apex a short stout
spine.

Female genitalia with signum a single, weak scobinate patch.
Ductus bursae swollen and strongly chitinized for over one-third
its length from genital opening.

A monotypic genus closely related to Aphanza.

BADEBECIA URTICANA (Hiibner)
(Figs. 43, 256, 391)

Tortrix urticana HUsBNER, Schmet. Eur. Tort., 1800, fig. 65.

Sericoris campestrana ZELLER, Verh. Zocl.-bot. Ges. Wien, vol. 25, 1875,
p. 282.

Olethreutes urticana STAUDINGER and REBEL, Cat. Lepid., vol. 2, no. 1921,
1901.—FERNALD, in Dyar List N. Amer. Lepid., no. 5061, 1903.

Olethreutes campestrana FERNALD, in Dyar List N. Amer. Lepid., no.
5066, 1908.—Dyar, Proc. U. S. Nat. Mus., vol. 27, 1904, p. 924—
Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 452.

Argyroploce utricana Barnes and McDunnovucH, Check List Lepid. Bor.
Amer., no. 6842, 1917.—Pierce and METCALFE, Genitalia Brit. Tort.,
1922, p. 49, pl. 16.

Argyroploce campestrana BARNES and McDuNNouGH, Check List Lepid.
Bor. Amer., no. 6864, 1917.

The name urticana Hiibner has b en a long time in our lists; but
most of the specimens that have been referred to it are anything
but that species. What Kearfott and others usually had so named
was Olethreutes deprecatoria Heinrich. The true urticana was con-
fused with puncticostana, glaciana (dealbana), and cespitana (in-
strutana) and when correctly determined only appeared under
campestrana Zeller. Strange to say the synonymy of the latter was
not suspected. The European series that I have seen were also
mixed indiscriminat-ly with wmbrosana Freyer. The confusion is
excusable; because the different species are all variable and at the
same time very similar in superficial appearance. They can only
be properly separated by the genitalia which fortunately are dis-
tinctive for each of them.

Male and female figured from specimens in National Collection
from Clear Creek, Colo. (male), and Europe (female).

Distribution.—Massachusetts, Maine, New Hampshire, Wisconsin,
North Dakota, Colorado, Arizona, Montana, Oregon, Washington,
British Columbia, Alberta, Manitoba, Ontario, Quebec.

Alar expanse.—15-19 mm.

126 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Types.—Location unknown (urticana) ; In Museum Comparative
Zoology (campestrana).

Type localities—Europe (urticana); “Maine or Massachusetts”
(campestrana).

Food plants—Rubus nutkanus (Dyar record), sugar beet, cran-
berry, “rum cherry” (larvae feeding upon the larves: U. S. Bur.
Ent. records), Betula, Populus tremuloides.

15. Genus PHAECASIOPHORA Grote
(Figs. 16, 61, 233)
Phaecasiophora Grote, Bull. Buffalo Soe. Nat. Sci., vol. 1, 1873, p. 90.

Genotype.—(Sericoris mutabilana Clemens)=Sciaphila confixana
Walker (North America).

Thorax with posterior tuft.

Forewing smooth; termen convex; 12 veins, all separate; 7 to
termen; 8 and 9 approximate; upper internal vein of cell from
between 10-11; 3, 4, and 5 not approximate at termen; 2 from cell
at 24, very slightly bent.

Hind wing with 8 veins; 6 and 7 approximate toward base; 3
and 4 connate; termen nowhere notched; in male with chitinous
ridge on inner margin.

Hind tibia of male heavily tufted with scales (less so in nivei-
guttana) and with strong yellow hair pencile from base.

Male genitalia with harpe elongate, broadened beyond base;
outer surface unspined; cucullus long and narrow, evenly spined
throughout, apex bluntly pointed; sacculus simple, weakly spined
toward base; spine clusters Spc and Spe? not developed. Uncus
absent. Socii large, oval-triangular, flexible, finely haired.
Gnathos a simple, weakly chitinized band, without subanal plate.
Aedoeagus short, stout, straight; cornuti a cluster of long, slender,
deciduous spines.

Female genitalia without signum. Ductus bursae moderately long,
chitinized for part of its length, toward genital opening.

Directly derived from Olethreutes. Contains two North American
species. The male character upon which the genus was originally
established (broadly tufted hind tibia) is decidedly less obvious in
niveiguttana than the type; but is present in both species.

KEY TO THE SPECIES OF PHAECASIOPHORA

1. Median band of fore wing not extending to dorsum; subapical bar very
broad; black dusting on disk rather in horizontal streaks than otherwise.

(1) confixana.

Median band extending to dorsum; subapical bar narrow; black scaling in
vertical streaks or outlining the dark pattern markings__ (2) niveiguttana.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 127

1. PHAECASIOPHORA CONFIXANA (Walker)
(Figs. 16, 61, 233, 394)

Sciaphila conficana Waker, Cat. Lepid. Heter. Brit. Mus., vol. 28, 1863,
p. 340.

Sciaphila? perductana WaLKER, Cat. Lepid. Heter. Brit. Mus., vol. 28, 1863,
p. 341. °

Sericoris mutabilana CLEMENS, Proe. Ent. Soe. Philadelphia, vol. 5, 1865,
p. 135.

-naecasiophora mutabilana Grote, Bull. Buffalo Soc. Nat. Sci., vol. 1, 1873,
p. 90.

Phaecasiophora confirana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus.,
vol. 4, 1879, p. 36.—FERNALD, in Dyar List N. Amer. Lepid., no. 5077,
1803—BarNneEs and McDunnovucH, Check List Lepid. Bor. Amer., no.
6877, 1917.—Forzses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 459.

This and the following (niveiguttana) are very similar in pattern
and appearance. Each has a white dot in cell on outer margin of
median band, the median band excavate behind, and the subapical
bar sending a spur from middle to termen. Their genitalia are very
similar. Those of confirana (male) have the harpes slightly the
broader.

Male and female genitalia figured from specimens in National Col-
lection from Plummer Island, Md., and Falls Church, Va. (August
Busck, May).

Distribution—New Hampshire, New Jersey, Pennsylvania, Mary-
land, Virginia, District of Columbia, Texas.

Alar exwpanse—17-20 mm.

Types—tin British Museum (confiwana, perductana); Academy
Natural Science (mutabilana).

Type localities —“ North America” (confixana perductana) ; Vir-
ginia (mutabilana).

2. PHAECASIOPHORA NIVEIGUTTANA Grote
(Figs. 231, 395)

Phaecasiophora ? niveiguttana Grote, Bull. Buffalo Soc. Nat. Sci., vol. 1,
1873, p. 91.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 459.

Olethreutes niveiguttana FrrnaLp, in Dyar List N. Amer. Lepid., no.
5075, 1903.—Kerarrott, Ins. New Jersey, 1910, p. 539.

Argyroploce niveiguttana Barnes and McDunnoucH, Check List Lepid.
Bor. Amer., no. 6873, 1917.

This species has the hind tibia of the male much less tufted with

scales than confizana and for this reason had been removed from

Phaecasiophora where Grote originally placed it. The genitalia

128 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

show, however, that it must go there. It also has the secondary
character, though not so prominently as confivana. The two species
are very close.

Male and female genitalia figured ao specimens in National
Collection from Plummer Island, Md. (Busck, July, male), and
Washington, D. C. (Busck, July, female, reared from Sassafrass).

Distribution.—Massachusetts, Connecticut, New Jersey, Pennsyl-
vania, Maryland, Virginia, District of Columbia, North Carolina,
Florida, Arkansas, Missouri, Ilinois.

Alar expanse.—14-17 mm.

Type.—tn National Collection.

Type locality —Pennsylvania.

Food plants.——Sassafras (also Hamamelis according to Kear-
fott).

16. Genus EXARTEMA Clemens

(Figs. 11, 14, 18, 42, 204)

Exartema CLEMENS, Proc. Acad. Nat. Sci. Philadelphia, 1860, p. 356.
Genotype—EHvartema nitidana Clemens (North America.)

Thorax with posterior tuft.

Fore wing smooth; termen convex; 12 veins; all separate; 7 to
termen; 8 and 9 approximate at base; upper internal vein of cell
from between 10-11; 3, 4, and 5 not approximate at termen; 2 from
cell at or beyond 24 but before 34, straight or very slightly bent.

Hind wing with 8 veins; 6 and 7 approximate toward base; 3
and 4 connate; 5 approximate to 4 at base; in male with termen
more or less notched at veins 5, 1c and 16 and with inner margin
developed into an extended lobe (fig. 18).

Hind tibia of male with long hair pencile from base.

Male genitalia with harpe elongate and with a row of flat, thin,
hair-like spines along outer surface; cucullus elongate, narrow,
densely spined; a cluster of extra heavy spines at base of cucullus
(Spe+); sacculus extended in an arch over and pocketing neck;
spine cluster (Spc*) always present and frequently upon a finger-
like process (digitus) projecting from the neck; spines at base of
sacculus short and weak. Uncus developed; weakly chitinized; tip
weakly spined. Socii developed; oval; broad; hairy and flexible.
Gnathos normal, a simple band with weakly chitinized subanal
plate. (ogee is short or only moderately long; straight or very
slightly curved; cornuti normally absent, sometimes one, two, or
three short rather weak ones present.

Female genitalia with signum present or absent; if present a single
small, weak, scobinate patch. Doctus bursae moderately long; un-
chitinized except near genital opening.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 129

A compact genus represented outside of North America as far as
I know only by a few Asiatic species. It derives directly from Oleth-
yeutes. The chief character separating it from that genus is the
strongly developed basal lobe on the hind wing of the male. This is
a prolongation and folding over of the membranous area of the wing
back of vein 1a, including within the fold a chitinous ridge morpho-
logically homologous with that upon the inner margin of the male
hind wing in Olethreutes and several other Olethreutine genera. The
mere presence or absence of such a lobe (it is a secondary character
that has been separately acquired in at least three other genera)
would not be a sufficient character for either separating or bringing
together species in generic grouping, did other facts not justify such
a proceedure. As it is, the character seems to be a good one; that is,
for separating species with such a lobe from those without it, not,
however, for grouping together all species having it.

Loxoterma, Cymolomia, and F’ccopsis each has a lobe similar to
that of Lxartema; but each is a good genus. On genitalia Loxvoterma
Busck (type, Tortrix latifasciana Haworth, fig. 390), might go very
well with H'zartema. It differs strikingly, however, in venation (hav-
ing veins 3-4 of hind wing separate). Cymolomia Lederer (type,
Sciaphila hartigiana Ratzeburg) also differs in venation, having
3-4 of hind wing stalked, a character which would place it outside
the subfamily did not genitalia and habitus show it to be a true
Olethreutine. Its genitalia are quite different from those of Hwar-
tema (see fig. 396). EH ccopsis Zeller (type, H'ccopsis wahlbergiana
Zeller) is at once differentiated by the much produced and strongly
chitinized socii of its genitalia (figs. 57, 397) and the branching of
the upper internal vein of cell of fore wing from between 9 and 10.
I have noticed what appears to be a tendency to this latter develop-
ment in one species of Lxartema (monetiferanum Riley, see p. 135) ;
but here I think the occasional branching of the internal vein from
between 9-10 is a mere freak occuring in very few specimens. I have
never noticed it in any other species of H'xartema,; and wherever else
it occurs it seems to be a consistent character of real generic value.

I am holding the genus /'xwartema because I believe it represents
a natural group. All the species with the exception of a few have
strikingly similar male genitalia of a peculiar type. The exceptional
species (those of Group () correspond in genitalia rather closely
with certain species of Olethreutes and probably represent the base
of the Exartema line where it branches from the main Olethreutes
stem. There is nothing to indicate that the group had other than a
single point of origin, or that its developmental tendency is in more
than one direction from that point. General habitus shows the
several species to be closer to each other than to any Olethreutinae

130 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

without the lobe. As far as identification goes there is little diffi-
culty. There is no such uncertainty about the lobe as there often
is, for example, about the costal fold in other groups. It is either
present and fully developed or absent (that is, the elongated, folded
structure, not the mere chitinized ridge at inner angle)."? Specific-
ally the group presents more difficulty. Many of the species are so
variable in color and, differ so slightly in structure (especially those
of Group @) that it is a hard problem to define their specifie limits.
To add to the difficulty mest of them have a number of food plants,
and many have common hosts. The trouble probably is that we are
recognizing too few or too many species. If the former is the case
then we are allowing too slight a range for structural variation.
Careful and extensive rearings from known parents will have to be
made before we can be really sure of our species.

The larvae as far as known are all leaf tiers, webbing together the
terminal leaves into a rather compact tie and feeding upon the en-
closed leaves and buds. Some of the species, especially those at-
tacking the berry-bearing plants (strawberry, raspberry, etc.), are
of economic importance as enemies of cultivated plants; but none has.
ever been recorded as an especially serious pest.

For convenience of identification I have divided the genus into
three groups according to the arrangement of spine group, Spc’, on
the harpe of the male genitalia, as follows:

Group A.—Spine group, Spe*?, upon a digitus projecting from
neck of harpe near sacculus.

Group &.—Spine group, Spc’, upon a digitus projecting from neck
of harpe near cucullus.

Group C.—Spine group, Spc”, not upon projecting digitus.

The chief structural differences between close species are in the
strength, number, and grouping of the spines of spine groups Spe?
and X on the harpes of the males and in the shape and chitinous de-
velopment of the genital plates of the females. These differences are
often slight. There is, however, little or no asymmetry in any given
species and the characters seem to be good. I do not attach any
great importance to the presence or absence of the signum. When
present it is very weak, often nothing but a vestigial patch which
might easily disappear within the limits of any given species.

As photographs of male genitalia show very little in this group,
only a few, representing markedly different forms, are given here.
A caution is also necessary in connection with the drawings of the
harpes (pls. 14, 15, 16, 17, 18). These are accurate as far as struc-

13The one possible exception (Olethreutes arcuella) to the contrary notwithstanding.
Walsingham states (Ann. Mag. Nat. Hist., ser 7, vol. 6, 1900, p. 128) that Japanese and
Korean species possess a short lobe, lacking in European examples.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 131

tures are concerned and they show the characters—such as they are—
separating the various species; but they were not made to any uni-
form scale and the differences in size are not significant.

ey:

10.

KEY TO THD SPECIES OF EXARTEMA

Dark pattern markings of fore wing much broken by rather large spots of
the pale ground color, giving wing a mottled appearance.
(1) monetiferanum.

Pore (wing) othermwises 426 N67. bite ure oeriyin hic eros foinnin ag 2A

. Fore wing ashy-gray-white with a red-brown basal patch and a large red-

brown quadrate spot upon midcosta, the latter fusing with subapical bar.
(48) ferriferanum.

Horeswingvetherwise sie fahs is a lurwria seity ese? Te “od fpotegmsielt 3%,
. Hind wing markedly: whitish toward base___-_-+-~+--_+_--4_-~+--L___ 4,
Hind wing uniformly dark or pale smoky fuscous; at most only faintly
Wilitish piowanrdnbaset=b4 aga? dees eer iesia Noy oy art _fpeshes oui tert & By:
shore wing. cinnamonsbrown sted 00st sn: sai to apie! yeu (19) bicoloranum.,

(39) submissanum.
MoOreawines pia ckishehuscOus sir xe3n ee Shee kh ef aye (20) tenebricum,

. Fore wing with median vertical band more or less obliterated by a central

longitudinal pale suffusion; or with a blackish suffusion covering basal
POUT ATG Sh:O fekyyaT eS Ee Bess eo eB ea hes on ote 6.
Fore wing with median vertical band defined; usually well contrasted —
against both antemedian and postmedian paler areas; if sometimes poorly
defined upon inner margin due to dark dusting in antemedian area, al-
ways at least somewhat contrasted against postmedian pale markings_ 18.

. Fore wing with a blackish suffusion covering basal four-fifths; outer fifth

whitish, whitish ocherous or ocherous, except for blackish subapical bar.
(44) concinnanum terminanum.

Roretwing noft‘se:markedorises fiw 44.3 os oivau seit sewal bry oye %.

. Fore wing snowy white somewhat spotted and dusted with black or blackish

fuscous toward lower inner angle, on outer half of costa, and toward
fenmen vanGdodpexine Ioan eiulur oiuiigie sal fie iy a (41) malanum.,
Fore wing more or less marked with dark patches and sometimes with
considerable suffusion of whitish ocherous on disk; but never with
PERSSON yy LOW on iets ee ee Be he A eee 8.

. Fore wing with a purplish fuscous or dark grayish fuscous blotch (con-

sisting of a fusion of basal patch, dorsal portion of median band and,
often, subtornal spot) covering most of dorsum nearly to tornus____ 9.
Fore wing more or less dark shaded or spotted; but never with a strongly
contrasted dark blotch continuous from base and reaching beyond
middlecof dorsumed leliitur pipeiiamos 2ueye ola i miboretaws Syn. - 10.

. Dark patch on dorsum of fore wing extending to but not including subtornal

spot or (where latter is absent) the area usually occupied by same; male
genitalia with spined digitus projecting from neck of harpe near sacculus.
(16) clavanum.

Dark patch on dorsum extending to tornus, including subtornal spot; male
genitalia with spined digitus projecting from neck of harpe near cucullus.
(25) nigranum (part).

Subapical bar and preternal spot of fore wing both obsolete_______-__ Ad
One or both of the above marks always present and definitely outlined,
Ehouch sometimes ftaintive colored= 22 - 2-2 a a eee 12.

54346—26—— 10

132 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

life

12.

13.

14.

15.

16.

17.

18.

19.

Fore wing olivaceous drab with a strong dark brownish fuscous bar or
patch at lower inner angle, a small dark brown dot at end of cell and a
whitish ocherous shade extending from base of costa longitudinally
through cell and suffusing preterminal area_____________ (14) cornanum.

Fore wing suffused olivaceous gray with a very faint black shade extending
diagonally from dorsum just beyond base to midecosta, and with a con-
spicuous black spot at apex; no whitish median shading and no dark

mark at, endvofrcellas a helt tite 9 wines were tes meine Bisetteves (18) exoletum.
Fore wing with basal patch, a midcostal spot, subapical bar and pretornal
spot concolorous and either raw sienna yellow or soudan brown. e

(29) ochrosuffusanum.
Above markings not concolorous; or if so, blackish or olivaceous fuscous,
Mevers sichna yellow; orSoudan browns. (2-2 =e a ee 13.
Subapical bar of fore wing strongly shaded with black toward costal ex-
tremity ; appreciably darker than dark markings near base of wing; pale
terminal area of wing tinted with ferruginous______~_ (27) merrickanum.
Subapical bar not shaded with black toward extremity; no darker than
dark markings at base of wing, often paler; cilia of fore wing sometimes
ferruginous but otherwise no appreciably ferruginous shading in ter-
MINA ATCO so i oo ee Ss etre? he eel eerie eae 14.
Cilia of fore wing strongly ferruginous__________________ (28) corylanum.
Cilia of fore wing not ferruginous except perhaps very slightly so at
IG CUTTS ei a se US ee eer ere 15.
Fore wing with a fine line of white scaling following the fold, and similar
fine white streaks outlining veins 1), 1c, 2, 3, and 4; male genitalia with
spined digitus projecting from neck of harpe near cucullus.
(21) quadrifidum.
Fore wing with considerable white or whitish scaling; but more diffused
and not outlining veins as above; male genitalia with digitus projecting
fremsneckrefharpe near sacculus 2.22222 oe ee ee 16.
Thorax and lower inner angle of fore wing strongly dusted with ferruginous
or. -brown-red! isemlesa2 $i¢ = ath te tapers oils (15) inornatanum.
Thoraxvand=forecwihneinotiso dustedic #_ ei yey Tope arrogurs® ile
Costa of fore wing at base shining white; basal patch heavily shaded with
black, the blackish scaling extended diagonally to middle of costa; dark
markings upon dorsum beyond basal patch, distinctly olivaceous.
(13) punctanum.
Costa of fore wing at base whitish ocherous; basal patch concolorous with
other dark markings of wing, ocherous drab; a large ocherous drab spot
on dorsum beyond basal patch_--_-_—~-_-_-_-_ ~~ _- (17) mediopartitum.
Median band of fore wing with two well defined, outwardly projecting
teeth, upper below costa and lower from middle of band; antemedian
and postmedian pale areas sometimes whitish but never both decidedly
WihDteyittacisygte-ceriy steel ad cing rs iene Seagios ayer Dh eg eerie eeeny ieee ceed see tg aS),
Median band with no such well defined teeth; if sometimes weakly indi-
eated, then antemedian and postmedian pale areas both decidedly

BUEN Hits Cpe a eae ee ce ee 43.
A fine, short, longitudinal whitish line cutting the median band above
lower toothy: dea eee red sesh cosy. oii Sha Pyles eo os fee eR 20.

No such whitish line cutting median band; when latter is broken it is by
invading extensions of the post median pale area_____-___-____-__ pl ie

20.

21.

22.

23.

24.

25.

26.

27.

28.

29.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 133

Dark pattern markings of fore wing brussels brown; pale areas dull leaden

OR Era eae tie a ee eh ey Se anh py aed pe eee le tS (23) sciotanum.
Dark pattern markings velvety black-brown; pale areas with a decided
17 CSN (GG Fy ao Ses 7 Ree Se Ce ie Pee em ss AcE eae. 28 eh eee Soe URS (24) trepidulum.
General color of fore wing (both pale and dark areas) markedly yel-
MO Vupee ee ee Seat cartes se eh te = esorneyers rer ese sunpeys gs Ooege Petty gs guy eee epee eo et Bw
ewer AMCOl OT OLNET WIS ei Soe aS ee ie Sl 23.

Dark markings of fore wing ocherous-orange; pale areas yellow, faintly
suffused with pinkish; male genitalia with spined digitus projecting from
NeGis.OlLMAT pe NCAT SACCUIUS S225 tet Se ee es eee (2) nitidanum.

Dark markings of fore wing cadmium yellow; pale areas paler yellow with-
out pinkish suffusion; male genitalia. with spined digitus projecting from
neck iof harpe nearcuculluscse2e2e. Serpe ae) sies os (31) ferrugineanum.

Median band of fore wing with both teeth long (three or more than three
times as long as wide); invading extension of postmedian pale area
between the teeth, deep and wedge shaped, deeper than distance separ-
ating the teeth; alar expanse always over 15 mm___-~--~-~-__+_--_+ 24.

Never will all above characters. Teeth of median band normally short or
only miderately long; sometimes lower long and upper correspondingly
short; where teeth are inclined to be long the space between them is
more round or oval than wedge shaped and equal to or greater than the
depth of the invading area separating them; alar expanse either over
or under 15 mm.; when both teeth are long expanse always less than
DE UND Ne ce en ae es eee a Sek ee ae erry S eee 26.

Pattern markings of fore wing brussels brown to dark brownish fuscous.

(25) nigranum (part).

Pattern markings of fore wing paler, more olivaceous (ocherous tawney

OL | CBWHEY-O1T Viel) =e ee ees! SG eh as NCE el bn Peper ere aE Tuer ias sp ey 2 Pas),
Pattern markings of fore wing pale rust red shading to olivaceous on mid-
dorsum and toward base______~____---___- (32) fagigemmeanum (part).
Thorax ocherous-tawney with little pale dusting except upon anterior
TERE I ECON Ge 187 es IS oe SP i em yf mR Oe (22) tilianum

(47) exaeresimum.

Thorax pale or whitish ocherous faintly cross barred with dark fuscous.
(32) fagigemmeanum (part).
Median band of fore wing much darker toward costa than toward
CLOT Sen Hah 2 68 et wears pass Fens Heise dary} eg as Tice bergeyey te ieee EES oes SEE sate 24.
Median band uniformly colored or but very little darker toward costa__ 30.
Dark dusting of median band confined to a small quadrate blackish fuscous

pateh< ons Costa snooet ys hss aa Lee ee te (43) concinnanum.
Dark dusting diffused over upper half of median band__-_____________ 28.
Dark part of medium band dark brown; pale areas of fore wing somewhat
ferrucinousiior purplish “ocherouss 22 ler wae Lael ee ee 29.
Dark part of median band blackish fuscous; pale areas of fore wing whitish
OCHEEGUS OIG. Dit Tete ie eRe AEM tee 20 (12) atrodentanum.

Lower tooth of median band same length as upper, well separated from
subapical bar; male genitalia with spined digitus projecting from neck of
BATHE MEAT SACCU LUISE Ms Bees ee eS eee ea (11) footianum.

Lower tooth of median band longer than upper, connecting with subapical
band; male genitalia with spined digitus projecting from neck of harpe
Hear euecwlwas MGR. Lore series sles 2 hoes eit (34) melanomesum.

134 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

30.

31.

32.

35.

36.

37.

38.

39.

40.

41.

Basal patch of fore wing triparted (longitudinally divided into three
RD SRIE GT GG) Si ka Te ge ae i et et ee i ie ee ree 31,
Basal patch sometimes interrupted below costa, or blotted out upon costal
margin by pale shading, or vertically striated; but never longitudinally
Ga ATG GE ese es ie Fae ese dob pepe EEN Ga) Sagas eh) Sale oes thrid co Taal ByE 32.
Dark pattern markings of fore wing brown; alar expanse 20 mm. or over.
(26) hippocastanum.

Dark pattern markings olivaceous; alar expanse under 17 mm.
(3) foedanum.
(4) furfuranum.

Alariexpanse 14amm.vorilessotier isles pales prey in eons spe L 33.
Marlexpansenl5i mim Jonioverd 222 i baet streets Eo ie epi Povey ale aaa a Rie 35.
. Dark areas of fore wing tawney ferruginous______________ (40) nananum.
Dark areas of fore wing ocherous-fuScous_______---__---____-_----_.__ 34.
. Thorax faintly and narrowly lined with fuscous, anterior margin ocherous.

(5) olivaceanum.

Thorax narrowly lined with ocherous, anterior margin fuscous.
: (46) troglodanum.
Dark markings of fore wing olivaceous or ocherous-fuscous; pale markings
whitish-ocherous, more or less lined with fuscous, but with no purplish,
ferrugineous, or pinkish suffusion; teeth of median band long and widely

Spaced Sa paiber ee lye 1 epi ve ee rae reyes eben wee Ppa ee & (38) permundanum.
Color pattern otherwise; if as above, then teeth of median band short and
WISETO Wl Ya SA COC 1 Eee 2 es a a alg Ee ak a aor se a Spl). eeeaee Seis 36.
General color of fore wing strongly ferruginous______-______-___-_____ ote
General color of fore wing not markedly ferruginous___________________ 38.

Underside of hind wing paler than underside of fore wing; male genitalia
with spined digitus projecting from neck of harpe near sacculus.
(10) zellerianum.
Underside of fore and hind wings concolorous; male genitalia with spined
digitus projecting from neck of harpe near cucullus.
(83) sericoranum (part).
Ante and post median pale areas of fare wing dark leaden purple; cilia of

hind? Wing fuscotiss2a2.220be yi ntes seria ¢ (30) brunneopurpuratum.
Ante and post median areas sometimes faintly purplish, but never dark
leaden! purple; cilia of hind wing whitishi--_==~~—-----.--+_-u-1u 2: 39.

Fore wing with central vertical dark line of antemedian pale area strongly
marked and with well-defined lines of fuscous scales in postmedian pale
GU Chet hn tad a bee ego brie Sse ee ent Dyan hegre) Seep eh ce heii yee ey (9) rusticanum,

All such lines rather weak and more or less discontinuous___________~_ 40.

Antemedian pale area of fore wing markedly whitish, in some specimens
with a faint pinkish suffusion ; when the latter, then basal patch, median
band and subapical bar rather strongly dusted with black__________~_ 41.

Antemedian pale area very pale sordid ocherous-fuscous or leaden metallic
with ocherous or pinkish suffusion; when pinkish, no pronounced black
shading/onidark ‘pattern Areas 40 sf: e0_ Sern ety ey Ee ee 42.

Teeth of median band narrowly separated by a small triangular extension
of the pale postmedian area; male genitalia with spined digitus project-
ing from neck of harpe near sacculus_.______________ (8) electrofuscum.

Teeth of median band more broadly saparated by a roundish extension of
the pale postmedian area; male genitalia with spined digitus projecting
fromyneck of harpe near cucnilus22 22042 sass ees (37) brevirostratum.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 135

42. Dark pattern markings of fore wing brussels brown; male genitalia with
spined digitus projecting from neck of harpe near sacculus.

(7) subnubdilum.

Dark pattern markings of various brownish shades, but never brussels

brown; male genitalia with spined digitus projecting from neck of harpe

jE EAU CCAD NS VS ch ell aes SR ee ne EN (33) sericoranum (part).

43. Fore wing with an appreciable shading of rusty yellow on outer half, espe-
Gidllyatoward tenmene <2 .4... 2 eres eee as (45) fasciatanum.
Nomuch yellowish shading, on foretwints 2. 227 ose se eS eee 44,

44. Median band of fore wing broken below costa and above dorsum; forming
MARCO tIELe SM ar ie ATK «SOLS see eee eee ee (6) fraternanum.

MICO LAEOU ANOMCOMPLEbes au ee ene ee Se ee Mee OA ee ee a 45.
45. Dark pattern markings of fore wing brown; antemedian and postmedian
paleareas cleariiwhiteyet 2b Sei eete Set beet y a sen pani Py epee reheE 46.
Dark pattern markings dark grayish fuscous; antemedian and postmedian
pale areas rather sordid white.) 8 (42) appendiceum.

46. .Costa of fore wing at, base white-—____-._-______-___ (36) versicoloranum.
Costa of fore wing at base dark brownish fuscous______-_-~_ (35) valdanum.

Group A.—GENITALIA OF MALE WITH A SPINED Dicitus (Spc) PROJECTING FROM
NECK OF HARPE CLOSE-TO SACCULUS

1. EXARTEMA MONETIFERANUM Riley
(Figs. 70, 221)

Exartema monetiferanum Ritry, Trans. St. Louis Acad, Sci., vol. 4, 1881,
p. 317.—FERNALD, in Dyar List N. Amer. Lepid., no. 5012, 1903.—
Barnes and McDunnoucH, Check List Lepid. Bor. Amer., no. 6795,
1917.

Cymolomia monetiferana Forspes, Memoir 68, Cornell Univ. Exp. Sta., 1924,
p. 4638.

The species may be at once distinguished by the character given 11.
the key. The upper internal vein of cell in fore wing is somewhat
variable. In most specimens it is normal (that is from between veins
10 and 11). In some, however, it comes from between 9 and 10,
and in a few is entirely lost. It is interesting to note that when
there is a variation from the normal in this respect it is not the same
in both right and left fore wings of any particular specimen.

Harpe of male genitalia figured from type; female from specimen
in National Collection from New Brighton, Pa. (H. D. Merrick,
“VI-804”).

Bursa of female without signum.

Distribution—Alabama, Pennsylvania, Ohio.

_ Alar expanse.—16-21 mm.

Type—tin National Collection.

Type locality—Kufaula, Ala.

136 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

2, EXARTEMA NITIDANUM Clemens
(Figs. 42, 64, 204, 399)

Heartema nitidana CLEMENS, Proc. Acad. Nat. Sci. Philadelphia, 1860,
p. 356.

Sericoris nitidana CLEMENS, Proc. Ent. Soc. Philadelphia, vol. 5, 1865,
p. 1383.

Exartema nitidanum FERNALD, in Dyar. List N. Amer. Lepid., no. 5010,
1903.—BarNnEs and McDuNNovucH, Check List Lepid. Bor. Amer., no.
6793, 1917.

Cymolomia nitidana Forbes, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 468.

Clemens’ description of this species is misleading. He speaks of
the fore wings as reddish brown with “markings pure brown.” The
type at Philadelphia, corresponding with what we have in the col-
lection as nétidanum, has the pale areas of the fore wing yellow with
an overshading of faint rose color and the dark markings brownish
yelllow. The general color of the insect is decidedly more yellow
than brown.

Male genitalia figured from specimen in National Collection from
Vermont; female from specimen in American Museum from Cin-
cinnati, Ohio. |

Bursa of female without signum.

Distribution.—Ohio, Pennsylvania, Vermont, Ontario.

Alar expanse.—15-20 mm.

Type.—in Academy Natural Science, Philadelphia.

Type locality—Pennsylvania.

3. EXARTEMA FOEDANUM (Clemens)
(Figs. 65, 209)

Sericoris foedana CLEMENS, Proc. Hnt. Soc. Philadelphia, vol. 5, 1865,
jo Gis

Ezartema concinnanum FERNALD, in Dyar List N. Amer. Lepid., no. 5017,
1903.—BarNeES and McDuNNoveH, Check List Lepid. Bor. Amer., no.
6800, 1917.

Cymolomia foedana Foxsses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 465.

This species has been incorrectly listed as a synonym of concinna-
num Clemens. It resembles the latter somewhat in pattern but is
quite different in genitalic structure. Strangely enough specimens of
the true foedanum have always been mixed with olivaceanum Fer-
nald in our collections.

Harpe of male genitalia figured from specimen in National Collec-
tion from Ocean View, Va.; female from a reared specimen without
locality label (“ July 27, 93”), also from the National Collection.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 137

Bursa of female without signum.

Alar expanse—11-14 mm.

Type—In Academy Natural Sciences, Philadelphia.
Type locality —Virginia.

Food plant.—Blackberry.

4. EXARTEMA FURFURANUM McDunnough
(Fig. 78)

Ezxartema furfuranum McDunnovueH, Can Ent., vol. 54, 1922, p. 38.
Cymolomia furfurana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 464.
_ Very close to foedanum and probably only a larger form of that
species. I am keeping it separate for the present as the genitalia
are considerably larger than those of typical foedanum.
So far represented only by the type and paratype from Ottawa
and Quebec in the Canadian National Collection.
Harpe of male genitalia figured from type.
Alar expanse.—16 mm.
Type-—Canadian National Collection.
Type locality —Ottawa, Ontario.

5. EXARTEMA OLIVACEANUM (Fernald)

(Figs. 66, 211, 400)

Eccopsis olivaceana FERNALD, Trans. Amer. Ent. Soc., vol. 10, 1882, p. 71.
Exartema olivaceanum FERNALD, in Dyar List N. Amer. Lepid., no. 5016,
- 1903.—BarNES and McDuNnNoucH, Check List Lepid. Bor. Amer.,
no. 6799, 1917.
Heartema bolandanum McDunnoueH, Can. Ent., vol. 54, 1922, p. 39.
Cymolomia olivaceana ForBes, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 464.

This species has been badly mixed in the collections. Over half
the specimens so named were either troglodanum or foedanum. In
general appearance, pattern, and color the three are very similar.
Clemens’ foedanum can be separated by the broken basal patch of
fore wing; but olivaceanum and trogiodanum are so much alike that
they can be definitely determined only by their genitalia. The
pattern characters given in the key hold in the main and will help
to separate most specimens; but they break down in some, especially
the darker examples of troglodanum, and when the thorax is rubbed
can not be used at all. Fernald had no idea that there were two
species involved, for he frequently determined specimens of tro-
glodanum as olivaceanum. Of the so-called “homotypes” in the
Kearfott Collection, one is troglodanum and the other olivaceanum.

138 BULLETIN 132, UNITBD STATES NATIONAL MUSEUM

McDunnough’s bolandanum is a straight synonym. It was distin-
guished by the thickness of the subapical bar of fore wing and its
relative nearness to apex, characters that are extremely variable
and not to be relied upon. There are no genitalia differences be-
tween it and typical olivaceanum.

Genitalia figured from specimens in National Collection from
Buffalo, New York (William Wild, “ VII-15-09,” male) and Ottawa,
Canada (C. H. Young, “28-VI-1905,” female). Bursa of female
without signum.

Distribution—New Jersey, Pennsylvania, New York, Massachu-
setts, Maine, Ontario, British Columbia.

Alar expanse.—12-14 mm.

Types.—In National Collection (olivaceanum) ; Canadian National
Collection (bolandanum).

Type localities—Massachusetts (olivaceanum) ; Ottawa, Canada
(bolandanum).

6. EXARTEMA FRATERNANUM McDunnough

Eaartema fraternanum McDuNNoucH, Can. Ent., vol. 54, 1922, p. 38.
Cymolomia fraternana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 464.
Apparently a valid species separable from others in this immediate
group by the broken median bar of fore wing.

Represented only by the unique type.

Harpe of male genitalia as in zellerianwm Fernald.

Alar expanse.—16 mm.

Type.—tn Canadian National Collection.

Type locality. Ottawa, Ontario.

7. EXARTEMA SUBNUBILUM Heinrich
(Figs. 68, 216)

Eerariema subnubilum HEInRIcH, Proc. Hnt. Soc. Washington, vol. 25, 1923,
p. 110.
Cymolomia subnubila Forses, Memoir 68, Cornell Uniy. Agr. Exp. Sta.
1924, p. 466.
Harpe of male genitalia figured from type; female from paratype
in the American Museum from Mountain Lake Park, Md.
Bursa of female with signum.
Distribution —New Jersey and Maryland.
Alar expanse.—15-17 mm.
Type.—In American Museum.
Type locality —Greenwood Lake, N. J.
Food plant.—Hazel.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 139
8 EXARTEMA ELECTROFUSCUM Heinrich
(Figs. 14, 75, 215)
Exartema electrofuscum HrInRricH, Proc. Ent. Soe. Washington, vol. 25,

1923, p. 110.

Cymolomia electrofusca Forspes, Memoir, 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 464.

Male and female genitalia figured from paratypes in National Col-
lection; male from Center Harbor, N. H. and female from Lake-
hurst N. J.

Bursa of female without signum.

Distribution—New Jersey, New Hampshire, Massachusetts.

Alar expanse.—15-17 mm.

Type—ain American Museum.

Type locality.— Lakehurst, N. J.

Food plant.—Sweet Fern.

9. EXARTEMA RUSTICANUM McDunnough
(Fig. 79)

Exartema rusticanum McDunnoucH, Can. Ent., vol. 54, 1922, p. 38.

Cymolomia rusticana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 463

Known only from the type and paratypes in the Canadian and
Barnes collections, all from the type locality. In the male genitalia
hardly to be distinguished from zellerianwm of which it may indeed
be a western race. It lacks, however, the decidedly ferruginous tint
in pale areas of the fore wing so characteristic of Zeller’s species.

Harpe of male genitalia figured from paratype in collection Barnes.

Alar expanse.—15-16 mm.

Type.—tIn Canadian National Collection.

Type locality—Onah, Manitoba.

10. EXARTEMA ZELLERIANUM (Fernald)
(Figs 63, 203)
Herartema nitidanum ZELLER (not Clemens), Verh. Zool.-bot. Ges. Wien,
vol. 25, 1875. p. 270.
Eccopsis zelleriana FERNALD, Trans. Amer. Ent. Soe., vol. 10, 1882, p. 29.

Eerartema zellerianum FERNALD, in Dyar List N. Amer. Lepid., no. 5014,

1903, BARNES AND McDuNNovuGH, Check List Lepid. Bor. Amer., no.
6797, 1917.

Cymolomia zelleriana Forsrs, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 465.

What this name really stands for is doubtful. Zeller described
and figured what he took to be a specimen of nétidanwm Clemens.
He evidently had not seen the type and was misled by Clemens’
faulty description; for what he figured under that name was not

140 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

nitidanum but an apparently undescribed species. Fernald recog-
nized this and proposed the name zellerianum for it. At the same
time he identified as zellerianwm specimens reared from Betula alba:
If Zeller’s description and figure are accurate this identification of
Fernald’s is also an error; for the Betula specimens match neither
figure nor description. JKearfott noticed this discrepancy and after
following Fernald for some years, suddenly applied the name zed/er-
ianum to quite a different looking insect (what I am describing
further on in this paper as Lxartema trepidulum (p. 147), and gave
the manuscript name betulana to the moths which he, Fernald and
others had previously identified as zellerianum. Kearfott’s speci-
mens, however, while they agree better than Fernald’s with Zeller’s
figure, do not altogether agree with his description, and for that
reason can not be accepted without considerable question. An ex-
amination of the type in London will settle the matter, for Fer-
nald’s name must apply to the specimen identified and labeled by
Zeller as nitidanum Clemens. In the mean time, so as not to add to
the confusion, I am continuing the use of the name as Fernald ap-
plied it and assuming that he was correct in identifying the Betula-
feeding, ferruginous species as the true zellerianwm.

Male and female genitalia figured from reared specimens in
National Collection; male from East River, Connecticut (C. R. Ely,
“ VII-26-12”’) and female (“K. 84”) without locality label but pre-
sumably from New Jersey.

Bursa of female with signum

Distribution.—Pennsylvania, New Jersey, Rhode Island, Con-
necticut, Maine, New Hampshire.

Alar expanse.—16-20 mm.

Type—tIn British Museum.

Type locality—Maine.

Food plant—Betula alba.

11. EXARTEMA FOOTIANUM (Fernald)
(Figs. 76, 205)

Eccopsis footiana FERNALD, Bull. Buffalo Soe. Nat. Sci., vol. 4, 1882, p. 53.

Exartema footianum FERNALD, in Dyar List N. Amer. Lepid., no. 5062,
1903.—BarNES and McDunnoucH, Check List Lepid. Bor. Amer., no.
6809, 1917.

Cymolomia footiana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 470.

A distinct species easily recognized by the characters given in the
key.

Harpe of male genitalia figured from reared specimen in National
Collection from Staunton, Va. (Quercus, 26 June, 1919, Heinrich) ;
female from American Museum from Montclair, N. J.

Bursa of female without signum.

' NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 141

Distribution.—Virginia, District of Columbia, Pennsylvania, New
Jersey, New York.

Alar expanse.—18-22 mm.

Type.—tIn National Collection.

Type locality —New York.

Food plants—Hamamelis, Quercus.

12. EXARTEMA ATRODENTANUM (Fernald)
(Figs. 77, 217)

Eccopsis atrodentana FERNALD, Trans. Amer. Ent. Soc., vol. 10, 1882, p. 71.

Exartema atrodentanum FERNALD, in Dyar List N. Amer. Lepid., no. 5020,
1903.—KerarrottT, Can. Ent., vol. 37, 1905, p. 206—Barnes and Mc-
DunnovueH, Check List Lepid. Bor. Amer., no. 6802, 1917.

Cymolomia atrodentana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 465.

In pattern much like footianum,; but of quite a different color.
The characters given in the key will readily separate the two species.

Fernald’s cotypes represent a mixed series. The Canadian sp:ci-
mens correspond to what we have always identified as atrodentanum,
while the Texas specimens represent a new species which is described
in this paper (p. 160) as exaeresimum. I am therefore restricting
the name to the form with blackish shading on costal half of median
band, and designating one of the cotypes from Ontario as the
holotype.

Male (harpe) and female genitalia figured from specimens in
National Collection from New Brighton, Pennsylvania (H. D. Mer-
rick, ‘“ VII-28-06” and “ VITI-9-07”).

Bursa of female without signum.

Distribution.—lowa, Pennsylvania, Massachusetts, Ontario, Mani-
toba.

Alar expanse.—17-19 mm.

Type.—tiIn National Collection.

Type locality —Ontario, Canada.

13. EXARTEMA PUNCTANUM Walsingham

(Figs. 71, 212, 401)

Ezrartema punctanum WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol.
4, 1879, p. 37.—FERNALD, in Dyar List N. Amer. Lepid. no. 5019,
1903.— Barnes and McDuNnNnNovueH, Check List Lepid. Bor. Amer., no.
6801, 1917.

Sericoris versicolorana CLEMENS (not Clemens, in part) Proc. Ent. Soe.
Philadelphia, vol. 4, 1865, p. 136.

Cymolonmia punctana Forsrs, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 467.

142 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Male and female genitalia figured from reared specimens in Na-
tional Collection from East River, Conn. (male, C. R. Ely, 11 July,
1912), and Medford, Mass. (female, W. O. Ellis, 21 June, 1921).

Bursa of female with signum.

Distribution—New York, New Jersey, Massachusetts, New Hamp-
shire, Ontario, Quebec.

Alar expanse.—17-18 mm.

Type.—In British Museum.

Type locality —Shasta County, Calif.

Food plant.—Cornus.

14. EXARTEMA CORNANUM Heinrich

(Fig. 67)
Heartema cornanum HEINRICH, Proc. Hnt. Soc. Washington, vol. 25, 1923,
Dee:
Cymolomia cornana ForBES, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 467.

Harpe of male genitalia figured from type. Female unknown.

Represented in the three collections by specimens from New Jer-
sey, and in the Canadian National Collection by one specimen from
Ottawa, Ontario.

Alar expanse —17.5-17 mm.

Lype.—In American Museum.

Type locality—Essex County Park, N. J.

Food plant.—Cornus canadensis.

15. EXARTEMA INORNATANUM Clemens
(Figs. 69, 213)

Exartema inornatana CLEMENS, Proc. Acad. Nat. Sci. Philadelphia, 1860,
p. 357.

Sericoris inornatana CLEMENS, Proc. Ent. Soc. Philadelphia, vol. 5, 1865,
p. 134.
Ezartema inmornatanum FERNALD, in Dyar List N. Amer. Lepid., no. 5024.,

1903.—BARNES and McDuNNoucH, Check List Lepid. Bor. Amer., no.
6807, 1917.

Cymolomia inornatana Kerarrort, Ins. of New Jersey, 1910, p. 589.—ForBEs,
Memoir 68, Corneil Univ. Agr. Expt. Sta., 1924, p. 469.

Male (harpe) and female genitalia figured from specimens in Na-
tional Collection from Oak Station, Penn. (F. Marloff, “ VII-17-10”
and “ VII-10-10”’).

Bursa of female without signum.

Distribution.—Pennsylvania, New York, New Jersey, Connecticut,
Massachusetts, New Hampshire, Ohio, Illinois, Kansas, Texas,
Quebec.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 143

Alar expanse.—15-22 mm.

Type—tLost (7%).

Type locality —Pennsylvania.

Food plants—Prunus serotina, Cornus, Quercus, Clethra alnifolia,
Juglans (Walnut).

16. EXARTEMA CLAVANUM (Walker)
(Figs. 80, 210)

Carpocapsa clavana WALKER, Cat. Lepid. Heter. Brit. Mus., vol. 28, 1863,
p. 395.

Exartema inornatanum FERNALD (not Clemens), in Dyar List N. Amer.
Lepid., no. 5024, 1903.—Barnes and McDunnovucHu, Check List Lepid.
Bor. Amer., no. 6807, 1917.

Cymolomia clavana Yorsres, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 469.

This species has been listed for a long time as a synonym of inor-
natanum, and specimens have been most frequently placed under that
name, though I have also seen several mixed with specimens of
nigranum Heinrich. Some forms of the latter are very hard to
separate from clavanum on pattern; but can be readily distinguished
by the genitalia. Walker’s species separates readily from inor-
natanum on both structure and color. It has the same pattern, but
lacks the distinct reddish scaling on thorax and lower inner angle
of fore wing so characteristic of Clemens’ species.

Harpe of male genitalia figured from specimen in National Col-
lection from Essex County, N. J. (Kearfott, “ 7-22-99”). In general
shape and structure the male genitalia are very like those of zelleri-
anum. Minor differences in the spining of the harpes of the two
species are shown in figures 63 and 80. Female genitalia figured
from specimen in National Collection from Onaga, Kans.

Bursa of female without signum.

Distribution—Kansas, Iowa, Illinois, New York, New Jersey,
Viassachusetts, Quebec.

Alar expanse.—15-17 mm.

Type—tn British Museum.

Type locality —* North America.”

17. EXARTEMA MEDIOPARTITUM Heinrich

(Fig. 72)
Exartema mediopartitum Hernricu, Proe. Ent. Soc. Washington, vol. 25,
1923, p. 118.
Cymolomia mediopartita Forses, Memoir 68, Cornell Uniy. Agr. Exp. Sta.,
1924, p. 465.

Similar in pattern to corylanum Fernald, but lacking the ferrugin-
ous fore wing cilia of the latter. Easily distinguished by the spin-

144 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

ing of the harpe of its genitalia. So far represented only by the
type and paratype in the National Collection.

Harpe of male genitalia figured from type. Female unknown

Alar expanse.—13 mm.

Type.—tn National Collection.

Type locality.—Virginia.

18. EXARTEMA EXOLETUM Zeller
(Figs. 73, 227)

Exzartema exoletum ZeLueR, Verh. Zool.-bot. Ges. Wien, vol. 25, 1875, p. ©
270.—FERNALD, in Dyar List N. Amer. Lepid., no. 5023, 1903——BarNESs
and McDunnoueH, Check List Lepid. Bor. Amer., no. 6806, 1917.

Cymolomia exoleta Krarrort, Ins. of New Jersey, 1910, p. 538.—Forsss,
Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 469.

A striking species, easily distinguished by the suffused olivaceous-
gray fore wing with its conspicuous, strongly contrasted and en-
larged black apical spot.

Male (harpe) and female genitalia figured from reared specimens
in National Collection from Orono, Me. (June 24 and 27, 1925; bred
from currant).

Bursa of female without signum.

Distribution.—Iowa, Ulinois, Ohio, Pennsylvania, New York, New
Jersey, Maine, Ontario, Quebec.

Alar expanse.—14-16 mm.

Type—In Museum Comparative Zoology.

Type locality —Massachusetts.

Food plant.—Gooseberry, currant.

19. EXARTEMA BICOLORANUM McDunnough
Exartema bicoloranum McDuNNouGH, Can. Ent., vol. 54, 1922, p. 40.

A small, dark, obscurely marked, cinnamon-brown species easily
distinguished by its white hind wings. So far represented only by
the male type and a specimen from Nova Scotia, and a doubtful
male from Duluth, Minnesota in the National Museum. (These
last two from the Fernald Collection.)

Alar expanse.—138-15 mm.

Type.—tIn Canadian National Collection.

Type locality—Barrington Passage, Nova Scotia.

20. EXARTEMA TENEBRICUM, new species
(Fig. 74)

Plapus whitish; second joint with a blackish fuscous dot on outer
side and a blackish fuscous shading at apex; third joint blackish.
Face, head, and thorax blackish fuscous with a slight admixture of

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 145

sorid ocherous scaling, especially on face and collar. Fore wing
blackish fuscous; maculation indistinct; antemedian and postmedian
areas leaden fuscous, poorly contrasted against the slightly darker
pattern markings; latter faintly dusted with scattered dull ocherous
(in male) or rusty brown scaling (in female paratpye) ; a few faint
whitish geminate marks on costa; cilia whitish blotched with blackish
and with a black basal band. Hind wing whitish hyaline toward
base shaded with blackish outwardly; cilia white with a black basal
band; in male slight notches at veins 15 and 5. Under side of fore
wing pale leaden fuscous; costa very faintly marked with white.
Under side of hind wing hyaline white with a dark shading at apex.

Harpe of male genitalia of type figured. Female genitalia as in
subnubilum except for slightly larger signum.

Alar expanse.—15-16 mm.

Type.—tn Collection Cornell University.

Paratype——Cat. No. 28037, U.S.N.M. Also in Cornell University
Collection.

Type locality——Argus Brook, McLean Reservation, N. Y.

Described from male type (20 July, 1924); one male paratype
from “Grass Bog 3,” McLean Reservation (15 July, 1924) ; and one
female paratype from West Ridge, McLean Reservation, New York
(25 July, 1924) all received from W. T. M. Forbes.

“A distinct species closest to swbmissanum McDunnough in pat-
tern; but with quite different genitalia. The suffused blackish
fuscous fore wing and the whitish hyaline base of hind wing easily
distinguish it from its nearest allies in Group A. (subnubilum and
bicoloranum.)

Grour B.—GENITALIA OF MALE WITH SPINED DiGitus PROJECTING FROM NECK OF
HARPE AWAY FROM SACCULUS (CLOSE TO CUCULLUS)

21. EXARTEMA QUADRIFIDUM Zeller
(Figs. 87, 226)

Exartema quadrifidum ZELLER, Verh. Zool.-bot. Ges. Wien, vol. 25, 1875,
p. 268.

Ezartema inornatanum Frrnatp (not Clemens) in Dyar List N. Amer.
Lepid., no. 5024, 1903.—Barnrs and McDunnoueH, Check List Lepid.
Bor. Amer., no. 6807, 1917.

Cymolomia quadrifida Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 469.

This species has also been wrongly listed along with clavana
Walker as a synonym of znornatanum. It resembles inornatanum.
somewhat in general appearance but has quite different genitalia.
The whitish scaling upon fore wing is also more confined to longi-
tudinal streaks along the veins and not so widely diffused through-
out the median and postmedian areas as it is in Clemens’ species.

146 BULLETIN 182, UNITED STATES NATIONAL MUSEUM

The male genitalia of the different species in this particular group
are very similar and there is little to distinguish them except minor
differences in the spining of the harpes; so, for most of the species
genitalia photographs are omitted, and drawings of the harpes
supplied.

Harpe of male genitalia figured from specimen in National Col-
lection from Ithaca, N. Y. (June 12, 1922). Female genitalia fig-
ured from specimen in Collection Cornell University from Peru,
N. Y. (W. T. M. Forbes).

Bursa of female with shght signum.

Specimens (males) in National Collection and collection Barnes
from Ithaca, N. Y., received through Dr. W. T. M. Forbes, who
writes that the species is not uncommon in that locality. There is
also a male from Bretton Woods, N. H., in the Barnes Collection,
a rubbed and doubtful specimen from Illinois in the American
Museum, a male from British Columbia in the collection of E. H.
Blackmore, and a short series (males and females) in the Canadian
National Collection from Manitoba, Ontario, and Quebec.

Alar expanse.—19-20 mm.

Type.—tin British Museum (?).

Type locality.—Massachusetts.

Food plant.—Cornus.

22. EXARTEMA TILIANUM Heinrich
(Figs. 97, 214, 398)
Exartema tilianum HeEtInricn, Proc. Ent. Soc. Washington, vol. 25, 1923,
p. 113.
Cymolomia tiliana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 468.

Close to permundanum Clemens and nigranwm Heinrich. From
the former it is distinguished by the length of the teeth of the median
bar of fore wing. These are long in ¢élianwm and short in permun-
danum. From both nigranum and permundanum it is distinguished
by the somewhat broadened tip of the uncus of its male genitalia.
The extremity of this organ is narrower in the other two species.

Male genitalia figured from type; female from paratype in Na-
tional Collection from Chicago, Ill. (Kwiat, “ 7-12-02”).

Bursa of female without signum.

Distribution.—New Jersey, Ohio, Illinois, Missouri.

Alar expanse.—18-20 mm.

Type—tin American Museum.

Type locality—Greenwood Lake, N. J.

Food plant—Tilia americana.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 147
23. EXARTEMA SCIOTANUM Heinrich
(Figs. 93, 207)

Ezartema sciotanum HEINRIcH, Proc. Ent. Soc. Washington, vol. 25, 1923,
joe aly
Cymolomia sciotana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 466.
Represented so far in the three collections only by the type material
from the type locality.
Female genitalia and harpe of male figured from paratypes in
National Collection and American Museum.
Bursa of female with signum.
Alar expanse.—19-21 mm.
Type.——Iin American Museum.
Type locality —Cincinnati, Ohio.

24. EXARTEMA TREPIDULUM, new species
(Fig. 81)

Palpus whitish ocherous; second joint with considerable fuscous
shading on outer side and at apex, upper edge sometimes with a
faint rosy tint; third joint fuscous. Face and head ocherous fuscous
shaded with smoky fuscous and with a slight admixture of purplish
s¢éales. Thorax tawny-rose or rosy fuscous; posterior tuft much
darker, blackish brown. Fore wing with pale areas tawny rose or
rosy fuscous, and with dark pattern markings a velvety black-
brown; basal patch reduced to a narrow bar extending obliquely
from dorsal margin just beyond base upward to fold, the outer
edge continuing as a narrow dark line beyond fold to middle;
extreme base of wing and basal half of costa pale, concolorous
with antemedian area; antemedian pale area with a fine median
dark line; median bar dark and well defined, narrow on costa,
broad on dorsum; upper tooth very short, lower long; just above
lower tooth a fine longitudinal ocherous line livides the median
band; lower tooth narrowly connected to dorsal triangular portion
of median band; subtornal spot practically obsolete, indicated only
by a few faint lines of dark scales; subapical bar distinct, broadest
at extremity, only obscurely joined to first costal dash; costal dashes
brownish, a trifle paler and fainter than other dark markings. Hind
wings tawny fuscous; cilia a trifle paler with a dark basal band;
in male a decided notch at vein 10 and a slight one at vein 5.

Harp of male genitalia of type figured.

Alar expanse.—16.5-19 mm.

Type.—tIn American Museum.

148 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Paraty pes—Cat. No. 28038, U.S.N.M. Also in American Mu-
seum and collection Barnes.

Type locality—Hampton, N. H.

Described from male type, five male and one female paratypes
(female without abdomen) from the type locality (S. A. Shaw,
July 16 to Aug. 4). These constituted part of a miscellaneous
series from various eastern localities that Kearfott had under the
name zellerianum Fernald (=nitidanum Zeller not Clemens). It is
quite possible that this may be the true zellerianum. The pattern
matches very well with Zeller’s figure; but the color is not right.
Zeller’s description calls for a species with a golden brown median
bar on fore wing. In trepidulum this and other dark pattern mark-
ings are an almost uniform blackish brown. In as much as another
and quite different species has been generally identified as zelleria-
num (see p. 140) and as the actual type is not at hand for com-
parison I feel that it is better to risk a possible synonym than an
identification which, if wrong, would only add to the confusion of
our literature. A synonym more or less does no harm; but a mis-
identification is always a source of trouble.

25. EXARTEMA NIGRANUM Heinrich
(Fig. 86)

Cymolomia ornatana KEARFOTT (ms.) Ins. New Jersey, 1910, p. 539.
Ezartema nigranum HEINRICH, Proce. Ent. Soc. Washington, vol. 25, 1923,
a lS.
pire aa nigrana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 468.

Kearfott gave the different forms of this variable species a number
of manuscript names, but published no descriptions and allowed only
the above name (ornatana) to get into print. Some of the forms are
strikingly different from each other in pattern; but there are many
intergrading specimens and the genitalia are alike in all.

Harpe of male figured from paratype in the National Collection
from the type locality. Female genitalia similar to those of foot-
zanum (compare fig. 205).

Bursa of female without signum.

Distribution.—Ohio, Illinois, Pennsylvania, New York, New Jer-
sey, Ontario.

Alar expanse.—17-22 mm.

Type.—\In American Museum.

Type locality—Cincinnati, Ohio.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 149
26. EXARTEMA HIPPOCASTANUM Kearfott
(Figs. 88, 224)

Exartema hippocastanum Kerarrott, Bull. Amer. Mus. Nat. Hist., vol. 23,
1907, p. 155.—Barnes and McDunnovueu, Check List Lepid. Bor.
Amer., no. 6812, 1917.

Cymolomia hippocastana Forbes, Memoir 68, Cornell Univ. Agri. Exp. Sta.,
1924, p. 466.

A large species with rich dark brown pattern markings, pale
areas of fore wing ferruginous-ocherous or rosy-ocherous, and basal
patch triparted (broken in three, longitudinally).

Aside from the type series from North Carolina in the American
‘Museum I have seen only one other specimen, a male in the National
Collection from Gainesville, Fla.

Female genitalia and harpe of male figured from paratypes in
American Museum.

Bursa of female with signum.

Alar expanse.—21-22 mm.

Type——tIn American Museum.

Type locality—Black Mountains, N. C.

Food plant—Aesculus (Buckeye).

27. EXARTEMA MERRICKANUM Kearfott
(Figs. 91, 222)

Evartema merrickanum KeEarForT, Bull. Amer. Mus. Nat. Hist., vol. 23,
1907, p. 156—Barnes and McDunnovueH, Check List Lepid. Bor.
Amer., no. 6810, 1917.

Cymolomia merrickana ForsBes, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 466.

Similar to corylanum Fernald; but distinguished by the dark shad-
ing on subapical bar of fore wing. The pretornal spot is normally
strongly shaded in merrickanum, never so in corylanum. Slight
differences in the genitalia are shown in the figures.

Female genitalia and harpe of male figured from paratypes in
the National Collection from the type locality (“ VII-13-03” and
“ VI-29-04,” H. D. Merrick).

Bursa of female without signum.

Distribution —District of Columbia, Pennsylvania, New Jersey,
New York, Wisconsin, Missouri, Iowa, Ontario.

Alar expanse.—18-21 mm.

Type—titn American Museum.

Type locality New Brighton, Pa.

Food plant.—Hamamelis.

150 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

28. EXARTEMA CORYLANUM (Fernald)
(Figs. 84, 228)

Eccopsis corylana FERNALD, Trans. Amer. Ent. Soc., vol. 10, 1882, p. 71..

Ezartema corylanum FrrRNap, in Dyar List N. Amer. Lepid., no. 5022,
1903.—BarNEs and McDunnovueH, Check List Lepid. Bor. Amer., no..
6805, 1917.

Cymolomia corylanad KEARFOTT, Ins. New Jersey, 1910, p. 538—ForsBes,
Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 465.

A species with pattern markings of forewing obscure, color a
much suffused olivaceous fuscous, a strong dark dot at apex, and
conspicuous ferruginous-ocherous cilia.

Harpe of male genitalia figured from specimen in American
Museum from Greenwood Lake, New Jersey (Kearfott, June 10,
1900) ; female from specimen in National Collection from Aweme,
Manitoba (Criddle, “12-VII-05”).

Bursa of female without signum.

Distribution.—New Hampshire, New Jersey, Pennsylvania, Dis-
trict of Columbia, Manitoba.

Alar expanse.—14-19 mm.

Type.—tin National Collection.

Type locality—White Mountains, N. H.

Food plant.—Corylus.

29. EXARTEMA OCHROSUFFUSANUM Heinrich
(Fig. 96)

Ezartema ochrosuffusanum HrtnricH, Proc. Ent. Soc. Washington, vol. 25,
1923. p. 117.
Cymolomia ochrisuffusana Forses, Cornell Univ. Agr. Exp. Sta., 1924, p. 469.

An obscurely marked species similar to corylanum but with fore
wing normally much more suffused with yellow and without contrast-
ingly colored cilia.

Harpe of male genitalia figured from type. Female genitalia like
those of hippocastanum Kearfott (see fig. 224.)

Bursa of female with signum.

Distribution.—Ohio, Illinois, Kansas. In the Barnes Collection
there is a specimen reared by Lindsey from larva taken feeding on
Buckeye at Decatur, Ill. (moth issued May 23, 1920). Specimens
reared from the same food plant have also been received from Theo-
dore H. Frison of the University of Illinois.

Alar expanse.—19-20.5 mm.

Type.—In American Museum.

Type locality.—Cincinnati, Ohio.

Food plant.—Aesculus (buckeye).

“NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 151

30. EXARTEMA BRUNNEOPURPURATUM Heinrich

(Fig. 208.)

Exartema brunneopurpuratum HernricH, Proc. Ent. Soc. Washington, vol.
25, 1923, p. 118.

Cymolomia brunneopurpurata Forses, Memoir 68, Cornell Univ. Agr. Exp.
Sta., 1924, p. 465. r

A small brown and metallic-purple species quite different in appear-
ance from anything else in the genus. So far represented only by
the type and paratype in the National Collection.

Female genitalia figured from type.

Bursa without signum.

Alar expanse.—14-14.5 mm.

Type.—tin National Collection.

Type locality —Falls Church, Va.

Food plant.—Alnus.

31. EXARTEMA FERRUGINEANUM Riley
(Figs. 94, 223)

Exartema ferrugineanum Ritey, Trans. St. Louis Acad. Sci., vol. 4, 1881,
p. 317.—FERNALD, in Dyer List N. Amer. Lepid., no 5011, 1903.—
Barnes and McDuNnNoucGH, Check List Lepid. Bor. Amer., no. 6794,
1917.

Cymolomia ferruginana ForsBes, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 468.

A brilliant ocher yellow species not easily confusable with any-
thing else in this group.

Harpe of male figured from type; female genitalia from specimen
in American Museum from Caldwell, N. J. (Kearfott).

Bursa of female without signum.

Distribution.—Missouri, New Jersey.

Alar expanse.—17-18 mm.

Type.—tn National Collection.

Type locality —St.Louis, Mo.

Food plant.—Plum.

32. EXARTEMA FAGIGEMMEANUM Chambers
(Fig. 92)

Exartema fagigemmaeana CHAMBERS, Can. Ent., vol. 10. 1878, p. 74.

Exartema fagigenmeanum Frrnatp, in Dyer List N. Amer. Lepid., no.
5009, 1903.—Barnes and McDunnoveu, Check List Lepid. Bor. Amer.,
no. 6792, 1917.

Cymolomia fagigemmeana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1925, p. 468.

A variable species as far as color is concerned. Typical specimens
have the pattern markings of fore wing on outer half pale rust red

152 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

and also a faint reddish suffusion on the postmedian pale area;
the dark markings on basal half being tawny white. In other speci-
mens, agreeing in all details of pattern and genitalia with the typical
form, these dark markings are tawny olive throughout and there is
no trace of any reddish shading.

Harpe of male genitalia figured from specimen in National Col-
lection from Cincinnati, Ohio (Braun “ VII-2-04”) ; female genita-
lia similar to those of troglodanum (see Fig. 219).

Bursa of female without signum.

Distribution.—Ohio, Pennsylvania.

Alar expanse.—15-19 mm.

Type—tn Museum Comparative Zoology.

Type locality—Kentucky.

Food plant.—Fagus sylvatica, Ostrya.

33. EXARTEMA SERICORANUM Walsingham
(Fig. 95)

Exartema sericoranum WaALSINGHAM, Illus. Lepid. Heter. Brit. Mus.,
vol. 4, 1879, p.-36.—FERNALD, in Dyar List N. Amer. Lepid., no. 5013,
1903.—BarRNES and McDunnoucGH, Check List Lepid. Bor. Amer., no.
6796, 1917.

Cymolomia sericorana Forses, Memoir 68, Univ. Agr. Exp. Sta., 1924,
p. 466.

Cymolomia myricana KEARFOTT (ms), Ins. New Jersey, 1910, p. 539.

Another variable species in color, but with rather uniform pattern
and structure. Most apt to be confused with melanomesum Hein-
rich from which it differs structurally in having a narrow uncus
and a different grouping of the spines (X’) on sacculus near the arch
of the neck of the harpe. It also has the subapical bar of fore wing
a uniform color throughout and not shading darker toward costa
as is the case in melanomesum. (Kearfott’s manuscript species
“myricana” I am unable to separate from what has generally been
identified as sericoranum.

Harpe of male genitalia figured from specimen in National Ccl-
lection from Lacy, New Jersey (Kearfott, “ VII-14-07”). Female
genitalia similar to those of troglodanum.

Bursa of female without signum.

Distribution.—Maryland, New Jersey, Connecticut, Massachusetts
New Hampshire, Quebec.

Alar expanse.—15-18 mm.

Type—tn British Museum.

Type locality—North America.

Food plant.—Mpyrica.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 158

34. EXARTEMA MELANOMESUM Heinrich
(Fig. 83)
EHzartema melanomesum HEINRICH, Proc. Ent. Soe. Washington, vol. 25,
1923, p. 119.
Cyolomia nortana Krarrott (ms), Ins. New Jersey, 1910, p. 539.
Cymolomia melanomesa ForBes, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 466.

Kearfott never published a description of his nortana and the
name therefore has no standing. Moreover, all the specimens he
had set aside under that name except two from New Jersey are from
Manitoba and are a different species which I am describing else-
where in this paper (p. 154) as brevirostratum. The eee to
which he refers in the New Jersey list I found in the Barnes’ collec-
tion after the description of melanomesum. Otherwise I should have
used his manuscript name.

Harpe of male genitalia of type figured.

Distribution—Maine and New Jersey. There is also a paratype
from Ithaca, New York in the collection at Cornell University.

Alar expanse.—15-17 mm.

Type.—tIn collection Barnes.

Type locality —Sebec Lake, Me.

35. EXARTEMA VALDANUM McDunnough
(Fig. 89)

Ezartema valdanum McDunnouen, Can. Ent., vol. 54, 1922, p. 38.
Cymolomia micantana ForBes, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 464.

This species is easily recognized by the narrow, clear white ante-
median and postmedian areas bordering the broad median fuscous
band on fore wing.

Harpe of male genitalia figured from specimen in National Col-
lection from District of Columbia (male, Schoenborn). Female
genitalia as in permundanum.

Distribution—District of Columbia, New York, Massachusetts,
Maine, New Hampshire, Quebec, Ontario, Manitoba. Except the
types, these specimens (several of them reared) represent a large
series which Kearfott had set aside as a new species under the manu-
script name micantanum but which he never described.

Alar expanse.—13-18 mm.

Type.—tIn Canadian National Collection.

Type locality —Fort Coulonge, Quebec.

Food plant.—Cornus.

154 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

_ 36. EXARTEMA VERSICOLORANUM Clemens

(Fig. 206)
Hzartema versicolorana CLEMENS, Proc. Acad: Nat. Sci. Philadelphia, 1860,
p. 357.
Sericoris versicolorana CLEMENS, Proc. Ent. Soc. Philadelphia, vol. 5,
1865, p. 136.

Heartema versicoloranum FERNALD, in Dyar List, N. Amer. Lepid., 5018,
1903.—BaRNES and McDunnovuGH, Check List Lepid. Bor. Amer., no.
6803, 1917.—McDunnoueH, Can. Ent., vol. 54, 1922, p. 39.

The specimens that have always appeared under this name in the
collections are appendiceum Zellar. Doctor McDunnough has re-
moved the latter from the synonymy and applied Clemens’ name to
a species with the pattern and color of valdanum but with the costa
of fore wing white at base. This appears to be a valid designation,
and there is no good reason for not accepting it since Dr. McDun-
nough’s specimen (from Trenton, Ontario) agrees very well with
Clemens’ description, and the type is lost.

Female genitalia figured from specimen in National Collection
from Buffalo, New York (W. Wild, “ VII-+-09”). This and the
Trenton example in the Canadian Collection are the only specimens
I have seen.

Bursa of female without signum.

Alar expanse.—16-17 mm.

Ty pe.—tLost.

Type locality—Pennsylvania (%)

37. EXARTEMA BREVIROSTRATUM, new species

In pattern very like valdanum McDunnough, with a similar white
antemedian band on fore wing and a whitish postmedian area. The
latter, however, is always more or less shaded with rosy or ferrugi-
nous ocherous. The basal patch, median band, subtornal spot, and
subapical bars are well defined, fuscous brown or ocherous brown
and usually containing some blackish dusting; teeth of median band
well separated, short and sharply pointed; subtornal spot more or
less fusing with dorsal portion of median band. Hind wing smoky
fuscous; cilia paler with a dark basal band; in male slight notches
at veins 1b, 1c, and 5.

Male genitalia similar to those of valdanum.

Alar expanse.—15-15.5 mm.

Type—In American Museum,

Paratypes—Cat. No. 28039, U.S.M.M. Also in American Museum,
Canadian National, and Barnes Collections.

Described from male type and 6 male paratypes from the type lo-
cality (dated July 20 to Aug. 5); and one male paratype labeled.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 155

“Canada.” One of the paratypes Kearfott had labeled “ olivaceanum
Fernald.” The rest formed part of a series which he had set aside
as a new species under the manuscript name “nortanum.” I have
not adopted the latter as it already appears as a manuscript name
in the New Jersey list in connection with two specimens representing
another species (melanomesum Heinrich).

E. brevirostratum is very close to valdanum McDunnough of
which it may very likely be nothing more than a western race.
Until the life histories in this group are more thoroughly worked
out, however, it had better be kept as a separate species.

38. EXARTEMA PERMUNDANUM Clemens
(Figs. 90, 225)

Ezartema permundana CLEMENS, Proc. Acad. Nat. Sci. Philadelphia, 1860,

p. 356.

Sciaphila meanderana Waker, Cat. Lepid. Heter. Brit. Mus., vol. 28,
1863, p. 341.

Sericoris permundana CLEMENS, Proc. Ent. Soc. Philadelphia, vol. 5, 1865,
p. 1384.

Ezartema permundanum ZELLER, Verh. Zool.-bot. Ges. Wien, vol. 25, 1875,
p. 273.—FERNALD, in Dyar List N. Amer. Lepid., no. 5015, 1903.—
BarRNES AND McDunnovueH Check List Lepid. Bor. Amer., no. 6798,
1917.

Cymolomia gaylussaciana Kearrott (ms), Ins. New Jersey, 1910, p. 539.

Cymolomia permundana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 466.

This is a very common species and of some economic importance
as an enemy of strawberry, raspberry, and huckleberry plants. It
seems to have a great variety of hosts. Clemens gives Spiraea as
the host of his type; and in the National Museum we have a speci-
men labeled as reared from hickory. It has also been recorded from
hazel. Published records are doubtful, however, as other species
are frequently identified as permundanum. Kearfott regarded the
huckleberry feeder as a distinct species and gave it the name
gaylussaciana. I can find nothing to separate it from typical
permundanum either in pattern or structure. At any rate the name
has no standing as a description was never published.

Female genitalia and harpe of male figured from reared speci-
mens in National Collection; female from District of Columbia
(“on raspberry, June 12—%9”); male from Waupaca, Wisconsin
(“on strawberry, Chittenden no. 6790”).

Bursa of female without signum.

Distribution.—District of Columbia, Maryland, Pennsylvania,
New York, New Jersey, Massachusetts, New Hampshire, Ohio,
Illinois, Wisconsin, Manitoba, Ontario, Quebec.

Alar expanse.—14-19 mm.

54546—26 11

156 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Types—In Academy Natural Science, Philadelphia (permun-
danum) ; British Museum (meanderana).

Type localities—Pennsylvania (permundanum) ; “ North Amer-
ica” (meanderana).

Food plants——Spiraea salicifolia, Corylus (%), Hicoria, black-
berry, raspberry, strawberry.

39. EXARTEMA SUBMISSANUM McDunnough
(Fig. 85)
Ezartema submissanum McDunnNouGH, Can. Ent., vol. 54, 1922, p. 40.

A suffused cinnamon brown species with pattern markings of
fore wing weakly contrasted against ante and postmedian paler
areas,.and with hind wing whitish hyaline toward base and dark
emoky fuscous toward apex and termen.

Represented by the type and paratypes in the Canadian National
Collection and one paratype (male) in collection Barnes, all from
the type locality.

Harpe of male genitalia figured from paratype in collection
Barnes.

Alar expanse.—16 mm.

Type.—tIn Canadian National Collection.

Type locality.—Ottawa, Ontario.

40. EXARTEMA NANANUM McDunnough

Erartema nananum McDunnoueH, Can. Hnt., vol. 54, 1922, p. 39.

Ezxartema quebecense HEINRICH, Proc. Ent. Soc. Washington, vol. 25,
1923, p. 119.

Cymolomia quebecensis Forsrs, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 464.

A tawny ferruginous species with uniformly dark brown hind
wings, and with fore wing markings defined by much restricted
antemedian and postmedian metallic bands. There are some slight
differences between the types of nananum and quebecense, but not
enough to justify us in holding the latter name.

Male genitalia very similar to those of sericoranuwm but with
uncus broader towards base.

Represented only by the types.

Alar expanse.—12-13 mm.

Types—tIn Canadian National Collection (nananum); in Amer- —
ican Museum (quebecense).

Type localities—Ottawa, Canada (nananum); Quebec, Canada
(quebecense).

‘NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 157

41. EXARTEMA MALANUM (Fernald)
(Figs. 82, 218)

Hecopsis malana FERNALD, Trans. Amer. Ent. Soc., vol. 10, 1882, p. 72.

EHzartema malanum FERNALD, in Dyar List N. Amer. Lepid. no. 5025, 1903.—
Barnes and McDunnovenu, Check List Lepid. Bor. Amer., no. 6808,
1917:

Cymolomia malana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 467.

A very uniformly marked species easily recognized by the charac-
ters given in the key. Feeds normally upon terminal buds and young
leaves of apple. There is in the American Museum a specimen
labeled “on plum”; but whether or not this is an authentic rearing
record I can not say.

Female genitalia and harpe of male genitalia figured from reared
specimens in National Collection from Olden, Mo. (J. P. Taylor,
“21—June—08 ”’).

Bursa of female with signum.

Distribution.—Illlinois, Iowa, Missouri, Pennsylvania, Massachu-
setts.

Alar expanse.—14-15 mm.

Type.—tn National Collection.

Type locality.—Ilinois.

Food plant—Apple, plum (?).

Group C.—MALEe witH SPINE CLUSTER (Spc’) ON HARPE OF GENITALIA WELL
DEVELOPED ; BUT NOT UPON A PROJECTING DiGiITUS

42. EXARTEMA APPENDICEUM Zeller
(Figs. 98, 230, 402)

Exartema appendiceum ZELLER, Verh. Zool.bot. Ges. Wien, vol. 25, 1875,
p. 275.—McDunnoveHu, Can. Ent., vol. 54, 1822, p. 39.

Erartema veriscoloranum FERNALD, in Dyar List N. Amer. Lepid., no. 5018,
1903.—K£ARFOTT, Bull. Amer. Mus. Nat. Hist., vol. 23, 1907, p. 157.—
Barnes and McDunnovueuH, Check List Lepid. Bor. Amer., no. 6803,
1917.

Cymolomia versicolorana KrEArrott, Ins. New Jersey, 1910, p. 539.

Cymolomia appendicea ForBes, Memoir 68, Cornell Uniy. Agr. Exp. Sta.,
1924, p. 464.

Doctor McDunnough resurrected this name from the synonymy
for the species we have been calling versicoloranum, restricting
Clemens’ name to a similar form with white costa at base of fore
wing. In this he is justified, as Clemens’ description covers both
forms and there is no type of versicoloranum in existence to other-
wise settle his concept. The usual food plant of appendiceum is oak
but it probably feeds upon many different plants. In the National

158 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Collection we have specimens reared from currant, raspberry, and
Dixonia.

Male and female genitalia figured from specimens in National
Collection from Oak Station, Pennsylvania (F. Marloff, “ VI-25-10”
and “June 17-04”).

Bursa of female with signum.

Distribution—Maine, Massachusetts, New Jersey, North Caro-
lina, Pennsylvania, Ohio. Colorado, Brith Columbia, Manitoba,
Ontario, Quebec.

Alar expanse.—14-16 mm.

Type.—tn British Museum.

Type locality.—Massachusetts.

Food plants.—Quercus, Dixonia punctalobia, currant, raspberry.

43. EXARTEMA CONCINNANUM (Clemens)
(Figs. 18, 99, 229, 403)

Sericoris concinnana CLEMENS, Proe. Ent. Soe. Philadelphia, vol. 5, 1865,
p. 1384.

Exartema concinnanum FERNALD, in Dyar List N. Amer. Lepid., no, 5017,
19038.—Barnes and McDunnoueH, Check List Lepid. Bor. Amer., no.
6800, 1917.

Cymolomia concinnana Forsrs, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 465.

An olivaceous ocherous species with black dusting upon basal
patch and median bar forming conspicuous blackish spots, one on
lower inner angle of fore wing and the other upon costa near middle.
There is also a rather prominent black spot at apex.

Male and female genitalia figured from specimens in National
Collection: Male from Oak Station, Pa. (Marloff, June 3-06) ; fe-
male from Plummer Island, Md. (Busck, June, 1903).

Bursa of female with signum.

Distribution —North Carolina, District of Columbia, Maryland,
Pennsylvania, New York, New Jersey, Ohio, Ontario.

Alar expanse.—11-15 mm.

Type.—tIn Academy Natural Sciences, Philadelphia.

Type locality.—Virginia.

Food plant.—Blackberry.

44, EXARTEMA CONCINNANUM TERMINANUM McDunnough
Hzeartema terminanum McDunnovueH, Can. Ent., vol. 54, 1922, p. 41.

This form differs from typical concinnanum only in color and
probably should not have even a varietal designation. I am keep-
ing the name on suspicion that the variety may represent a food
plant race. For a long time we have had series in the three collec-
tions under a Kearfott manuscript name (“doacana”’).

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 159

Genitalia as in typical concinnanum.

Distribution.—Maryland, Pennsylvania, Ohio, New Jersey. On-
tario.

Alar expanse—11-15 mm.

Type.—tIn Canadian National Collection.

Type locality —Ottawa, Ontario.

45. EXARTEMA FASCIATANUM Clemens
(Figs. 102, 232, 404)

Ezartema fasciatana CLEMENS, Proc. Acad. Nat. Sci. Philadelphia, 1860,

p. 357.

Sciaphila decisana Waker, Cat. Lepid. Heter. Brit. Mus., vol. 28, 1863,
p. 340.

Sericoris fasciatana CLEMENS, Proc. Ent. Soc. Philadelphia, vol. 5, 1865,
p. 134.

Ezartema albofasciatum ZELLER, Verh. Zool.-bot. Ges. Wien, 1875, p. 272.

Ezartema fasciatanum FERNALD, in Dyar List N. Amer. Lepid. no. 5021,
1908.—BarRNnES and McDunnovucH, Check List Lepid, Bor. Amer.,
no. 6804, 1917.

Cymolomia fasciatana Krarrott, Ins. New Jersey, 1910, p. 539.—ForBzs,
Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 463.

This species is quite common in the vicinity of Washineton during
June and July. It is quite distinct from anything else in the genus
and is easily recognized by the characters given in the key and by
its peculiar genitalia.

Male and female genitalia figured from specimens in National Col-
lection from Plummer Island, Md. (R. C. Shannon, June 7-14,
male) and Allegheny County, Pa. (F. Marloff, “ VI-22-10,”
female).

Bursa of female with signum.

Distribution.—Wllinois, Iowa, Missouri, Kansas, Ohio, Pennsyl-
vania, Maryland, Virginia, North Carolina, District of Columbia,
New York, New Jersey, New Hampshire, Connecticut, Massachu-
setts, Quebec, Ontario.

Alar expanse.—13-16 mm.

Types—In Academy Natural Science, Philadelphia (fasciata-
num) ; British Museum (decisana, albofasciatum).

Type localities —Pennsylvania (fasciatanwm) ; “ North America ”
(decisana) ; Ohio (albofasciatum).

Food plant.—Rumex.

46. EXARTEMA TROGLODANUM McDunnough
(Figs. 103, 219, 405)
Exartema troglodanum McDunnovueH, Can. Ent., vol. 54, 1922, p. 37.

Very like olévaceanum in pattern and often confused with that
species. Easily distinguished by its male genitalia.

160 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Male genitalia figured from type (photograph) and specimen in
American Museum from Greenwood Lake, New Jersey (Kearfott,
June 10, 1900, drawing of harpe) ; female from specimen in National
Collection from Essex County Park, New Jersey (Keartott, July
4). Bursa of female without signum.

Distribution.—_New Jersey, Pennsylvania, Quebec.

Alar expanse.—14-18 mm.

Type.—tIn Canadian National Collection.

Type locality—Meach Lake, Quebec.

47. EXARTEMA EXAERESIMUM, new species
(Figs. 101, 406)

In pattern and color like ¢édianwm Heinrich, but with very dif-
ferent genitalia.

Antenna ocherous with a brownish fuscous spot on upper side
of first joint at base; second joint brownish fuscous above. Palpus
whitish ocherous; second joint with one or two fuscous spots op
outer side; apical joint blackish brown. Head ocherous, shading to
smoky fuscous on top. Fore wing with median areas, outer costal
and pretornal spots and subcostal bar ocherous tawny; basal patch
somewhat darker (more brownish) than outer pattern markings, not
reaching costa, and below costa divided longitudinally by a fine whit-
ish ocherous line; antemedian bar dividing toward dorsum, cutting
deeply into basal patch and inclosing a dorsal spot detached from
basal patch; costa at base whitish ocherous; antemedian and _ post-
median pale areas leaden metallic, faintly interlined with fuscous;
median band broken below Jower tooth, tceth long and narrowly
separated, upper tooth touching or nearly touching subapical bar,
the lower equally long and bluntly pointed, dorsal portion of median
band roughly diamond shaped, separated from or barely touching
pretornal spot; cilia tawny fuscous more or less shaded with dull
ocherous and with a dark basal band. Hind wing ochraceous-
tawny; cilia sordid whitish with a dark basal band; in male a slight
notch at vein 5 and decided notches at veins 1c and 10.

Male genitalia figured from type.

Alar expanse.—18-20 mm.

Type and paratypes—Cat. No. 28040, U.S.N.M. Paratypes also
in American Museum and collection Barnes.

Type locality —Dalias, Tex.

Food plant.—Cornus.

Described from male type and two male paratypes from the type
locality; one male paratype from Kerryville, Tex.; and one male
paratype from Victoria, Tex. (labeled, “on Cornus, pupated
IV-8-07, adult V-4+07, J. D. Mitchell”). The Dallas specimens
originally formed part of the types series of Fernalds’ atrodentanum,

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 161

which I have elsewhere in this paper (p. 141) restricted to the north-
ern form with blackish fuscous dusting on dorsal half of median
band.

In unrubbed specimens the basal patch is darker in exaeresimum
than in ¢zlianum and more completely obliterated on costa by pale
scaling; but in slightly rubber specimens these differences disappear.
The two, however, have such distinct genitalia that there is no con-
fusing them. Those of exaeresimum have the general harpe shape
of Group B&B; but the digitus is entirely lacking and there is a stout
hair brush from the base of the sacculus arch different from any-
thing else in the genus.

48. EXARTEMA FERRIFERANUM (Walker)
(Figs. 100, 220, 407)

Sciaphila? ferriferana Waker, Cat. Lepid. Brit. Mus., vol. 28, 1863, p. 348.

Sericoris gratiosana CLEMENS, Proc. Ent. Soc. Philadelphia, vol. 5, 1865,
p. 134.

Grapholitha (Poecilochroma) usticana ZELLER, Verh. Zool.—bot. Ges. Wien,
vol. 25, 1875, p. 293.

Ezartema ferriferanum WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol.
4, 1879, p. 37.— FERNALD, in Dyer List N. Amer. Lepid., no. 5027, 1903.—
Barnes and McDunnoucH, Check List Lepid. Bor. Amer., no. 6811,
1917.

_Cymolomia ferriferana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 469.

A strikingly marked species easily distinguished on paattern and
genitalia from all other Yxartema.

Male and female genitalia figured from reared specimens in Na-
tional Collection from Bluemont, Virginia (H. B. Kirk, 1—-June-
1914).

Distribution.—District of Columbia, Virginia, North Carolina,
Pennsylvania.

Alar expanse.—16-18 mm.

Types.—tin British Museum (ferviferanum) ; In Academy of Natu-
ral Sciences, Philadelphia (gratiosana); location unknown (us¢i-
cana).

Type localities —*‘ North America” (ferriferanum and usticana) ;
Virginia (gratiosana).

Food plant.—H ydrangea.

17. Genus HEDIA Hiibner
(Fig. 62)
Hedia Hutsner, Verz. Schmet., 1826, p. 380. (Penthina Treitschke).
Genotype.—Phalaena Tinea salicella Linnaeus (Hurope).

Characters as in Olethreutes except:
Male genitalia with cucullus usually more than half the length
of harpe; uncus rather well spined inwardly at apex; cornuti absent

162 BULLETIN 132, UNITED STATES NATIONAL MUSEUM
or much reduced. Female with two signa in bursa, developed as
short thorns or (rarely) slightly impressed scobinate patches.

KEY TO THE SPECIES or HEDIA

1. Fore wing sordid white, finely streaked longitudinally with brownish Jines
and with a moderately broad, hook-shaped bar from midcosta:
(6) lineana.

PERO aN Vat) oO LELOL LS OE Re ae a a Pe 2:
2. Fore wing with no white or whitish areas or markings._______ (5) cyanana.
Fore wing with one or more conspicuous white areas or patches________ oe
3. Terminal area of wing whitish from costa to dorsum____-~-__-_-_______ 4,

Terminal area of wing not white; white markings limited to a large semi-
oval white patch on outer half of costa, but not reaching apex.

(4) chionosema.

4. Outer margin of dark area nearly vertical, at least from costa to middle; a

conspicuous isolated blackish spot in disk beyond it__-___ (1) separatana.

Outer margin of dark area slightly angulate and outwardly slanting from

costa to middle; no such black spot in disk or, if somewhat indicated,

fhen touching ane levokouter Gark marcine =o oe ee eae 5.

5. Pale terminal area white; nonmetallic scaling at tornus__. (2) ochroleucana,

Pale terminal area white with a faintly bluish tinge; at tornus a couple of

bluish metallic bars meeting to outline a faint simitriangular dark patch.

(3) variegana.

1. HEDIA SEPARATANA (Kearfott)
(Figs. 263, 416)

Olethreutes separatana KerarrotT, Trans. Amer. Ent. Soc., vol. 33, 1907,
p. 13; Ins. New Jersey, 1910, p. 539.—Forses, Memoir 68, Cornell Univ.
Agr. Exp. Sta., 1924, p. 455.

Penthina dimidiana Frernatp (not Sodoffsky), Trans. Amer. Ent. Soc., vol.
10, 1882, p. 31.

Olethreutes dimidiana FERNALD, in Dyar List. N. Amer. Lepid., no. 5034,
1903.—KEARFoTT, Can. Ent., vol. 37, 1905, p. 207.

Argyroploce separatana Barnes and McDunNouGH, Check List Lepid. Bor.
Amer., no. 6825, 1917.

The genitalia show as Kearfott contended that this species is dis-
tinct from the European dimidiana. Both, however, go in Hedia.

Male and female genitalia figured from specimens in National
Collection from New Brighton (H. D. Merrick, “ 6—2-03,” male para-
type) and Oak Station, Pa. (Fred Marloff, “ VI-4-11,” female).

Distribution.—Massachusetts, New Hampshire, New Jersey, Mary-
land, Pennsylvania, Ohio, Missouri, Arkansas, Ontario, Manitoba.

Alar expanse.—10-16 mm.

Type.—tIn American Museum.

Type locality —Cincinnati, Ohio.

Food plants—Wild black cherry, larkspur, thorn.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAB 163

2. HEDIA OCHROLEUCANA (Hiibner)
(Figs. 264, 417)

Tortriz ochroleucana Htsner, Samm. Europ. Schmet. Tort., 1814, fig. 304.

Antithesia nimbatana CLEMENS, Proce. Acad. Nat. Sci. Philadelphia, 1860,
p. 346.

Penthina contrariana WALKER, Cat. Lepid., Heter. Brit. Mus., vol. 28,
1863, p. 374.

Penthina consanguinana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus.,
vol. 4, 1879, p. 30.

Olethreutes ochroleucana STAUDINGER and REBEL, Cat. Lepid., vol. 2, no.
1874, 1901.—Pirrcr: and MeETca.Lre, Genitalia Brit. Tort., 1922, p. 43,
pl. 15.

Penthina nimbatana CHITTENDEN, Ent. Bull. U. S. Dept. Agr., no. 27, 1901,
p. 83.

Olethreutes nimbatana FERNALD, in Dyar List N. Amer. Lepid., no. 5031,
1903.—Kerarrotr, Can. Ent., vol. 37, 1905, p. 207—Forses, Memoir
68, Cornell Univ. Agr. Exp. Sta., 1924, p. 455.

Olethreutes consanguinana FERNALD, in Dyar List N. Amer. Lepid., no.
5032, 1903.

Argyroploce nimbatana Barnes and McDuNNoucH, Check List Lepid.
Bor. Amer., no. 6821, 1917.

Argyroploce consanguinana BARNES and McDunnovueH, Check List Lepid.
Bor. Amer., no. 6824, 1917.

There is no real difference in structure or pattern between the
European ochroleucana and our supposed American species. The
latter is variable in size and to some extent in color; but there is a
complete series of intergrades between the smallest eastern nim-
batana and the largest California consanguinana, and also between
both these and typical ochroleucana. 'The genitalia show no varia-
tions of any significance. At most our American form can be no
more than a local race. I do not see anything however to separate
it even as such.

Male and female genitalia figured from specimens in National
Collection from Plumas County, Calif. (June).

Distribution.—Missouri, Kentucky, Virginia, Maryland, Penn-
sylvania, New York, New Hampshire, Connecticut, Massachusetts,
Ontario, Nova Scotia, Manitoba, Alberta, Saskatchewan, British
Columbia, Washington, California, Colorado, Utah, Nevada,
Arizona.

Alar expanse.—13-22 mm.

Types.—Location unknown (ochroleucana); in. Academy Nat-
ural Science (némbatana); in British Museum (contrariana and
consanguinana) .

Type localities—LEurope (ochroleucana) ; Massachusetts (nimba-
tana); Nova Scotia (contrariana); California (consanguinana).

Food plants.—Rose and apple.

54346—26——12

164 BULLETIN 132, UNITED STATES NATIONAL MUSEUM
3. HEDIA VARIEGANA (Hiibner)
(Figs. 262, 418)

Tortrix veriegana Hiner, Samm. Europ. Schmet. Tort., 1814, fig. 14.

Olethreutes variegana STAUDINGER and REBEL, Cat. Lepid., vol. 2, no.
1872, 1901.—Pierce and Mercatre, Genitalia Brit. Tort., 1922, p.
43, pl. 15.

Argyroploce variegana KENNEL, Palaeark, Tort., Lfg. 3, Zoologica, vol. 21,
Heft 54, 1918, p. 379. |

A few examples of this species have been in our collections for
several years either unidentified or under consanguinana Walsing-
ham. There is no appreciable difference between American and
European specimens.

Male and female genitalia figured from specimens in National
Collection from Providence, R. I. (reared under Quaintance No.
1636, May 22, 1916, from plum, J. F. Collins).

Distribution —Rhode Island, Nova Scotia, British Columbia.

Alar expense.—17-20 mm.

Type—tLocation unknown.

Type locality —Kurope.

Food plants——Plum (in Europe: Pyrus, Sorbus, Prunus, Betula,
Myrica, Quercus).

4. HEDIA CHIONOSEMA (Zeller)
(Figs. 261, 419)

Penthina chionosemaé ZELLER, Verh. Zool.-bot. Ges. Wien, vol. 25, 1875,
p. 265.—Murtretpt, Bull. 23, Div. Ent. (old series), U. S. Dept. Agr.,
1891, p. 51.

Olethreutes chionosema FERNALD, in Dyar List N. Amer. Lepid., no. 5047,
1903.—WELLHOUSE, Cornell Univ. Memoir, 56, 1922, p. 1077.—ForBEs,
Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 456.

Argyroploce chionosema BARNES and McDunnovueH, Check List Lepid.
Bor. Amer., no. 6847, 1917.

A striking species easily distinguished by the character given in
the key.

Male and female genitalia figured from specimens in National
Collection from Vienna, Va. (Cushman, “4-30-12,” male) and un-
known locality (female, labeled: “on apple”).

Distribution —Missouri, Kansas, North Carolina, Virginia, Penn-
sylvania, New Jersey, New York, New Hampshire, Connecticut,
Vermont, Ontario, Quebec.

Alar expanse.—14-18 mm.

Type.—In collection Rossler.

Type locality.—North America.

Food plants—Apple, Crataegus, Amelanchier.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 165

5. HEDIA CYANANA (Murtfeldt)
(Figs. 259, 415)

Penthina cyanana MurtTFELDT, Amer. Hnt., vol. 3, 1880, p. 14.

Olethreutes cyanana FERNALD, in Dyar List N. Amer. Lepid., no. 5039,
1903.—HEtnRIcH, Ins. Ins. Mens., vol. 7, 1919, p. 66.—Forses, Memoir
68, Cornell Univ. Agr. Exp. Sta., 1924, p. 458.

Argyroploce cyananad BARNES and McDunnoucH, Check List Lepid. Bor.
Amer., no. 6828, 1917.

A dark semilustrous bluish-purple species with pattern mark-
ings faint and more brownish; similar in general appearance to
Olethreutes agilana Clemens and sia approximana Heinrich but
structurally different.

Male genitalia figured from specimen in National Collection from
Missouri (Murtfeldt, “6—-25”) ; female from specimen in American
Museum from unknown locality.

Distribution.—Missouri, Kansas, Iowa, Indiana, New Jersey,
Manitoba.

Alar expanse.—8-15 mm.

Type.—tn collection Cornell University.

Type locality—Missouri.

Food plant.—Rose,

6. HEDIA (7?) LINEANA (Fernald)
(Fig. 260)

Eucosma lineana FERNALD, Journ. New York Ent. Soe., vol. 9, 1901, p. 501;
in Dyar List N. Amer. Lepid., no. 5116, 1903.—Barnes and Mc-
DunnovueH, Check List Lepid. Bor. Amer., no. 6952, 1917.

This species does not fit into any of the genera we have, and is
only temporarily referred here. In the absence of a male I do not
feel justified in proposing a new genus, but have no doubt that one
must eventually be erected. In female genitalia characters (the
two thorn-like signa) lineana would go easily enough into Hedia;
but the wing shape and venation are wrong; the fore wing termen is
concave, vein 2 is strongly bent, running up to termen parallel to
8, and the upper internal vein of cell is from between 9 and 10.
The pattern, on the other hand, suggests Hartema.

Aside from the female type (without abdomen) I have seen only
one other specimen, a female from Porto Bello, Panama (August
Busck, March, 1911), also in the National Collection. Genitalia of
this latter figured.

Alar expanse.—15-16 mm.

Type.—tIn National Collection.

Type locality—Palm Beach, Fla.

Food plant.—Anona laurifolia.

166 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

18. Genus OLETHREUTES Hiibner
(Figs. 13, 40)

Olethreytes HtUsnrer, Tentamen, 1806 (—Rozana Stephens).
Genotype.—Phalaena Tinea arcuella Clerck (Europe).

Argyroploce HtpBner, Verz. Schmet., 1826, p. 379.
Genotype.—Phalaena Tinea arbutella Linnaeus (Europe).

Phiaris HUsBNER, Verz. Schmet., 1826, p. 381.
Genotype.—Tortriz micana HUBNER (Europe).

Celypha HUBNER, Verz. Schmet., 1826, p. 382. (= Sericoris Treitschke).
Genotype—Phalaena rivulana Scopoli (= conchana Hiibner)

(Europe).

Orthotaenia STEPHENS, Cat. Brit. Ins., vol. 2, 1829, p. 181.
Genotype.—Tortriz striana Denis and Schiffermiiller (Europe).

Selenodes GUENEE, Eur. Microlep. Index Method., 1845, p. 26.
Genotype.—Selenodes dalecarliana Guenée (Europe).

Mizodia GUENEE, Eur. Microlep. Index Method., 1845, p. 26.
Genotype.—Pyralis schulziana Fabricius (Europe).

Thorax with posterior tuft.

Forewing smooth; termen convex or straight; 12 veins; all sep-
arate; 7 to termen; 8 and 9 normally approximate at base, rarely
much separated (costimaculana) ; upper internal vein of cell from
between 10-11; 3, 4, and 5 not aproximate at termen; 2 from cell
usually before 2/3, rarely beyond 2/3 (griseoalbana), usually straight,
rarely bent upward beyond middle (costémaculana).

Hind wing with 8 veins; 6 and 7 approximate toward base; 3 and
4 connate; 5 approximate to 4 at base; termen without appreciable
notches at veins 5, 1c, or 10; inner margin simple or bearing, in male,
a slight chitinous ridge or thickening of the membrane.

Hind tibia of male with hair pencile from base.

Male genitalia with harpe elongate and sometimes bearing a short
row of flat hair-like spines on outer surface, usually without such;
cucullus seldom more than half the length of harp, narrow, densely
spined; sacculus normally extended in an arch over and pocketing
neck; spine clusters Spc! and Spc? usually present and strongly de-
veloped, latter (Spc’) variously modified; spines at base of sacculus
short and weak. Uncus present; sometimes abbreviated but normally
developed; weakly chitinized; tip weakly spined (except in costi-
maculana, denotana, mengelana, and a few exotic species). Socii
present, often small; hairy and _ flexible. Gnathos normal.
Aedoeagus short or long; very slightly bent; cornuti present or ab-
sent, if present one, two or a small cluster of short spines.

Female genitalia with signum present or absent; when present
a scobinate patch occasionally produced into a hollow spine. Ductus
bursae moderately long; unchitinized except near genital opening.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 167

As it stands this genus is still considerable of a lump and probably
should be further divided; but as yet I have found no characters
that will satisfactorily group its conflicting elements. It represents,
I believe, the parent stem of the family.

The type species (arcuella) is strikingly different in pattern from
anything else in the genus, and should its far eastern variety prove
to have a real H'zartema lobe on the male hind wing as claimed
by Walsingham (see p. 130), I should be inclined to restrict the
genus to that species and apply the name Argyroploce to the Amer-

ican species.
KEY TO THD SPECIES OF OLETHREUTES

1. Greater part of fore wing whitish; conspicuous dark markings a bluish
black basal patch and a similar colored subtornal spot.
(1) griseoalbana.

ORCS Witte 0 LACT PLS 0 aes ase ee Re Eat Ne 2.

2. Fore wing brownish with a few small black spots on median and basal
areas and a conspicuous square, pinkish-white blotch on costa beyond .
middle; other pale areas suffused with dark scaling__ (28) costimaculana.
SOT OREN Og OBIE TW LS Cte cep ee a SE RI Le rs ee Ay

3. Fore wing with a clear white semicircular spot on dorsum just beyond base;
terminal quarter of wing white, faintly marked with brownish fuscous;

BES GAO Gaga Sea POW at er ee ees Ce Se (29) devotana.
OKC gwinen Ob Or WwAS esac oh eS a i ee Re 4,
4. Fore wing with a single, rather broad, clear white antemedian fascia and
a more or less extended white postmedian area defining a distinct
brownish or dark grayish fuscous median band______________________ 5.
Antemedian and postmedian areas otherwise; when whitish more or less
suffused or mottled with ocherous or fuscous, or broken into narrow

TENOR UMUC SS ee tee ie OP oe ee EE eae he te ke 10.
5. Postmedian white area a narrow outwardly angulate band, narrower than
aniemedian sw hites DANO! Seen ss See) ie Se (16) carolana.

Postmedian white area broad and occupying greater part of outer quarter
of wing; when reduced by terminal dark shading and markings, in-

WEIL Ua an Cua ake ae aA ee ek I ee he es Se 6.
6. Dark markings olivaceous fuscous_______________ (31) buckellana albidula.
Darkomarkimes dark STeVISHMSEUSCOMG 6 22 ee yo eke ee ee ice

7. Dark markings in terminal area of fore wing obsolete, or only a faint
shading indicating remains of subapical bar; median band divided at
middle by a fine longitudinal white streak___-__-______ (19) dilutifuscana.

(18) bipartitana (part).
Dark markings in terminal area more or less pronounced; subapical bar
at least present, and usually a dark shade between it and apex; no white

Aikenk.Giwaune MeGian DANG. 2 556s ee. 2 ee eee 8.
Pepin ttnaeobandegs 22.22 s Soe Sek Hee eS ee ee ee ee 9.
Elmdstiniae noe, bamcde dis: Se ss See ait ae (18) bipartitana (part).

9. Outer margin of median dark band distinctly toothed below costa.
(17) polluxana.
Outer margin of median band not toothed below costa_____~_ (20) glaciana.

168 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

10.

12.

18.

14.

15.

16.

ite

18.

19.

20.

21.

22.

23.

Fore wing with median area ferruginous ocherous; postmedian area darker.
(2) osmundana.

Fore wing otherwise; if rather pale ferruginous brown in middle, than
with postmedian area paler or broken by irregular white lines______ aie
Fore wing showing to the naked eye a number of shining metallic bluish
Spots Or! Tmmegh see Ces eee Cae De eee OE a iy,
Metallic markings obscured or obliterated; not apparent to the naked
CVC Sa sees oie ee Se a a eee EN Be et 20.
Cilia of fore wing conspicuously white toward apex; dark on lower half
Of COT CN a ee re eT I ne ee Se eee Re 13.
Cilia of fore wing rather uniformly colored; if somewhat whitish, not
conspicuously so toward ‘apex. =o vst oe Sw ee ae ees 14.
Hind wing uniformly dark throughout on upper and under side; cilia oniy
WALIEC BOW TG ea Kees a0 eh RE ee ee ee ee (6) siderana chalybeana.
Hind wing whitish toward base; dark smoky fuscous beyond, the dark
areas narrower on under than on upper side of wing; cilia shining white

almost cO-Inner angles (22 Se ee ee eee (5) albiciliana.
Fore wing no paler in postmedian area than at middle__-__-__-______ 15.
Fore wing paler in postmedian area than at middle_____-___-_________ 16.
Head bright ocherous toward front_____-_-_____________~_ (3) auricapitana.
VOR CC ATC LU SCOUMS iLO VsetT Cl ensdile OT fos eee aes eS (4) agilana.
Basal patch, median band, subtornal spot and subapical bar of fore wing

alhGisemetly denned! to the Make: CYC aaa ee ee ee Alt(e

One or more of the above markings obscured by dark dusting on pale ante-
median area or pale suffusion on dark areas; median band often fusing

WAMel aT kis vt GiGSU LOTMA) post ti CS ees ee ee 18.
Pale dusting on dark areas of fore wing ferruginous brown_ (7) sordidana,
Pale dusting on dark areas of fore wing light ocherous_____~_ (8) galaxana.

Underside of fore and hind wings concolorous, dark shining smoky fuscous.
(11) coruscana.
Underside of hind wing much paler than underside of fore wing,

Ar FT FS) 1a ae i fr ra Poe ce Re ear aL Ae. bayer nen es oh eee eer 19;
Fore wing with termen nearly vertical, slanting only slightly; tornus rather
EV POTSUT TO Gly gr TO LNT CLC Cl oes toe ce es ee (9) constellatana.
Fore wing with termen slanting in even curve from apex to around tornus;
LOENS evenly TOUNnMEed 8 2h Ven LNey oe oes ee eee (10) astrologana.
Fore wing with antemedian and postmedian areas whitish ocherous and
Shanoly. Genninena DO Sy Mec ene bo ela Ces ee ee eee es mile

Fore wing with antemedian and postmedian areas more or less whitish,
but with no ocherous tint; dark median band sometimes sharply defined

but often mottled and broken by pale scaling_________________-_____ 24.
Median dark band of fore wing produced at middle into a single outwardly
DLOJECEII COOL TEs eae a aa) (24) septentrionana.

Median dark band not produced at middle; or, if so, into two teeth___ 22.
Subapical bar of fore wing extending in a broad band from tornus to costa.
(12) puncticostana.

(18) puncticostana major.

Subapical bar from above tornus, and term‘nating, as an appreciable band,

LOY =U ECO) gf Seek CLOYSH Us Oe ean ok aR aa ie = ype aes ne Eten od glue oad on Men Sule) ald ae ee ao.
Median band partially divided by a faint longitudinal pale streak (male) ;
or completely inclosing 48 white discal dot (female)____ (14) deprecatoria.

No such longitudinal pale streak in median band; white discal dot, where
distinguishable, but partially inclosed by median band___ (15) cespitana.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 169

24, Dark median band complete and sharply defined, outwardly biangulate, well
separated from, subtornal spote— 22-222 (21) nordeggana.
Dark median band usually mottled and broken by pale patches, poorly de-
fined, and very irregular in outline; when complete and defined, fused
with subtornal spot (in species having a mottled and broken median
band the subtornal spot is frequently well separated from the remains of
CEES W EL Se PLES ATI OPAL OFT age Nata es Ay ys ra A mr ane eS NM Cate ae 20:
25. Fore wing a suffused mottling of ashy gray and blackish fuscous with no
well-defined whitish areas except a white dot at end of cell, and with

none of the usual dark pattern markings distinctly defined.
(23) intermistana.
Fore wing more or less mottled; but always with one or more definite
whitish areas beside the occasional white dot at end of cell, and with one

or more of the usual dark pattern markings distinctly defined____-__~_ 26.

- 26. Subtornal dark spot of fore wing distinctly separated from dark median
band and more or less extended. —.. ~~~ 2_. + ee VAG
Subtornal dark spot fused with median band______--__---____--__-__ 20:

27. Fore wing with a well-defined subapical bar touching at its discal extremity
the’ apex of subtormalspote lL 2S ih sisi es (22) schulziana.

Fore wing without subapical bar, or latter represented only by a small dark

snot Gn texmeniapove tOFnUS=. 2 2 a eee 28.

28. Fore wing wider toward termen than at middle_______-___ (25) inquietana.
Fore wing no wider toward termen than at middle_______ (26) bowmanana.

29. Postmedian whitish area of fore wing most extended toward costa.
(27) mengelana.
Postmedian whitish area most extended toward tornus____ (30) buckellana.
1. OLETHREUTES GRISEOALBANA (Walsingham)
(Figs. 236, 422)
Exartema griseoalbana WaALSINGHAM, Illus. Lepid. Heter. Brit. Mus.,
vol. 4, 1879, p. 38.

Olethreutes griseoalbana FERNALD, in Dyar List N. Amer. Lepid., no.

5036, 1903.—Forpes, Memoir 68, Cornell Uniy. Agr. Exp. Sta., 1924,
p. 457.

Argyroploce griseoalbana BARNES and McDuNnnoueH, Check List Lepid.
Bor. Amer., no. 6815, 1917.

In this species vein 2 of fore wing branches from the cell very
close to outer three-fourths, further out in fact than it does in any
other species in the genus. However, there doesn’t seem to be any
good reason for generic separation.

Male genitalia figured from specimen in National Collection from
Hampton, N. H. (S. A. Shaw, “ VITI-14-1907”); female from
specimen in American Museum from Cincinnati, Ohio (A. F.
Braun).

Bursa of female with weak signum.

Distribution.—Ohio, Pennsylvania, New Hampshire, North Caro-
lina.

Alar expanse.—14-15 mm.

Type.—tIn British Museum.

Type locality —Eastern States of North America.

170 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

2. OLETHREUTES OSMUNDANA (Fernald)
(Figs. 234, 423)

Penthina osmundana FERNALD, Can. Ent., vol. 11, 1879, p. 156.

Olethreutes osmundana FERNALD, in Dyar List N. Amer. Lepid., no. 5043,
1903.—ForsBres, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 457.

Olethreutes ochromediana KEARFoTT, Trans. Amer. Ent. Soc., vol. 33, 1907,
p. 11—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 457.

Argyroploce osmundana BARNES and McDuNNoucH, Check List Lepid.
Bor. Amer., no. 6850, 1917.

Argyroploce ochromediana BARNES and McDuNNovuGH, Check List Lepid.
Bor. Amer., no. 6851, 1917.

There is an authentic male of osmundana from Maine in the
Cornell collection labeled by Fernald. It agrees in pattern and
genitalia with Kearfott’s ochromediana. I can not find any of
the differ:nces that Kearfott uses to separate his supposed species.

Male and female genitalia figured from specimens in National Col-
lection from Hampton, N. H. (male, S. A. Shaw, “ VIII-14-1907 ”)
and Samons, Mass. (female, reared by W. O. Ellis from Osmunda
cinnamomea, 28 May, 1921).

Bursa of female without signum.

There is a male in the National Collection reared from larvae
feeding on Pteridium aquilinum (“7-7-16,” Witesbog, N. J., Quani-
tance no. 12743, H. H. Scammell, collector). Kearfott also records
(Ins. New Jersey, 1910, p. 540) seeds of Ambrosia trifida as a food
plant.

Distribution —District of Columbia, Pennsylvania, New Jersey,
New Hampshire, Massachusetts, Maine.

Alar expanse.—11-13 mm.

Types.—In National Collection (osmundana); in American
Museum (ochromediana).

Type localities——Orono, Me. (osmundana) Hazelton, Pa. (ochro-
mediana).

Food plants—Osmunda regalis, O. cinnamomea, Pteridium aquil-
inum, Ambrosia trifida (%).

3. OLETHREUTES AURICAPITANA (Walsingham)
(Figs. 238, 424)

Sericoris auricapitana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol. 4,
1879, p. 33.

Olethreutes auricapitana FrRNaLp, in Dyar List N. Amer. Lepid. no. 5052,
1903.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 458.

Argyroploce auricapitana Barnes and McDunnoveH, Check List Lepid.
Bor. Amer., no. 6830, 1917.

Male and female genitalia figured from specimens in National
Collection; male from Great Falls, Va. (Busck) ; female from Essex
County Park, N. J. (Kearfott).

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 171

Bursa of female with signum.

Distribution —New Jersey, Virginia, Nova Scotia.
Alar expanse.—10-12 mm.

Type.—tIn British Museum.

Type locality Pennsylvania (?).

Food plant.—Betula lutea.

4. OLETHREUTES AGILANA (Clemens)
(Figs. 235, 439)

Endopiza ? agilana CLemMENs, Proc. Acad. Nat. Sci. Philadelphia, 1860,
. 359.

note agilana FERNALD, in Dyar List N. Amer. Lepid., no. 5053,
1903.—KeEarFoTT, Ins. New Jersey, 1910, p. 540—Forses, Memoir 68,
Cornell Univ. Agr. Exp. Sta., 1924, p. 458.

Olethreutes abiciliana Busck (not Fernald), Proc. Ent. Soc. Washington,
vol. 11, 1909, p. 98.

Argyroploce agilana BARNES and McDuNNouGH, Check List Lepid. Bor.
Amer., no. 6831, 1917.

A specimen in the collection of the Philadelphia Academy has
been labeled by Fernald as the probable type. Since it matches the
description and agrees with the general concept of the species it
may be so regarded.

The metallic spots are duller in agilana than in the other dark
species with similar markings; but in fresh unrubbed specimens are
easily seen without a lens. They only become completely obscured
in greasy specimens.

Male and female genitilia figured from specimens in National
Collection from Ithaca, N. Y. (male, reared, E. H. Hausman, May,
1920) and Oak Station, Pa. (female, F. Marloff, “June 10-07”).

Bursa of female without signum.

Distribution.—Ohio, Pennsylvania, New York, New Jersey, Mary-
land, Virginia, Massachusetts, Ontario.

Alar expanse.—11-14 mm.

Type.—In Academy Natural Science, Philadelphia.

Type locality.—Pennsylvania (?)

Food plant.—Impatiens (larva a stem borer).

5. OLETHREUTES ALBICILIANA (Fernald)
(Figs. 237, 425)

Sericoris albiciliana FrERNAtLD, Trans. Amer. Ent. Soc., vol. 10, 1882, p. 70.

Olethreutes albiciliana FERNALD, in Dyar List N. Amer. Lepid., no. 5054,
1903.—Forpes, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 458.

Argyroploce albiciliana Barnes and McDuNnNovucH, Check List Lepid. Bor
Amer., no. 6832, 1917.

172 BULLETIN 182, UNITED STATES NATIONAL MUSEUM

Male and female genitalia figured from specimens in National Col-
lection from Aweme, Manitoba (Criddle, “13-VII-05,” male) and
Bretton Woods, White Mountains, N. H. (“July 8-15,” female).

Bursa of female with signum.

Distribution—South Carolina, Pennsylvania, New Jersey, New
Hampshire, Maine, Ohio, Manitoba.

Alar expanse.—12-15 mm.

Type.—tIn National Collection.

Type locality —Orono, Me.

Food plant.—S piraea salicifolia (Kearfott notes).

6. OLETHREUTES SIDERANA CHALYBZANA (Walsingham)

(Fig. 427)
Sericoris chalybeana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol. 4,
1879, p. 34.
Olethreutes chalybeana FERNALD, in Dyar List. N. Amer. Lepid., no. 5055,
1903.

Argyroploce chalybeana BARNES AND McDunnoueH, Check List Lepid.
Bor. Amer., no. 6833, 1917.

In structure and: pattern Walsingham’s chalybeana matches very
closely the typical European stderana Treitschke (fig. 426). Speci-
mens of the latter in the National Museum, however, are a trifle
different in color. The pale shade in cilia of fore and hind wings
is whiter in American than European specimens, the dark line at
base of fore wing cilia is also uninterrupted while in the European
specimens it is partially obliterated below apex by encroachment of
the pale shading of the cilia, and the yellow dusting of fore wing
is paler—less orange colored—than in European examples. It is
doubtful if such slight differences will hold consistently through
large series from any given locality. I have only seen three Ameri-
can and a half dozen European specimens, and in these they do.
I am therefore keeping the Walsingham name for the present, upon
suspicion that it may designate a distinguishable local race. Both
forms occur in rather high altitudes. In Europe stderana feeds
upon Spiraea. None of the American specimens have been reared.

Male genitalia figured from cotype in National Collection. Fe-
male genitalia similar to those of albiciliana.

Besides the cotype, we have in the National Collection a male from
Kaslo, British Columbia. There is also a male from Glacier Na-
tional Park, Mont., in the Barnes Collection, and two specimens
(male and female) from Kaslo in the Canadian National Collection.

Alar expanse.—15-16 mm.

Type.—tn British Museum.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 178

Type locality.—Siskiyou Mountains, on the borders of Oregon and
California.
Food plant—Unknown (probably Spiraea).

7. OLETHREUTES SORDIDANA (McDunnough)
(Fig. 428)
Argyroploce sordidana McDunnoueH, Can. Ent., vol. 54, 1922, p. 48.

In genitalia close to constellatana Zeller. In pattern and general
habitus resembling some of the smaller specimens of galaxana and
constellatana, but much darker, above and beneath, and more brown-
ish than either. So far represented only by the type material in the

‘Canadian National and Barnes Collections.

Male genitalia figured from paratype in collection Barnes.

Alar expanse.—16 mm.

Type—tiIn Canadian National Collection.

Type locality —Coliseum Mountain, Nordegg, Alberta.

5d)
8. OLETHREUTES GALAXANA Kearfott
(Figs. 241, 431)

Olethreutes galaxana Krarrotr, Trans. Amer. Ent. Soc., vol. 33, 1907, p. 9.

Olethreutes glitranana Kerarrott, Trans. Amer. Ent. Soc., vol. 33, 1907,
yi

eae cies galaranad BARNES and McDunnovuGH, Check List Lepid. Bor.
Amer., no. 6836, 1917.

Argyroploce glitranana BARNES and McDuNNovuGH, Check List Lepid. Bor.
Amer., no. 6837, 1917.

There are no real differences in genitalia, color, or pattern be-
tween Kearfott’s supposed two species. In size the more easterly
form (glitranana) averages smaller than specimens from British
Columbia; but in Manitoba and Vancouver Island there are inter-
grades, so the forms can not be distinguished even as races.

Male and female genitalia figured from specimens in National
Collection from Victoria, British Columbia (male paratype of ga-
laxana), and Regina, Saskatchewan (female paratype glitranana).

Bursa of female with signum developed as.a scobinate patch with
a small central spine.

Distribution —Colorado, Saskatchewan, Manitoba, Alberta, Brit-
ish Columbia, Vancouver, Alaska.

Alar expanse.—16-23, mm.

Types.—In American Museum.

Type localities—Vernon, British Citas din (galaxana); Re-
gina, Saskatchewan (glitranana).

174 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

9. OLETHREUTES CONSTELLATANA (Zeller)
(Figs. 240, 429)

Sericoris constellatana ZELLER, Verh. Zool.-bot. Ges. Wien, vol. 25, 1875,
p. 279.

Olethreutes constellatana FERNALD, in Dyar List N. Amer, Lepid., no.
5057, 1903.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 454.

Argyroploce constellatana Barnes and McDunnovueH, Check List Lepid.
Bor. Amer., no. 6835, 1917.

Very similar to the larger form of astrologana Zeller and fre-
quently confused with it. The two species have different genitalia
and may also be distinguished by the shape of the fore wing. The
latter is broader and has the termen a little more vertical in con-
stellatana than in astrologana.

Male and female genitalia figured from specimens in National
Collection from Kansas (male) and Arendtsville, Pa. (S. W. Frost,
June 8, 1919, female).

Bursa of female with signum a short hollow spine.

Distribution.—New York, New Hampshire, Massachusetts, Penn-
sylvania, Maryland, Virginia, West Virginia, Ohio, Illinois, Iowa,
Kansas, Wisconsin, Ontario, Quebec.

Alar expanse.—18-22 mm.

Type.—tIn British Museum ?

Type locality —Ohio.

10. OLETHREUTES ASTROLOGANA (Zeller)
(Figs. 239, 433)

Sericoris astrologana ZELLER, Verh. Zool.-bot. Ges. Wien, vol. 25, 1875,
p. 281.

Olethreutes astrologana FERNALD, in Dyar List N. Amer, Lepid., no. 5062,
1903.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 454.

Olethreutes coronana Kearrott, Trans. Amer. Ent. Soc., vol. 33, 1907, p.
10.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 454.

Argyroploce astrologana BarRNES and McDunnovueH, Check List Lepid.
Bor. Amer., no. 6843, 1917.

Argyroploce coronana BaRNES and McDunnoucH, Check List Lepid. Bor.
Amer., no. 6838, 1917.

The larger western specimens of this species closely resemble con-
stellatana Zeller and the smaller eastern specimens are very similar
to coruscana with which astrologana has been frequently confused.
In superficial characters it differs from the former in having termen
of fore wing more slanting, and from the latter in having under side
of hind wing distinctly paler than under side of fore wing. The
genitalia are characteristic. The species is variable as to size and
intensity of dark scaling on the fore wing. The small eastern and
large western forms could be easily separated as local races, did not

“NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 175

intergrading specimens occur throughout the middle states. Kear-
fott’s coronana is only a dark specimen of the western form. In
genitalia it is a typical astrologana.

Male and female genitalia figured from specimens in National
Collection from Long Island, New York (male) and Kaslo, British
Columbia (Dyar, “No. 20792,” female).

Bursa of female with signum a scobinate patch with a small weak
central spine.

Distribution—Maryland, Virginia, New York, Maine, Illinois,
Iowa, Missouri, New Mexico, Texas, Ontario, Alberta, Manitoba,
British Columbia.

Alar expanse.—14-19 mm.

Types—tIn British Museum (astrologana); American Museum
(coronana).

Type localities—Texas (astrologana); Aweme, Manitoba (coro-
nana).

11. OLETHREUTES CORUSCANA (Clemens)
(Figs. 242, 480)

Antithesia coruscana CLEMENS, Proc. Acad. Nat. Sci. Philadelphia, 1860,

p. 346.

Carpocapsa ferrolineana WALKER, Cat. Lepid. Heter. Brit. Mus., vol. 28,
1863, p. 395.

Sericoris argyroelana ZELLER, Verh. Zool.-bot. Ges. Wien, vol. 25, 1875,
p. 277.

Olethreutes coruscana FERNALD, in Dyar List N. Amer. Lepid., no. 5056,
1903.—Dyar, Proc. U. 8. Nat. Mus., vol. 27, 1904, p. 925.—KEaRForT,
Can. Ent., vol. 37, 1905, p. 207—Forpes, Memoir 68, Cornell Uniy.
Agr. Exp. Sta., 1924, p. 453.

Argyroploce coruscana BARNES and McDuNNouGH, Check List Lepid. Ber.
Amer., no. 6834, 1917.

Very like smaller eastern specimens of astrologana but with very
different genitalia and with under side of fore and hind wings both
of the same dark shade. The male genitalia are similar to those of
constellatana,; but differs consistently in one very significant detail;
in constellatana the spining on the cucullus of harpe extends along
the arch over the neck of the harpe and fuses in with spine group,
Spe; while in coruscana it ends abruptly at the beginning of the
arch and the spine group, Spe,’ is isolated.

Male and female genitalia figured from specimens in National
Collection from Washington, District of Columbia (‘‘28-5-85”
and “ 24-5-79”’).

Distribution —North Carolina, Virginia, Maryland, District of
Columbia, Pennsylvania, New York, New Jersey, Massachusetts,
New Hampshire, [llinois, Iowa.

Alar expanse.—i4-17 mm.

176 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Types.—In Academy Natural Sciences, Philadelphia (coruscana) ;
British Museum (ferrolineana and argyroelana).

Type localities—Pennsylvania? (coruscana) ; “ North America’
(ferrolineana) ; New York ( argyroelana).

12. OLETHREUTES PUNCTICOSTANA (Walker)
(Fig. 254)

Sciaphila puncticostana WALKER, Cat. Lepid. Heter. Brit. Mus., vol. 28,

1863, p. 389.
Penthina murina PackagrD, Proc, Boston Soc. Nat. Hist., vol. 11, 1867, p. 60.
Sericoris puncticostana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol.
4, 1879, p. 33.
Olethreutes murina FERNALD, in Dyar List N. Amer. Lepid., no. 5042,
1903.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 457.
Olethreutes puncticostana FERNALD, in Dyar List N. Amer. Lepid., no. 5063,.

1903.
Argyroploce puncticostana BARNES and McDunNoucH, Check List Lepid.

Bor. Amer., no. 6844, 1917.
Argyroploce murina BARNES and McDuNNovueH, Check List Lepid. Bor.

Amer., no. 6857, 1917.

According to Knight’s figure in the Kearfott Collection at the
American Museum this species is the eastern equivalent of major
Walsingham. In the collections specimens of cespitana, urticana,
and deprecatoria have been indiscriminately referred to it. I was
inclined to apply the name to the large northern and western form
of cespitana Hiibner (instrutana Clemens) ; but according to August
Busck, who examined the type, puncticostana lacks the dark shading
on the pale terminal area of fore wing which is present in all speci-
mens of cespitana.

We have in the National Collection two males from New York and
Quebec received through Dr. W. T. M. Forbes. These agree in geni-
talia with major Walsingham. In the Canadian National Collec-
tion there are several small specimens (15-17 mm.) from New York,
and a series of larger (17-20 mm.) specimens from Ontario and Cen-
tral Alberta (Edmonton). ‘There is also a male in the Barnes Col-
lection from Mount Washington, N. H. (“July 24-31, 4,000 ft.”).
Packards’ murina is a straight synonym. The type at Cambridge
agrees in both pattern and genitalia with what we have under the
Walker name.

Female genitalia figured from specimen in the Cornell Univer-
sity collection from North Twin Brooks, N. Y. (W. T. M. Forbes,
“10-7-18”).

Bursa of female with signum a small impressed scobinate patch.

Alar expanse.—15-20 mm.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 177

Types—tIn British Museum (puncticostana); in Museum Com-
parative Zoology (murina).

Type localities—Nova Scotia (puncticostana); Caribou Island,
Labrador (murina).

13. OLETHREUTES PUNCTICOSTANA MAJOR (Walsingham)
(Fig. 444)

Penthina major WALSINGHAM, Trans. Ent. Soe. London, 1895, p. 502.

Olethreutes major FERNALD, in Dyar List N. Amer. Lepid., no. 5058, 1903.—
Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta. 1924, p. 454.

Argyroploce major BaRNES and McDunNouGcH, Check List Lepid. Bor.
Amer., no. 6839, 1917.

A somewhat larger Rocky Mountain and Pacific coast variety of
puncticostana. It probably does not deserve to be distinguished even
as a race; but should be kept as such until intergrading forms are
recovered. In pattern and genitalia metallicana Hiibner is very
close. It is possible that our American forms are varieties of the
European species. I believe, however, that they are distinct.

Male genitalia figured from specimen in National Collection
(“ Colo. 2168”).

Distribution.—British Columbia, Alberta (Banff). There are
also male and female specimens from Atlin, British Columbia, in
the collection of Mr. E. H. Blackmore.

Alar expanse.—23-25 mm.

Type.—tiIn British Museum.

Type locality.—Loveland, Colo.

14. OLETHREUTES DEPRECATORIA, new species
(Figs. 247, 486)

Palpi, face, and head whitish ocherous; some faint fuscous dusting
on palpus toward apex of second joint; third joint pale. Thorax
pale acherous with slight peppering or cross streaking of dark fus-
cous. Fore wing dull whitish ocherous with brownish or grayish
fuscous pattern markings; pale antemedian and postmedian areas
finely lined with fuscous; dark markings dusted with ocherous
scales, giving them a rather dull ashy fuscous shade, especially in
the males (the females have the dark areas less overlaid with pale
scaling and are more brilliant and more sharply contrasted against
the pale areas) ; dark basal patch sharply excavated below middle,
vertically lined with ocherous; median band narrow on costa, out-
wardly rounded at end of cell and inclosing a faint whitish dot, more
or less coalescing with pretornal spot on dorsum (especially in males)

178 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

and usually partially interrupted at middle by a faint pale streak
(not noticeable in most of the females but rather distinct in the
males) ; subapical bar from termen near tornus to vein 8; costa with
four outer costal spots; a faint fuscous shading between apical spot
and subapical bar; cilia whitish with a fuscous patch at apex and
opposite base of subapical bar. Hind wing smoky fuscous; cilia
whitish with a dark basal band. Underside of hind wing whitish,
much paler than under side of fore wing. Hair pencile on hind
tibia of male pale fuscous.

Male genitalia of type figured. Female genitalia figured from
paratype in National Collection from Shasta Retreat, Calif.

Bursa of f male with signum a weak scobinate patch.

Alar expanse.—15-19 mm.

Type and paratypes.—Cat. No. 28041, U.S.N.M. Paratypes also
in Canadian National Collections, American Museum, and collec-
tion Barnes.

Type locality Wellington, British Columbia.

Described from male type, 5 male and 4 female paratypes from
the type locality (June and July); 3 male paratypes from Hot
Springs, Green River, Wash.; 1 male paratype from Oysterville,
Wash. (taken in cranberry bog by H. K. Plank, ‘ 6-21-18,” Quain-
tance No. 15533); 1 female paratype from Grayland, Wash.
(Quaintance No. 15502, H. K. Plank, collector, “ 8-15-18”) ; 1 male
paratype from Clatsop, Wash. (Quaintance No. 15510, H. K. Plank,
“7-9-18”); and 2 male and 2 female paratypes from Shasta Re-
treat, Siskiyou County, Calif.

In addition to the above I have before me four males from New
Hampshire (“Mount Washington, 4,000 ft., July 24-31,” 2 speci-
mens; and Glen House, White Mountains, 1,600 ft., Aug. 1-7,” 2
specimens) which belong here but which I hesitate to include among
the types as they may represent a distinct eastern race. The pat-
tern and genitalia are typical but the white areas of for: wing show
little or none of the strong ocherous shading characteristic of west-
ern specimens.

In the collections this species has appeared most frequently
under the name urticana Hiibner. It has also been identified as
puncticostana, and one of the cotypes had been labeled chalybeana
Walsingham by Kearfott. In genitalia and gen:ral appearance it
most closely resembles the European lacunana Duponchel and
umbrosana Freyer. It differs from both however in the spining
of the harpe of the male genitalia. The differences are slight but
appear to be constant.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 179

15. OLETHREUTES CESPITANA (Htbner)
(Figs. 245, 434)

Tortrix cespitana Hiiener, Samm. Bur. Schmet. Tort., 1814, figs. 244-245.

Rhyacionia flavofasciana WeEstwoop and HUMPHREYS, vol. 2, 1844, p. 173,
pl. 99, fig. 16.

Sericoris instrutana CLEMENS, Proc. Ent. Soc. Philadelphia, vol. 5, 1865,
p. 135.

Sericoris poana Zeer, Verh. Zool.-bot. Ges. Wien, 1875, p. 282.

Eucosma cespitana Meyrick, Hand Book Brit. Lepid., 1895, p. 468.

Olethreutes cespitana STAUDINGER and REBEL, Cat. Lepid., vol. 2, no. 1927,
1901.

Olethreutes instrutana FERNALD, in Dyar List N. Amer. Lepid., no. 5064,
1903.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 453.

Olethreutes flavofasciana Krarrott, Ent. News, vol. 20, 1909, p. 53; Ins.
New Jersey, 1910, p. 539.

Argyroploce instrutana Barnes and McDunnoucH, Check List Lepid.
Bor. Amer., no. 6862, 1917.

Celype cespitana Pierce and Metcatr, Genitalia Brit. Tort., 1922, p. 52,
pls 1%

There are no differences in pattern or genitalia b- tween the Euro-
pean cespitana and the American instrutana and I have no doubt
that they are but one species. Our American specimens vary con-
siderably in size. In Colorado and western California there is a
large variety measuring from 17 to 20 mm. alar expanse. Speci-
mens of this have been identified—and I now think incorrectly—as
puncticostana Walker. I am not differentiating it as a race because
there are many Colorado specimens intergrading in size between it
and the smaller eastern form. Colorado specimens range all the
way from 12 to 20 mm. expanse; typical eastern specimens from 11
to 16 mm. The genitalia of all are alike, not varying in the
slightest degree.

Male and female genitalia figured from specimens in National
Collection from Clear Creek, Colo. (Oslar, male), and Washington,
D. C. (Busck, female).

Bursa of female with signum a weak scobinate patch.

Distribution —North Carolina, District of Columbia, Pennsyl-
vania, New Jersey, New York, New Hampshire, Massachusetts,
Ohio, Kentucky, Tennessee, Iowa, Nebraska, Missouri, Wisconsin,
South Dakota, Colorado, Utah, California, Washington, British
Columbia, Alberta, Manitoba, Ontario.

Alar expanse.—11-20 mm.

Types.—Location unknown (cespitana and flavofasciana) ; Acad-
emy Natural Science, Philadelphia (énstrutana); British Museum
(poana).

180 BULLETIN 1832, UNITED STATES NATIONAL MUSEUM

Type localities —Europe (cespitana); England (flavofasciana) ;
Virginia (instrutana) ; Ohio (poana).
Food plants—Clover, grass, horse chestnut (in Europe, Spar-
tuum).
16. OLETHREUTES CAROLANA (McDunnough)

(Figs. 248, 438)

Argyroploce carolana McDunnNoucH, Can. Ent., vol. 54, 1922, p. 46.
Olethreutes carolana Forsres, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 451.
A distinct species. Represented only by the type material in the
Canadian National Collection.
Genitalia figured from type (male) and paratype (female) from
Trenton, Ontario (“11-6-11, Evans”).
Alar expanse.—14 mm.
Type.—tn Canadian National Collection.
Type locality —Ottawa, Ontario.

17. OLETHREUTES POLLUXANA (McDunnough)

(Figs. 244, 445)

Argyroploce polluxana McDuNNovGH, Can. Ent., vol. 54, 1922, p. 45.
Olethreutes polluxana Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta.,
1924, p. 452. ;

Male genitalia figured from paratype in collection Barnes from
the type locality; female from specimen in National Collection from
Mount Washington, N. H.

Bursa of female with signum a scobinate patch. The ductus
Bursae is chitinized for a considerable part of its length, rather
an unusual character for this genus.

Distribution —Alberta, Ontario, New Hampshire. There is also
a male from Peru, N. Y., in the Cornell University Collection.

Alar expanse.—18-19 mm.

Type.—In Canadian National Collection.

Type locality —Nordegg, Alberta.

18. OLETHREUTES BIPARTITANA (Clemens)
(Figs. 249, 435)

Antithesia bipartitana CLEMENS, Proce. Acad. Nat. Sci., Philadelphia, 1860,
p. 346.—Packarpb, Guide Study Ins., 1869, p. 333.

Penthina similisana WALKER, Cat. Lepid. Heter, Brit. Mus., vol. 28, 1863,
p. 373.—WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol. 4, 1879,
p. 34.

Sericoris caesialbana ZELLER, Verh. Zool.-bot. Ges. Wien, vol. 25, 1875,
p. 285.—FERNALD, Trans. Amer. Ent. Soec., vol. 10, 1882, p. 34.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 181

Sericoris bipartitana Davis, Bull. Michigan Agr. Sta., no. 102, 1893, p. 30.

Olethreutes bipartitana FrrNap, in Dyar List N. Amer. Lepid., no. 5071,
1903.—Forsrs, Memoir 68, Cornell Univ. Agr. Exp. Sta. 1924, p. 451.

Olethreutes similisana FERNALD, in Dyar List N. Amer. Lepid., no. 5072,
1903.

Olethreutes dilutifuscana Krarrott (not Walsingham), Can. Ent., vol.
37, 1905, p. 208.

Argyroploce bipartitana Barnes and McDunnouGH, Check List Lepid.
Bor. Amer., no. 6869, 1917.

Argyroploce similisana BARNES and McDunNovuGH, Check List Lepid, Bor.
Amer., no. 6870, 1917.

Walsingham referred simélisana Walker to the synonymy in 1879;
but in this was not followed by our cataloguers, why I do not know
as there does not seem to be any reason for doubting the correctness
of his reference. From the description and figure dilutifuscana
Walsingham also appears to be a synonym or variety of bipartitana.
Zeller’s caesialbana is certainly equal, as Fernald had it, to bzparti-
tana. ‘There are cotypes of the former in the National Collection
which agree in every detail with Clemens’ type in Philadelphia.
In general appearance bipartitana Clemens and glaciana Moschler
are very similar and easily confused. Both species are variable in
the extent and intensity of the dark markings in the postmedian area
of the fore wing. The characters given in the key will help to dis-
tinguish most specimens; but the only sure way to separate the two
is by their genitalia which are quite different.

Male and female genitalia figured from specimens in National
Collection from East River, Conn. (C. R. Ely, July 13, 1907, male),
and Sebec Lake, Me. (June, female).

Bursa of female with signum a scobinate patch.

Distribution.—Maine, New Hampshire, Vermont, Massachusetts,
Connecticut, New York, New Jersey, Pennsylvania, Montana, Nova
Scotia, Quebec, Ontario, Manitoba, Saskatchewan, Alberta, Alaska.

Alar expanse.—15.5-20 mm.

Types.—In Academy Natural Science, Philadelphia (bipartitana) ;
British Museum (similisana) ; Museum Comparative Zoology (cae-
sialbana) .

Type localities—Massachusetts (bipartitana, caesialbana):; St.
Martins Falls, Albany River, Hudsons Bay (sindlisana).

Food plant.—Celery.

19. OLETHREUTES DILUTIFUSCANA (Walsingham)

Sericoris dilutifuscana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol. 4,
1879, p. 33.

Olethreutes dilutifuscana FERNALD, in Dyar List N. Amer. Lepid., no. 5070,
1903; not Kearfott, Can. Ent., vol. 37, 1905, p. 208.

Argyroploce dilutifuscana BARNES and McDunnovueH, Check List Lepid.

Bor. Amer., no. 6868, 1917.

182 BULLETIN 182, UNITED STATES NATIONAL MUSEUM

Specimens from Saskatchewan determined by Kearfott as this
species are bipartitana. I am inclined to agree and to refer dilutifus-
cana to the synonymy, except for the fact that Walsingham has taken
particular pains to distinguish his species from bipartitana. The
variability of the latter, however, invalidate the characters he uses
for separation. On the other hand the type locality of dilutifus-
cana seems to be somewhat outside the known range of bipartitana.

There is a specimen from Walsingham in the Fernald Collection
labeled “ Type” which equals urticana Hiibner. I have some doubts,
however, that this supposed paratype is correctly identified, and
until the genitalia of the actual type can be examined it were better
to keep the Walsingham name out of synonymy.

Alar expanse.—16 mm. 3

Type.—in British Museum.

Type locality —Southern Oregon.

20. CLETHREUTES GLACIANA (Mobschler)
(Figs. 248, 432)

Sericoris glaciana-M6scHLER, Wien, Ent. Monat., vol. 14, 1860, p. 380.

Penthina dealbana WALKER, Cat. Lepid. Heter. Brit. Mus., vol. 28, 1863,
p. 374.

Sericoris fuscalbana ZELLER, Verh. Zool.-bot. Ges. Wien, vol. 25, 1875, p. 284.

Sericoris dealbana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol. 4,
1879, p. 84; Trans. Ent. Soc. London, 1884, p. 1386.

Olethreutes fuscalbana FERNALD, in Dyar List N. Amer. Lepid., no. 5067,
1903.—KeEarFoTtT, Can. Ent., vol. 37, 1905, p. 208; Ins. New Jersey,
1910, p. 540.—¥Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924,
p. 451

Olethreutes glaciana FERNALD, in Dyar List N. Amer. Lepid., no. 5068,
1903.—KeEaRFoTT, Can. Ent., vol. 37, 1905, p. 208.

Olethreutes dealbana FERNALD, in Dyar List N. Amer. Lepid., no. 5069,
1908.

Argyroploce fuscalbana Barnes and McDuNNouGH, Check List Lepid. Bor.
Amer., no. 6865, 1917.

Argyroploce glaciana BaRNES and McDuNNovuGH, Check List Lepid. Bor.
Amer., no. 6866, 1917.

Argyroploce dealbana BaRNES and McDunnovueH, Check List Lepid. ie
Amer., no. 6867, 1917.

Argyroploce castorana McDuNNouGH, Can. Ent., vol. 54, 1922. p. 54.

I do not think there can be any reasonable doubt of the above
synonymy. We have a paratype of fuscalbana in the National
Collection, and it agrees in genitalia with cvstorana and what Fernald
and others have identified as glaciana. Walker’s dealbana I am
placing here on the strength of Walsingham’s description and figure
and Knight’s very careful figure in the American Museum. Kear-
fott usually determined specimens of urticana as dealbana. This 1

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 183

think is certainly an error. He also had specimens of glactana as
mengelana and inquietana, obvious misidentifications.

Male and female genitalia figured from specimens in Nationai
Collection from Massachusetts (paratype of fuscalbana, male) and
Hot Springs, Washington (female).

Bursa of female with signum a somewhat impressed scobinate
patch.

Distribution —North Carolina, Pennsylvania, New Jersey, Massa-
chusetts, New Hampshire, Nova Scotia, Labrador, Ontario, Alberta,
Saskatchewan, Manitoba, British Columbia, Washington, Colorado,
Arizona.

Alar expanse.—12-19.5 mm.

Types.—In collection Staudinger? (glaciana); British Museum
(dealbana, fuscalbana) ; Canadian National Collection (castorana).

Type localities——Labrador (glaciana) ; “ North America” (deal-
bana); Ohio (fuscalbana); Nordegg, Alberta (castorana).

21. OLETHREUTES NORDEGGANA (McDunnough)
(Figs. 255, 437)
Argyroploce nordeggana McDunnoucH, Can. Ent., vol. 54, 1922, p. 46.

A distinct species, so far represented only by the type material in
the Canadian National Collection.

Male and female genitalia figured from type and paratype.

Bursa of female with signum a slight, impressed, scobinate patch.

Alar expanse —20 mm.

Type.—tin Canadian National Collection.

Type locality —Nordegg, Alberta.

OLETHREUTES RIVULANA (Seopoli)

This European name should be dropped from our lists as the
species probably does not occur in North America. Nothing that
I have seen under the name in the collections agrees with European
specimens.

22. OLETHREUTES SCHULZIANA (Fabricius)

(Fig. 440)

Pyralis schulziana Fasrictus, Gen. Ins., 1777, p. 293.

Orthotaenia bentleyana Curtis, Appendix, Ross Second Arctic Voyage,
1835, p. 74.

Olethreutes schulziana STAUDINGER and REBEL, Cat. Lepid., vol. 2, no. 1912,
1901.—FERNALD, in Dyar List N. Amer. Lepid., no. 5076, 1903.—Forsgs,
Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 452.

Argyroploce schulziana BARNES and McDuNNouGH, Check List Lepid. Bor.
Amer., no. 6874, 1917.

Phiaris schulziana PIERcE and METCALFE, Genitalia Brit. Tort., 1922, p. 50,
Diels:

184 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

In genitalia schulziana and intermistana are very similar. The
European species is somewhat larger and often more reddish; but
otherwise I can see no real difference. The question of synonymy,
however, will have to remain in abeyance until a typical specimen
can be recovered from Arctic America. In the Canadian National
Collection there are a number of specimens from Quebec, Alberta,
and Quebec, Labrador under the name. They are quite dark how-
ever (blackish rather than red) and very variable. I have little
doubt but that they are color or local varieties of schulziana.

Male genitalia figured from European specimen in National Col-
lection.

Alar expanse.—18-25 mm.

Type.—Location unknown.

Type locality.—Europe.

Food plant.—Pinus sylvestris (in Europe).

23. OLETHREUTES INTERMISTANA (Clemens)
(Figs. 258, 441)

Penthina turfosana MéscHLrr (not Herrich-Schaefer) Ent. Monat., vol. 8,

1864, p. 199.

Mizodia ? intermistana CLEMENS, Proc. Ent. Soc. Philadelphia, vol. 5, 1865,
p. 140.

Penthina tessellana PacKarD, Proc. Boston Soe. Nat. Hist., vol. 11, 1867,
p. 58.

Olethreutes intermistana FERNALD, in Dyar List N. Amer. Lepid., no. 5028,
1903.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 452.

Olethreutes turfosana FERNALD, in Dyar List N. Amer. Lepid., no. 5065,
1903.—Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 452.

Argyroploce intermistana BaRNES AND McDunnoueu, Check List Lepid.
Bor. Amer., no. 6813, 1917.

Argyroploce turfosana BARNES AND McDuNnNoueH, Check List Lepid. Bor.
Amer., no. 6863, 1917.

In size, color, and general appearance intermistana and turfosana
are much alike. The two differ genitalically in the length of the
spined lower margin of the harpe, and there is no doubt of their being
distinct species. It is my opinion that the European species does not
occur in our fauna, and that what Méschler took for it was a speci-
men of tntermistana.

Male genitalia figured from specimen in Canadian Collection from
Labrador; female from specimen in National Collection from Mount
Washington, N. H. (“Aug. 1-7”).

Bursa of female with signum a week scobinate patch.

Distribution.—Labrador, Alberta, New Hampshire.

Alar expanse.—16-20 mm.

Types.—In Academy Natural Sciences Philadelphia (intermis-
tana) ; Museum Comparative Zoology (tessellana).

Type locality.—Labrador (intermistana, tessellana).

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 185

24. OLETHREUTES SEPTENTRIONANA (Curtis)
(Fig. 443)

Orthotenia septentrionana CurTIs, Ross Second Voyage N. W. Passage,
Appendix, 1831, p. 74.—WaLsINGcHAM, Illus. Lepid. Heter. Brit. Mus.,
vol. 4, 1879, p. 35.

? Sciaphila primariana WaAtker, Cat. Lepid. Heter. Brit. Mus., vol. 28,
1863, p. 336.

?Penthina fulvifrontana PackarpD, Proc. Boston Soc. Nat. Hist., vol. 11,
1867, p. 59.

Olethreutes septentrionana FERNAND, in Dyar List N. Amer., Lepid., no.
5029, 1903.—BarNES AND McDunnouGcH, Check List Lepid. Bor.
Amer., no. 6814, 1917.—ForBes, Memoir 68, Cornell Univ. Agr. Exp.
Sta., 1924, p. 452.

There is considerable doubt about the correctness of the above
synonymy. From Knight’s figure in the American Museum it does
not seem that primaviana can be the same as fulvifrontana; and it
is also a question whether either is equivalent to septentrionana.
Until some one can examine the genitalia of Walkers’ and Curtis’
types the synonymy may as well stand as it is.

O. fulvifrontana has genitalia similar to those of inguietana, ex-
cept that the spining of cucullus fuses more completely with that at
th: apex of the. extended sacculus, and that there is less of a break
between cucullus and extended sacculus than in the latter species.
The fore wing pattern reminds strongly of that of some specimens
of schulziana. Packard’s species, however, lacks any sign of a round
white dot at end of cell; the basal patch is well marked, strongly
angulate outwardly; the median dark band narrow on costa, broad
on dorsum and fused with subtornal spot, and has only one project-
ing tooth and that from middle: the apical dark markings are
somewhat variable, being different in type and paratype and even
on the right and left fore wings of the type; the dark markings are
a rick brown shaded with black, and the pale antemedian and post-
median areas a whitish ochreous.

Besides the type and paratype at Cambridge there is a broken
and faded paratype (female without abdomen) in the Academy of
Natural Sciences at Philadelphia, and two small Labrador speci-
mens from the Fernald collection in the National Museum.

Male genitalia figured from type (fulvifrontana).

Alar expanse.—14-15 mm.

Types.—? (septentrionana); in British Museum (primariana) ;
Museum Comparative Zoology (fulvifrontana).

Type localities—Arctic American (septentrionana, primariana) ;
Sloop Harbor, Labrador (fulvifrontana).

186 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

25. OLETHREUTES INQUIETANA (Walker)
(Figs. 252, 442)

Paedisca inquietana WaLKER, Cat. Lepid. Heter. Brit. Mus. vol. 28, 1863,

p. 378.
Sericoris inquietana WALSINGHAM, Illus. Lepid. Heter. Brit. Mus., vol.

4, 1879, p. 35.

Penthina septentrionana MOscHter, Stett. Ent. Zeit., vol. 44, 1883, p. 124.

Olethreutes boreana Reset, Stauding and Rebel, Cat. Lepid., vol. 2, no.
1913, 1901.

Olethreutes inquietana FERNALD, in Dyar List N. Amer. Lepid., no. 5074,
1903.

Argyroploce inquietana BARNES AND McDunnoucH, Check List Lepid.
Bor. Amer., no. 6872, 1917.

McDunnough has suggested the synonymy of boreana and inquie-
tana. In this he is most probably correct.

We have a single male in the National Collection (without locality
label) which agrees with specimens in the Canadian National Col-
lection from Greenland and Northwest Territory and with Knight’s
figure in the American Museum. There are also several specimens
from Greenland in.the Barnes collection.

Genitalia figured from specimens in National Collection (male)
and collection Barnes (female).

Bursa of female without signum.

Alar expanse.—24—-26 mm.

Types.—In British Museum (inguietana) ; collection Staudinger?
(boreana).

Type localities —* Arctic America” (tnguietana); Labrador
(boreana).

26. OLETHREUTES BOWMANANA (McDunnough)

(Fig. 447)
Argyroploce bownanana McDuNNouGH, Can. Ent., vol. 55, 1923, p. 165.

Similar to zntermistana in pattern and color, but with narrower
wings. Can be recognized by the strongly contrasted shining white
spot at upper outer angle of cell on fore wing.

Male genitalia of type figured.

In addition to the material in the Canadian National Collection
there is a male from Moraine Lake, Alberta (“’7—-VIII-1923”) in
the United States National Museum donated by Dr. McDunnough.

Alar expanse.—15 mm. .

Type.—tiIn Canadian National Collection.

Type locality —Nordegg, Alberta.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 187

27. OLETHREUTES MENGELANA (Fernald)
Figs. 251, 446)

Sericoris mengelana Frernaup, Ent. News., vol. 5, 1894, p. 131.

Penthina groenlandicana Bane-Haas, Vid. Medd., 1896, p. 190.

Olethreutes groenlandicana STaupINGER and Reset, Cat. Lepid., vol 2,
no. 1884, 1901.

Olethreutes mengelena STAUDINGER and REBEL, Cat. Lepid., vol. 2, no. 1885,
1901.—FERNALD, in Dyar List N. Amer. Lepid., no. 5059, 1903.—
Forses, Memoir 68, Cornell Univ. Agr. Exp. Sta., 1924, p. 452.

Argyroploce groenlandicana KENNEL, Palaeark. Tort., Lfg. 3, Zoologica, vol.
21, Heft 54, 1918, p. 377.

Argyroploce mengelana BaRNES and McDunnovueH, Check List Lepid. Bor.
Amer., no. 6840, 1917.—(?) Biackmorg, Report Proc. Mus. Nat. Hist.
British Columbia, 1921, pl. 4, p. 33.

There is an authentic male of groenlandicana from Greenland in
the Barnes collection, received from Bang-Haas. It agrees with
Fernald’s type of mengelana. Superficially it is much like the
Aphania of the tertiana-afficticia group; but is structurally quite
different and a good Olethreutes. The Barnes collection also has
two other males and a female, all from Greenland.

The male genitalia of this Band-Haas type is figured. Female
genitalia figured from specimen in National Collection, also from
Greenland.

Aside from the above (and the Fernald types), I have seen one
other example, in poor condition, in the American Museum. The
other specimens Kearfott had under the name are glaciana.

Alar expanse.—16-20 mm.

Types—tIn National Collection (mengelana) ; collection Staud-
inger (groenlandicana).

Type locality Greenland.

28. OLETHREUTES COSTIMACULANA (Fernald)
(Figs. 246, 421)

Penthina costimaculana FERNALD, Trans. Amer. Ent. Soc., vol. 10, 1882,
p. 70.

Olethreutes costimaculana FERNALD, in Dyar List N. Amer. Lepid., no.
5046, 1903.—Forses, Memoir 68 Cornell Univ. Agr. Exp. Sta., 1924,
p. 456.

Argyroploce costimaculana BARNES and McDunnovucGH, Check List Lepid.
Bor. Amer., no. 6846, 1917.

This species fits rather badly in the genus. In many ways—
especially on characters of the male genitalia—it fits better in
Hedia; but the absence of signa in the female bursa seems to forbid
such reference. Veins 8 and 9 of fore wing are more widely sepa-
rated, 2 is more bent than in other Olethreutes, and the spining on

54346—26——_13 ;

188 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

the uncus of the male genitalia is a little too strong, more like that
of typical Hedia.

Male and female genitalia figured from specimens in National
Collection from Meach Lake, Quebec (C. H. Young, “25-5-03,”
male), and Mer Bleue, Ottawa, Ontario (‘25-5-03 ” female).

Bursa of female without signum.

Distribution.—Maine, Ontario, Quebec, Manitoba, Alberta, Lab-
rador.

Alar expanse. —13-14 mm.

Type—tIn National Collection.

Type locality.—Maine.

OLETHREUTES ROSEOMACULANA (Herrich-Schaefer)

Is probably wrongly credited to North America and should be
omitted from our lists. Fernald’s costimaculana is very similar in
pattern and color and is very hkely what Moschler had from Labra-
dor. It averages a trifle smaller than typical specimens of roseo-
maculana, but could easily be mistaken for that species.

29. OLETHREUTES DEVOTANA Kearfett
(Figs. 250, 420)

Olethreutes devotana WKearrort, Trans. Amer. Ent. Soc., vol. 33, 1907,

p. 16.
Argyroploce devotana BARNES and McDunnoucH, Check List Lepid. Bor.

Amer., no. 6848, 1917.

This species also, like costimaculana, would seem rather to belong
in Hedia than Olethreutes. The uncus is rather strongly spined;
but the bursa shows no trace of signa.

Represented by the type material in the American Museum and
National Collections, and by specimens in the Barnes and National
Collections from Florida.

Male genitaha figured from type, female from paratype in Col-
lection Barnes from Hastings, Fla. (“ VI-8”).

Alar expanse.—14-15 mm.

T'ype.——In American Museum.

Type locality.—* Ch. Harbor, Fla.”

30. OLETHREUTES BUCKELLANA (McDunnough)
(Fig. 448)
Argyroploce buckellana McDuNNoucuH, Can. Ent., vol. 54, 1922, p. 48.

So far represented only by the Canadian Museum material, type
and one other male from the type locality. Easily recognized by
the male genitalia which has a short stiff spur projecting from sac-

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 189

culus near incurvation of neck. This is somewhat obscured in the
photograph by a slight tuft of hair.

Male genitalia of type figured.

Alar expanse.—15 mm.

Type.—tIn Canadian National Collection.

Type locality—Salmon Arm, British Columbia.

31. OLETHREUTES BUCKELLANA ALBIDULA, new variety
(Fig. 253)

Similar to buekellana but with antemedian and post median areas
of fore wing distinctly white.

Palpus white; terminal joint and end of second joint black.
Face white, with projecting scales above black. Head black with
a scattering of ferruginous and whitish scales. Thorax blackish
fuscous, faintly banded with white. Fore wing with dark areas
olivaceous fuscous faintly spotted with black; median band of
nearly uniform width throughout and including subtornal patch,
oblique, outer margin somewhat irregular, two small black dots in
middle; subapical bar weak, narrow, curved; a faint dark shading
at apex; termen ferruginous above tornus; cilia whitish, shaded
with ferruginous b yond base. Hind wing pale smoky fuscous;
cilia white with a dark basal band. Underside of hind wing con-
siderably paler than underside of fore wing, whitish.

Male genitaha as in buckellana.

Female genitalia figured from paratype in National Collection
from the type locality.

Bursa of female without signum.

Alar expanse.—15.5-17 mm.

Type.—tin collection Barnes.

Paratypes—Cat. No. 28042, U.S.N.M.; also in American Mu-
seum, Barnes and Canadian National coll ctions.

ype locality.—Inyo County, Calif.

Described from male type, 7 male and 4 female paratypes from
the type locality (“June 5-30-22,” “June 1-15,” “June 15-30,

1922,” O. G. Poling) ; two male paratypes from Elk Point, S., Dak.
(hue 1913, C. N. Ainslie); and one male paratype from Saskal om
Saskatchewan (“21—-VII-1923,” Kenneth M. King).

19. EVORA, new genus
(Figs. 55, 201)

Genotype —Euchromia hemidesma Zeller (North America).
Thorax with posterior tuft.

190 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

Fore wing smooth; termen straight; 12 veins, all separate; 8 and
9 separate; upper internal vein of cell from between 10-11; 3, 4, and
5 not approximate at termen; 2 from cell beyond 34, straight.

Hind wing with 8 veins; 6 and 7 approximate toward base; 3
and 4 connate; 5 approximate to 4; termen concave below apex; in
male a chitinous ridge on inner margin.

Hind tibia of male with yellow hair pencile.

Male genitalia with harpe elongate; broadened at base of cucullus;
outer surface unspined; cucullus short, strongly spined; spine
clusters Spe, Spe? strongly developed; sacculus weakly haired
toward base. Uncus short, broad, rounded, heavily spined beneath.
Socii short, flexible, hairy. Gnathos normal, a simple band with
weakly chitinized subanal plate. Aedoeagus fairly long, rather
stout, straight; cornuti absent.

Female genitalia without signum. Ductus bursae moderately
long, unchitinized except at genital opening.

A derivative of Olethreutes. Contains only the one North
American species.

The position of vein 2 of fore wing would seem to place it in the
Phaloniidae; but the typically olethreutine genitalia forbid. This
is the only olethreutid as far as I know which shows such a
character.

EVORA HEMIDESMA (Zeller)

(Figs. 55, 201, 411)

Euchromia hemidesma ZELLER, Verh. Zool.-bot. Ges. Wien, vol. 25, 1875, p. 261.

Olethreutes hemidesma FERNALD, in Dyar List N. Amer. Lepid., no. 5041,
1903.—KeEarrFotT, Can. Ent., vol. 37, 1907, p. 207.—Forses, Memoir 68,
Cornell Univ. Agr. Exp. Sta., 1924, p. 458.

Argyroploce hemidesma BaRrNes and McDuNNouGH, Check List Lepid. Bor.
Amer., no. 6856, 1917.

A reddish brown species with narrow, darker median band on
fore wing; somewhat resembling dark specimens of MAfelissopus
latiffereanus Walsingham.

Male and female genitalia figured from specimens in National
Collection from Cartwright, Manitoba (KE. F. Heath, male) and
Kentucky (August Busck, August, female).

Distribution——Maine, New Hampshire, Massachusetts, Connecti-
cut, New Jersey, Pennsylvania, Virginia, Kentucky, Illinois, Iowa,
Manitoba, Ontario.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 19]

Alar expanse.—13-17 mm.

Type.—tin British Museum.

Type locality —Massachusetts.

Food plants—Spiraea tormentosa, Solidago (larvae in flower

heads).
SPECIES REFERABLE ELSEWHERE

Aphelia? inquadrana WALSINGHAM, Trans. Ent. Soc. London, 1884, p 134.—
FERNALD, in Dyar List N. Amer. Lepid., no. 5008, 1903 (Bactra).—
Barnes and McDunnovueH, Check List Lepid. Bor. Amer., no 6791,
1917 (Bacira).

A narrow-winged Hucosma similar in pattern to pulveratana
Walsingham, which species it precedes in our arrangement. The
resemblance to Bactra is only superficial, and there is a well-marked
costal fold which Walsingham overlooked. A series of males from
Sells Post Office, Pima County, Arizona (April-May, 1923), has
been received recently through Dr. William Barnes. These are the
first authentic specimens I have seen in any American collection.

The following species now listed with the Olethreutinae are ref-
erable to the Tortricidae. The numbers before each are those of
the Barnes and McDunnough list.

6820—Olethreutes wellingtoniana Krarrott, Trans. Amer. Ent. Soc., vol. 33,

190 Ds 13:

6845—Olethreutes gogana Krarrott, Trans. Amer. Ent. Soc., vol. 33, 1907,
p. 8.

6875—Olethreutes provana KEARFOTT, Trans. Amer. Ent. Soc., vol. 33, 1907,
Dp: 16:

EXPLANATION OF PLATES

The drawings accompanying this paper were made under the author’s
supervision by Miss Eleanor Armstrong and Mr. Harry Bradford, of the
Bureau of Entomology. The photographs were taken by Mr. J. G. Pratt, of
the Bureau of Entomology.

Terms used in description of male genital organs:

Dsp=spines arising from outer surface of harpe.

Gn=Gnathos.

ScSp=spines on base of sacculus.

Si=socii.

Spc’ and Spe’*=heavy spine clusters on or near sacculus of harpe.
U=Uncus.

X=spine group on arch of sacculus (in Hxvartema).

192

Fic.

Fie.

(OU Ne) os

10.

BULLETIN 132, UNITED STATES NATIONAL MUSEUM

JPipyNaug) <1

Structural characters in Laspeyresiinae

. Metathoracic leg of Gymnandrosoma punctidiscanum Dyar, Male.
. Metathoracic leg of Melissopus latiferreanus (Walsingham), Male.
. Partially denuded male hind wing of G. punctidiscanum, showing

venation, pocket-like development of inner margin, and sex scaling.

. Hind wing venation of Ricula maculana (Fernald), Male.
. Abdomen of G. punctidiscanum (male), dorsal view showing pair of

hair tufts from caudal edge of second segment.

. Hind wing venation (male) of Hemimene populana (Fabricius).
. Caudal segments of denuded male abdomen of Grapholitha fana (Kear-

fott), Showing hair tufts from sternite of eighth segment.

. Hind wing venation (male) of Dichrorampha kana (Busck).
. Denuded hind wing (male) of WM. latiferreanus, showing venation,

thickening at inner margin, and pocket inclosing pecten.
Denuded hind wing (male) of Medytolopha insiticiana Zeller, showing
venation and pocket and hair pencile on vein 1°.

PLATE 2

Structural and pattern characters in Olethreutinae

. Inside view of metathoracie leg of male Exartema showing hair pencile

from base of tibia.

. Fore wing venation of Polychrosis viteana (Clemens).
. Fore wing venation of Olethreutes arcuella (Clerek).
. Fore wing of Hxartema electrofuscwm Heinrich, showing characteristic

Olethreutid pattern: B=basal patch; I/=median band; St=sub-
tornal spot; Sa=subapical bar.

. Anterior segment of denuded male abdomen of Polychrosis cypripediana

Forbes, showing pockets (#) of papilliform hairs on basal segment.

. Metathoracie leg (male) of Phaecasiophora confixana (Walker).
. Head and thorax of Sciaphila duplex (Walsingham) ; side view show-

ing posterior thoracic tuft (TT).

. Denuded male hind wing of Hxartema concinnanum (Clemens) show-

ing venation and anal lobe.

. Fore wing venation of Polychrosis botrana (Schiffermtiller).

. Hind wing venation of P. botianda.

. Fore wing venation of Hpisimus transferranus (Walker).

. Fore wing of EH. transferranus, showing characteristic Hpisimus pattern.

PLATE 3

Male genitalia (Laspeyresiinae)

. Satronia tantilla Heinrich.

. Goditha bumeliana Heinrich.

. Rieula maculana (Fernald).

. Ethelgoda taxanana (Walsingham).
. Hemimene populana (Fabricius).

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 193

PLATE 4
Male genitalia (Laspeyresiinae )

4G. 28. Dichrorampha plumbagana (Treitschke).
29. Sereda lautana (Clemens).
30. Ofatulena duodecemsiriata (Walsingham).

PLATE 5

Male genitalia (Laspeyresiinae)

ic. 31. Tegumen and aedoeagus of Melissopus latiferreanus (Walsingham) ;
variety G.
32. Tegumen and aedoeagus of WM. latiferreanus; variety B.
33. Tegumen and aedoeagus of M. latiferreanus; variety C.
34. Tegumen and aedoeagus of M. latiferreanus; variety D.
35. Tegumen and aedoeagus of M. latiferreanus; variety PF.
36. Melissopus latiferreanus (Walsingham) ; variety A.
37. Carpocapsa pomonella (Linnaeus).
38. Gymnandrosoma punctidiscanum Dyar.

PLATE 6
Male genitalia (Olethreutinae)

Fic. 39. Episimus transferranus (Walker).
40. Olethreutes arcuella (Clerck).

PLATE 7
Male genitalia (Olethreutinae)

Fic. 41. Polychrosis botrana (Schiffermitiller).
42. Exartema nitidanum Clemens. ;

PLATE 8

Male genitalia (Olethreutinae)

Fie. 43. Badebecia urticana (Htibner).
44, Bactra lanceolana (Hitibner) ; right harpe.
45. Bactra furfurana (Haworth).
46. Bactra verutanea albipuncta Heinrich ; right harpe.
47. Bactra verutana verutana Zeller; right harpe.
48. Endothenia gentianana (Htibner).
49. Bactra verutana chrysea Heinrich; right harpe.

PEATE; 9
Male genitalia (Olethreutinae)

Fic. 50. Taniva albolineana (Kearfott).
51. Sciaphila duplex (Walsingham).

194 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

PLATE 10

Male genitalia (Olethreutinae)

Fig. 52. Hulda impudens (Walsingham).
53. Tia vulgana (McDunnough).
54. Aphania scriptana (Hiibner).

PLATE 11

Male genitalia (Olethreutinae)

Fie. 55. Evora hemidesma (Zeller).
56. Hsia approximana (Heinrich).
57. Eccopsis wahlbergiana Zeller.

PrArn sl?

Male genitalia (Olethreutinae)

Fic. 58. Ahmosia galbinea Heinrich.
59. Zomaria interruptolineana (Fernald).
60. Humarozia malachitana (Zeller).

PLATE 13

Male genitalia (Olethreutinae)

Fic. 61. Phaecasiophora confirtana (Walker).
62. Hedia salicella (Vinnaeus).

PLATE 14

Right harpes of male genitalia (Hzartemu)

Fic. 63. Hxartema zellerianum (Fernald).

64. Hxartema nitidanum Clemens.

65. Exartema foedanum (Clemens).
66. Ezrartema olivaceanum (Fernald).
67. EHxartema cornanum Heinrich.

68. Havartema subnubilum Heinrich.
69. Hxeartema inornatanum Clemens.
70. Hxartema monetiferanum Riley.
71. Exartema punctanum Walsingham.

PLATE 15
Right harpes of male genitalia (Hawartema)

Fig. 72. Hxartema mediopartitum Heinrich.
73. Exartema exoletum Zeller.
74. Exartema tenebricum Heinrich.
75. Hxartema electrofuscum Heinrich.
76. Exartema footianum (Fernald).
77. Heartema atrodentanum (Fernald).
78. Exartema furfuranum MeDunnough.
79. Exartema rusticanum McDunnough.
80. Exartema clavanum (Walker).

Fig.

Fic.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE

81.

98.
99.
100.
101.
102.
103.

104.
105.
106.
107.
108.
109.
110.
111.

PLATE 16
Right harpes of male genitalia (Hzartema)

Exartema trepidulum Heinrich.

. Hxartema malanum (Fernald).

. Exartema melanomesum Heinrich.

. Hzartema corylanum (Fernald).

. Hzartema submissanum McDunnough.
. Hxzartema nigranum Heinrich.

. Hzartema quadrifidum Zeller.

. Exzartema hippocastanum Kearfott.

PLATE 17

Right harpes of male genitalia (Hzartema)

. Hxzartema valdanum McDunnough.

. Hxartema permundanum Clemens.

. Hzartema merrickanum Kearfott.

. Hzartema fagigemmeanum Chambers.
. Exartema sciotanum Heinrich.

. Hzartema ferrugineanum Riley.

. Hxartema sericoranum Walsingham.
. Hzartema ochrosuffusanum Heinrich.
. Hxartema tilianum Heinrich.

PLATE 18
Right harpes of male genitalia (Hzartema)-

EHzrartema appendiceum Zeller.
Heartema concinnanum Clemens.
Eaartema ferriferanum (Walker).
EHaartema exaeresimum Heinrich.
Ezartema fasciatanum Clemens.
EHeartema troglodanum McDunnough.

PLATE 19
Female genitalia (Dichrorampha, Ricula)

Dichrorampha bittana (Busck).
Dichrorampha leopardana (Busck).
Ricula maculana (Fernald).

Dichrorampha sedatana (Busck).
Dichrorampha incanana (Clemens).
Dichrorampha radicicolana (Walsingham).
Dichrorampha dana (Kearfott).
Dichrorampha capitana (Busck).

195

196 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

PLATE 20
Female genitalia (Melissopus, Talponia, Eedytolopha, Hemimene)

Hig, 112. Melissopus latiferreanus (Walsingham) ; variety £.
113. Melissopus latiferreanus (Walsingham) ; variety A.
114. Talponia plummeriana (Busck).

115. HEcdytolopha insiticiana Zeller.
116. Hemimene felicitana (Heinrich).
117. HEcdytolopha mana (Kearfott).

PLATE 21
Female genitalia (Sereda, Ofatulena, Gymnandrosoma, Ethelgoda)

Fig. 118. Sereda lautana (Clemens).
119. Ofatulena duodecemstiata (Walsingham).
120. Ofatulena luminosa Heinrich.
121. Gymnandrosoma punctidiscanum Dyar.
122. Hthelgoda texanana (Walsingham).
123. Gymnandrosoma desotana Heinrich.

PLATE 22
Female genitalia (Grapholitha)

Fic. 124. Grapholitha eclipsana Zeller.
125. Grapholitha caeruleana Walsingham.
126. Grapholitha fana (Kearfott).
127. Grapholitha angleseana (Kearfott).
128. Grapholitha lunatana Walsingham.
129. Grapholitha molesta (Busck).
130. Grapholitha prunivora (Walsh).
1381. Grapholitha vitrana Walsingham.
182. Grapholitha packardi Zeller.
133. Grapholitha conversana Walsingham.
134. Grapholitha imitativa Heinrich.

PLATE 23
Female genitalia (Grapholitha, Laspeyresia)

Fie. 185. Grapholitha dyarana (Kearfott).
136. Grapholitha tristrigana (Clemens).
137. Grapholitha lana (Kearfott).
138. Laspeyresia youngana (Kearfott).
139. Grapholitha interstinctana (Clemens).
140. Laspeyresia nigricana (Stephens).
141. Laspeyresia erotella (Heinrich).
142. Laspeyresia flexiloqua Heinrich.

Fig.

Fic.

Fie.

Fic.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE

143.

144.
145.
146.
147.
148.
149.
150.

151.
152.
153.
154.
155.
156.
157.
158.

159.
160.
161.
162.
163.

164.

165.
166.
167.
168.
169.

170.
171.
172.
173.
174.

PLATE 24
Female genitalia (Laspeyresia)

Laspeyresia garacana (Kearfott).
Laspeyresia prosperana (Kearfott).
Laspeyresia populana Buseck.
Laspeyresia caryana (Fitch).
Laspeyresia albimaculana (Fernald).
Laspeyresia ninana (Dyar).
Laspeyresia flavicollis (Walsingham).
Laspeyresia membrosa Heinrich.

PLATE 25
Female genitalia (Laspeyresia)

Laspeyresia bracteatana cornutana (Dyar).
Laspeyresia candana Forbes.

Laspeyresia rana Forbes.

Laspeyresia cupressana (Kearfott).
Laspeyresia laricana Busck.

Laspeyresia tana (Kearfott).

Laspeyresia leucobasis Busck.

Laspeyresia americana (Walsingham).

PLATE 26

Female genitalia (Laspeyresia, Hedulia )

Laspeyresia toreuta (Grote).
Laspeyresia miscitata Heinrich.
Laspeyresia ingens Heinrich.
Hedulia injectiva Heinrich.
Laspeyresia piperana (Kearfott).

PLATE 27

Female genitalia (Laspeyresia, Carpocapsu)

Laspeyresia obnisa Heinrich.
Laspeyresia inopiosa Heinrich.
Laspeyresia gallaesaliciana (Riley).
Laspeyresia colorana (Kearfott).
Laspeyresia lautiuscula Heinrich.
Carpocapsa pomonella (Linnaeus).

PLATE 28
Female genitalia (Bactra)

Bactra furfurana (Haworth).
Bactra verutana Zeller.
Bactra priapeia Heinrich.
Bactra maiorina Heinrich.
Bactra sinistra Heinrich.

197

198 BULLETIN 1382, UNITED STATES NATIONAL MUSEUM
PLATE 29

Female genitalia (Polychrosis)

Fie. 175. Polychrosis liriodendrana Kearfott.
176. Polychrosis rhoifructana Keurfott.
177. Polychrosis vernoniana Kearfott.
178. Polychrosis spiraeifoliana Heinrich.
179. Polychrosis slingerlandana Kearfott.
180. Polychrosis aemulana Heinrich.
181. Polychrosis carduana Busck.
182. Polychrosis viteana (Clemens).

PLATE 30

Female genitalia (Polychrosis, Episimus, Ahmosia)

Fie. 1838. Episimus argutanus (Clemens).
184. Ahmosia aspasiana (McDunnough).
185. Polychrosis cyclopiana Heinrich.
186. Ahmosia galbinea Heinrich.

187. Hpisimus tyrius Heinrich.

PLATE 31

Female genitalia (Taniva, Endothenia)

Fic. 188. Hndothenia hebesana (Walker).
189. Taniva albolineana (Kearfott).
190. Endothenia montanana (Kearfott).
191. Endothenia melanosticta (Walsingham).
192. HEndothenia antiquana nubilana (Clemens).

PLATE 32
Female genitalia (Hndothenia, Hulda, Tia, Eumarozia)

Fie. 1938. Hulda impudens (Walsingham).
194. Humarozia malachitana (Zeller).
195. Tia vulgana (McDunnough).

196. Endothenia sordulenta Heinrich.
197. Hndothenia rubipuncta (Kearfott).

PLATE 33

Female genitalia (Zomaria, Evora, Esia)

Fie. 198. Zomaria andromedana (Barnes and McDunnoug)).
199. Zomaria interruptolineana (Fernald).
200. Zomaria rosaochreana (Kearfott).
201. Hvora hemidesma (Zeller).
202. Hsia approximana (Heinrich).

Fig.

Fic.

EIG.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE

203.
204.
205.
206.
207.
208.

209.
210.
211.
212.
213.
214.
215.
216.
217.

218.
219.
220.
221.
222.
223.
224.
225.
226.

227.
228.
229.
230.
21.
232.
233.

Eazartema

PLATE 34

Female genitalia (Hzartema)

zellerianum (Fernald).

Ezartema nitidanum Clemens.

Exzartema
Eeartema
EHzartema
Ezartema

Eaartema
Ezxartema
EHzartema
Ezartema
Ezartema
Exzartema
Ezartema
Ezartema
Exzartema

Hxeartema
Exartema
Ezartema
Ezartema
Ezxartema
Ezartema
Ezartema
Ezxartema
Ezxartema

footianum (Fernald).
versicoloranum Clemens.
sciotanum Heinrich.
brunneopurpuratum Heinrich.

PLATE 35

Female genitalia (Hzartema)

foedanum (Clemens).
clavanum (Walker).
olivaceanum (Fernald).
punctanum Walsingham.
inornatanum Clemens.
tilianum Heinrich.
electrofuscum Heinrich:
subnubilum Heinrich.
atrodentanum (Fernald).

PLATE 36

Female genitalia (Hzartema)

malanum (Fernald).
troglodanum McDunnough.
ferriferanum (Walker).
monetiferanum Riley.
merrickanum Kearfott.
ferrugineanum Riley.
hippocastanum Kearfott.
permundanum Clemens.
quadrifidum Zeller.

PLATE 37

Female genitalia (H«xartema, Phaecasiophora)

Exartema exoletum Zeller.
Ezartema corylanum (Fernald).
Ezartema concinnanum (Clemens).
Exartema appendiceum Zeller.
Phaecasiophora niveiguttana Grote.
Ezartema fasciatanwm Clemens.
Phaecasiophora confizana (Walker).

199

200

Vic.

Fic.

ic.

Fig.

Ia.

234.

235.
236.
237.
238.

239.

240.
241.
242.
243.

244.

245.

246.
247.
248.
249.

254.
255.
256.
257.
258.

BULLETIN

Olethreutes
Olethreutes
Olethreutes
Olethreutes
Olethreutes

Olethreutes
Olethreutes
Olethreutes
Olethreutes
Olethreutes
Olethreutes

. Olethreutes
. Olethreutes
. Olethreutes
. Olethreutes

132, UNITED STATES NATIONAL MUSEUM
PLATE 38

Female genitalia (Olethreutes)

osmundana (Fernald).
agilana (Clemens).
griseoalbana (Walsingham).
albiciliana (Fernald).
auricapitana (Walsingham).

PLATE 39

Female genitalia (Olethreutes)

astrologana (Zeller).
constellatana (Zeller).
galazana Kearfott.
coruscana (Clemens).
carolana (MecDunnough).

PLATE 40

Female genitalia (Olethreutes)

polluxana (McDunnough).
cespitana (Hubner).
costimaculana (Fernald).
deprecatoria Heinrich.
glaciana (Moéschler).
bipartitana (Clemens).

PLATE 41

Female genitalia (Olethreutes)

devotana Kearfott.
mengelana (Fernald).
inquietana (Walker).
buckellana albidula Heinrich.

PLATE 42

Female genitalia (Olethreutes, Badebecia, Sciaphilu)

Olethreutes puncticostana (Walker).
Olethreutes nordeggana (McDunnough).
Badebecia urticana (Hiibner).

Sciaphila duplex (Walsingham).
Olethreutes intermistana (Clemens).

NORTH AMERICAN LASPEYRESHINAE AND OLETHREUTINAR 201
PLATE 43
Female genitalia (Hedia)

Wie. 259. Hedia cyanana (Murtfeldt).
260. Hedia? lineana (Fernald).
261. Hedia chionosema (Zeller).
262. Hedia variegana (Hiibner).
263. Hedia separatana (Kearfott).
264. Hedia ochroleucana (Hiibner).

PLATE 44
Female genitalia (Aphania)

Fig. 265. Aphania capreana (Hiibner).
266. Aphania removana (Kearfott).
267. Aphania frigidana (Packard).
268. Aphania deceptana (Kearfott).

PLATE 45
Female genitalia (Aphania)

Fic. 269. Aphania apateticana (McDunnough).
270. Aphania youngana (McDunnough).
271. Aphania infida Heinrich.
272. Aphania albeolana (Zeller).

PLATE 46
Male genitalia (Dichroramphea)

Fie. 273. Dichrorampha kana (Busck).
274. Dichrorampha capitana (Busck).
275. Dichrorampha britana (Busck).
276. Dichrorampha simulana (Clemens).

PLATE 47

Male genitalia (Dichrorampha)

Fic. 277. Dichrorampha bittana (Busck).
278. Dichrorampha banana (Busck).
279. Dichrorampha incanana (Clemens).
280. Dichrorampha leopardana (Busck).
281. Dichrorampha piperana (Busck).
282. Dichrorampha radicicolana Walsingham.
283. Dichrorampha sedatana (Busck).

202 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

PLATE 48

Male genitalia (Ricula, Satronia, Talponia, Ethelgoda, Sereda, Hemimene,
Goditha)

Fic. 284. Ricula maculana (Fernald).
285. Satronia tantilla Heinrich.
286. Talponia plummeriana (Buseck).
287. EHthelgoda texanana (Walsingham).
288. Sereda lautana (Clemens).
289. Hemimene paula Heinrich.
290. Hemimene signifera Heinrich.
291. Hemimene ocliferia Heinrich.
292. Hemimene felicitana (Heinrich).
293. Goditha bumeliana Heinrich.

PLATE 49
Male genitalia (Grapholitha)

Fic. 294. Grapholitha libertina Heinrich.
295. Grapholitha packardi Zeller.
296. Grapholitha prunivora (Walsh).
297. Grapholitha fana (Kearfott).
298. Grapholitha imitativa Heinrich.
299. Grapholitha caeruleana Walsingham.
300. Grapholitha lunatana Walsingham.
801. Grapholitha conversana Walsingham.
302. Grapholitha eclipsana Zeller.
303. Grapholitha vitrana Walsingham.

PLATE 50
Male genitalia (Grapholitha, Ofatulena)

Fic. 304. Grapholitha angleseana (Kearfott).
305. Grapholitha molesta (Busck).
306. Grapholitha interstinctana (Clemens).
307. Grapholitha tristrigana (Clemens).
808. Grapholitha lana (Kearfott).
309. Ofatulena duodecemstriata (Walsingham).
310. Ofatulena luminosa Heinrich.

PLATE 51
Male genitalia (Laspeyresia)

Fie. 311. Laspeyresia bracteatana (Fernald).
312. Laspeyresia bracteatana (Fernald). (=pallidibasalis.)
313. Laspeyresia garacana (Kearfott).
314. Laspeyresia rana Forbes.
315. Laspeyresia ingrata Heinrich.
316. Laspeyresia larimana (Walsingham).
317. Laspeyresia laricana Busck.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE

Fie. 318.
319.
320.
321.

Fie. 322.

Fi@. 326.
327.
328.
329.

Fie. 330.
331.
332.
333.
334.

Fig. 335.
. Laspeyresia americana (Walsingham).
. Laspeyresia colorana (Kearfott).

Fie. 338.

339.
340.
341.

54346—26

PLATE 52
Male genitalia (Laspeyresia )
Laspeyresia multilineana (Kearfott).
Laspeyresia populana Busck.
Laspeyresia grandicula Heinrich.
Laspeyresia fletcherana (Kearfott).
PuATH 5s

Male genitalia (Laspeyresia)

Laspeyresia memborsa Heinrich.

. Laspeyresia caryane (Fitch).
. Laspeyresia prosperana (Kearfott).
. Laspeyresia candana Forbes.

PLATE 54
Male genitalia (Laspeyresia, Carpocapsa)

Laspeyresia cupressana (Kearfott).
Laspeyresia tana (Kearfott).
Laspeyresia nigricana (Stephens).
Carpocapsa pomonella (Linnaeus).

PLatTe 55
Male genitalia (Laspeyresia, Hedulia)

Laspeyresia youngana (Kearfott).
Laspeyresia gallaesaliciana (Riley).
Laspeyresia leucobasis Busck.
Laspeyresia piperana (Kearfott).
Hedulia injectiva Heinrich.

PLATE 56
Male genitalia (Laspeyresia)

Laspeyresia ninana (Dyar).

PLATE 57
Male genitalia (Laspeyresia, Eeéytolopha, Gymnandrosoma>

Laspeyresia erotella (Heinrich).
Ecdytolopha islandana (Kearfott).
Ecdytolopha insiticiana Zeller.
Gymnandrosoma punctidiscanum Dyar.

203

204

Fig. 342.
343.
344.
345.
346.
347.
348.
349.

Fig. 350.
351.
352.
353.
354.
355.
356.

357.

Fie. 358.
359.
360.
361.
362.
363.
364.

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BULLETIN 182, UNITED STATES NATIONAL MUSEUM

PLATE 58
Male genitalia (Bactra, Endothenia)

Bactra lanceolana (Hiibner).

Bactra furfurana (Haworth).
Bactra maiorina Heinrich.

Bactra priapeia Heinrich.

Bactra verutana Zeller (typical).
Bactra verutana albipuncta Heinrich.
Bactra verutana chrysea Heinrich.
Endothenia montanana (Kearfott).

PLATE 59
Male genitalia (Hndothenia)

Endothenia hebesana (Walker).
Endothenia daeckeana (Kearfott).

Endothenia
Endothenia
Endothenia

sordulenta Heinrich.
melanosticta (Walsingham).
conditana (Walsingham).

Endothenia infuscata Heinrich.

Endothenia
Endothenia

Polychrosis
Polychrosis
Polychrosis
Polychrosis
Polychrosis
Polychrosis
Polychrosis

Polychrosis

. Polychrosis
. Polychrosis
. Polychrosis
. Polychrosis
. Polychrosis
. Polychrosis
. Polychresis

rubipuncta (Kearfott).
antiquana nubilana (Clemens).

PLATE 60
Male genitalia (Polychrosis)

botrana (Schiffermiiller).
slingerlandana Kearfott.

spiraeifoliana Heinrich.

liriodendrana Kearfott. ‘
viteana (Clemens).

carduana Buseck.

cyclopiana Heinrich.

PLATE 61
Male genitalia (Polychrosis)

aruncana Kearfott.
monotropana Heinrich.
cypripediana Forbes.
blandula Heinrich.
aemulana Heinrich.
yarakana Kearfott.
vernoniana Kearfott.
rhoifructana Kearfoot.

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 205
PLATE 62

Male genitalia (Aphania)

Fic. 373. Aphania deceptana (Kearfott) ; aedoeagus and anellus omitted.
374. Aphania youngana (McDunnough).
375. Aphania capreana (Htibner).

PLATE 63
Male genitalia (Aphania)

Hie. 376. Aphania frigidana (Packard).
377. Aphania tertiana (McDunnough).
378. Aphania afficticta Heinrich.

PLATE 64
Male genitalia (Aphania)

Fig. 379. Aphania apateticana (McDunnough).
380. Aphania infida Heinrich.
381. Aphania removana (Kearfott).

PLATE 65
Male genitalia (Aphania)

Fig. 882. Aphania strigosa Heinrich.
383. Aphania albeclana (Zeller).
384. Aphania dextrana (MceDunnough).

PLATE 66
Male genitalia (Ahmosia, Sciaphila, Hulda, Taniva, Lovoterma, Badebecia)

Fie. 385. Ahmosia aspasiana (McDunnough).
386. Ahmosia galbinea Heinrich.
387. Sciaphila duplex (Walsingham).
388. Hulda impudens (Walsingham).
389. Taniva albolineana (Kearfott).
390. Lozoterma latifasciana (Haworth).
391. Badebecia urticana (Htibner).

PLATE 67
Male genitalia (Hpisimus, Phaecasiophora, Cymolomia, Eccopsis)

Fig. 392. Episimus argutanus (Clemens).
393. Episimus augmentanus (Zeller).
394. Phaecasiophora confizana (Walker). =
395. Phaecasiophora niveiguttana Grote.
396. Cymolomia hartigiana (Ratzeburg).
397. Eccopsis wahlbergiana Zeller.

206 BULLETIN 132, UNITED STATES NATIONAL MUSEUM

PLATE 68
Male genitalia (Hrartema)

Fic. 398. Hxartema tilianum Heinrich.
399. Havartema nitidanum (Clemens).
400. Hzartema olivaceanum (Fernald).
401. Hxartema punctanum (Walsingham).

PLATE 69
Male genitalia (H.wrartema)

Fie. 402. Hxrartema appendiceum Zeller.
403. Exartema concinnanum (Clemens).
404. Exartema fasciatanum Clemens.
405. Exartema troglodanum McDunnough.
406. Hxartema eraeresimum Heinrich.
407. Exartema ferriferanum (Walker).

PLATE 70
Male genitalia (Zomaria, Evora, Tia, Eumarozia, Hsia, Hedia)

Fie. 408. Zomaria interruptolineana (Fernald).

409. Zomaria rosaochreana (Kearfott).
410. Zomaria andromedana (Barnes and McDunnough).
411. Evora hemidesma (Zeller).

- 412. Tia vulgana (McDunnough).
413. Humerozia malachitana (Zeller).
414. Hsia approzimana (Heinrich).
415. Hedia cyanana (Murtfeldt).

PLATE 71
Male genitalia (Hedia, Olethreutes)

Fic. 416. Hedia separatana (Wearfott).
417. Hedia ochroleucana (Hiibner).
418. Hedia variegana (Hiibner).
419. Hedia chinosema (Zeller).
420. Olethreutes devotana Kearfott.
421. Olethreutes costimaculana (¥ernald).

PLATE 72
Male genitalia (Olethreutes)

Fig. 422. Olethreutes griseoalbana (Walsingham).
423. Olethreutes osmundana (Fernald).
424. Olethreutes auricapitana (Walsingham).
425. Olethreutes albiciliana (Fernald).
426. Olethreutes siderana (Treitschke) ; Huropean specimen.
427. Olethreutes siderana chalybeana (Walsingham).

NORTH AMERICAN LASPEYRESIINAE AND OLETHREUTINAE 207

PLATE 73
Male genitalia (Olethreutes)

Fic. 428. Olethreutes sordidana (McDunnough).
429. Olethreutes constellantana (Zeller).
430. Olethreutes coruscana (Clemens).

431. Olethreutes galarana Kearfott.
432. Olethreutes glaciana (Moschler).
433. Olethreutes astrologana (Zeller).

PLATE 74
Male genitalia (Olethreutes)

Fie. 434. Olethreutes cespitana (Hiibner).
435. Olethreutes bipartitana (Clemens).
436. Olethreutes deprecatoria Heinrich,
437. Olethreutes nordeggana (McDunnough).
438. Olethreutes carolana (McDunnough).
439. Olethreutes agilana (Clemens).

PLATE 75
Male genitalia (Olethreutes)

Fig. 440. Olethreutes schulziana (Fabricius).
441. Olethreutes intermistana (Clemens).
' 442. Olethreutes inquietana (Walker).
448. Olethreutes septentrionana (Curtis) (=fulvifrontana Packard).
444. Olethreutes puncticostana major (Walsingham).
445. Olethreutes polluxana (McDunnough).

PLaTE 76
Male genitalia (Olethreutes)

Fie. 446. Olethreutes mengelana (Fernald) (=groenlandicana Bang-Haas).
447. Olethreutes bowmanana (McDunnough).
448. Olethreutes buckellana (McDunnough).

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U. S. NATIONAL MUSEUM

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U. S. NATIONAL MUSEUM ; BULLETIN 132 PL.

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STRUCTURAL AND PATTERN CHARACTERS IN OLETHREUTINAE

FOR EXPLANATION OF PLATE SEE PAGE 192

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 3

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MALE GENITALIA OF LASPEYRESIINAE

FOR EXPLANATION OF PLATE SEE PAGE 192

U. S. NATIONAL MUSEUM BULLETIN 132 PL.

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MALE GENITALIA OF LASPEYRESIINAE

FOR EXPLANATION OF PLATE SEE PAGE 193

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 5

36 Melhssopus

Sy) Carpocapsa

Li id

58 Gymnandrosoma .

MALE GENITALIA OF LASPEYRESIINAE

FOR EXPLANATION OF PLATE SEE PAGE 193

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 6

J9 Ie) pisimus

AQ Olethreutes

MALE GENITALIA OF OLETHREUTINAE

FOR EXPLANATION OF PLATE SEE PAGE 193

Hes 7

132

BULLETIN

U. S. NATIONAL MUSEUM

Folychrosis

Exartema

MALE GENITALIA OF OLETHREUTINAE

FOR EXPLANATION OF PLATE SEE PAGE 193

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 8

ot wie ie = NN ¥ | ; -\ 46 albipuncta
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MALE GENITALIA OF OLETHREUTINAE

FOR EXPLANATION OF PLATE SEE PAGE 193

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 9

Sy! we raphila

MALE GENITALIA OF OLETHREUTINAE

FOR EXPLANATION OF PLATE SEE PAGE 193

U. S. NATIONAL MUSEUM BULLETIN 132 PL. [0

SA = Agpeaa

MALE GENITALIA OF OLETHREUTINAE

FOR EXPLANATION OF PLATE SEE PAGE 194

U. S. NATIONAL MUSEUM BULLETIN 132 PL. II

= Z ou se = ewe
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MALE GENITALIA OF OLETHREUTINAE

FOR EXPLANATION OF PLATE SEE PAGE 194

U. S. NATIONAL MUSEUM

IS Ahmosia

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MALE GENITALIA OF OLETHREUTINAE

FOR EXPLANATION OF PLATE SEE PAGE 194

Eumarozia

BULLETIN

132

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U. S. NATIONAL MUSEUM BULLETIN 132

PL. 13

MALE GENITALIA OF OLETHREUTINAE

For EXPLANATION OF PLATE SEE PAGE 194

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 14

635 zellerianum 64

nitidanum

6 7 cornmanum

69 inornatanum 10 ~monetiferanum Wy punctanum

RIGHT HARPES OF MALE GENITALIA OF EXARTEMA

FOR EXPLANATION OF PLATE SEE PAGE 194

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 15

72 mediopa rbitum
735 exoletum 74 tenebricurn

7T71~— atrodentanum

78 furfuranum 79° rusticanum 80 clavanum

RIGHT HARPES OF MALE GENITALIA OF EXARTEMA

FOR EXPLANATION OF PLATE SEE PAGE 194

U. S. NATIONAL MUSEUM BULLETIN 132 PL. I6

82 malanum

3835 melanomesum

3! trepidulum

SS hippocastanum

RIGHT HARPES OF MALE GENITALIA OF EXARTEMA

FOR EXPLANATION OF PLATE SEE PAGE 195

U. S. NATIONAL MUSEUM BULLETIN 132 PL. IT

95 serricoranum 96 ochrosuffusonum 97 Khanum

RIGHT HARPES OF MALE GENITALIA OF EXARTEMA
FOR EXPLANATION OF PLATE SEE PAGE 195

54546—26——16

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 18

98 appendiceum

99 concinnanum

103

RIGHT HARPES OF MALE GENITALIA OF EXARTEMA

10! exaeresimum troglo danum

FOR EXPLANATION OF PLATE SEE PAGE 195

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 19

maculana

04 bittona
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capitana

& \

FEMALE GENITALIA OF DICHRORAMPHA AND RICULA

FOR EXPLANATION OF PLATE SEE PAGE 195

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 20

D> ee 14 SS

lohferreanus < / plummeriana

\ 16

felcitana

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INSItIC/ana

GENITALIA OF MELISSOPUS, TALPONIA, ECDYTOLOPHA, AND HEMIMENE

FOR EXPLANATION OF PLATE SEE PAGE 196

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 21

8 leutana

punctidiscanum

bs we ;
Hye,
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Bact ad

l2? texanona

GENITALIA OF SEREDA, OFATULENA, GYMNANDROSOMA, AND ETHELGODA

FOR EXPLANATION OF PLATE SEE PAGE 196

U. S. NATIONAL MUSEUM BULLETIN 132

Pie?)

125,

coeruleana F

is 28 lunatene

pie

131 vitrana [32 packard [33 conversana 34 imitative

FEMALE GENITALIA OF GRAPHOLITHA

FOR EXPLANATION OF PLATE SEE PAGE 196

BULLETIN 132 PL. 23

U. S. NATIONAL MUSEUM

[37 lena

ae
/Al erotella S nee

/40 nigricana /42 Hexilogua

FEMALE GENITALIA OF GRAPHOLITHA AND LASPEYRESIA

FOR EXPLANATION OF PLATE SEE PAGE 196

BULLETIN 132 PL. 24

U. S. NATIONAL MUSEUM

1435 garacana :
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FEMALE GENITALIA OF LASPEYRESIA

FOR EXPLANATION OF PLATE SEE PAGE 197

BULLETIN 132 PL. 25

U. S. NATIONAL MUSEUM

cornutana

[EVA

CUES OIE!

SSS

americana

(SU) leucobasis

FEMALE GENITALIA OF LASPEYRESIA

FoR EXPLANATION OF PLATE SEE PAGE 197

U. S.

NATIONAL MUSEUM

1§9 toreuta

6/ ingens

l62 | injectiva

BULLETIN 132 PL. 26

163 piperana

FEMALE GENITALIA OF LASPEYRESIA AND HEDULIA

FOR EXPLANATION OF PLATE SEE PAGE I97

BULLETIN 132 PL. 27.

U. S. NATIONAL MUSEUM

165 In0pIos@

I64 obrusa

169 pomonelia

l67 colrena
FEMALE GENITALIA OF LASPEYRESIA AND CARPOCAPSA

FOR EXPLANATION OF PLATE SEE PAGE |97

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 28

IT] veru/ana

[72 pa ela

114 sinistra

IIS

malorina

FEMALE GENITALIA OF BACTRA

FOR EXPLANATION OF PLATE SEE PAGE I97

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 29

116 rhoifructana

Crt yvernoavana

/ 1S Iiriodendrana & 1

ISO aemulana 18! carduana 182 wileana

FEMALE GENITALIA OF POLYCHROSIS

FOR EXPLANATION OF PLATE SEE PAGE 193

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 30

IS4. aspasiana

(SS cyclopiana

IS6 galbinea

fo) 7 fyrius

FEMALE GENITALIA OF POLYCHROSIS, EPISIMUS, AND AHMOSIA

FOR EXPLANATION OF PLATE SEE PAGE 198

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 31

191 melanosticta

192 nubilana

FEMALE GENITALIA OF TANIVA AND ENDOTHENIA

FOR EXPLANATION OF PLATE SEE PAGE 198

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 32

udens

193 mp

ee e, IDS vulgana

196 cot ae
sordulenta 197 rubipuncta

GENITALIA OF ENDOTHENIA, HULDA, TIA, AND EUMAROZIA

FOR EXPLANATION OF PLATE SEE PAGE 198

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 33

198 andromedana

200 rosaochreana

OZ a /pproximana
201 hemidesma

GENITALIA OF ZOMARIA, EVORA, AND ESIA

FOR EXPLANATION OF PLATE SEE PAGE 198

54346—26——_17

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 34

204. nifidanum

203 zellerianum

sit lta age a 207 sciotanum 1 ey eee
208 brunneopurpuratum

206 versicolranum

FEMALE GENITALIA OF EXARTEMA

FOR EXPLANATION OF PLATE SEE PAGE 199

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 35

4 PAG
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209 foedanum 210 clavanum PA if

ohvaceanum

2/2 punctanum | 214. filanum

2/13 inornatanum

ZS) electrotuscum 2/6 subnubilum el] atrodentanum

FEMALE GENITALIA OF EXARTEMA

FOR EXPLANATION OF PLATE SEE PAGE 199

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 36

221 monetiferanum Es LESS ferrugineanum

222 merrickanum

224 hyppocastanum YS permundanum 226 quadritidum

FEMALE GENITALIA OF EXARTEMA

FOR EXPLANATION OF PLATE SEE PAGE 199

U.S. NATIONAL MUSEUM BULLETIN 132 PL. 37

exoletum 225 corylanum 229 concinnanum

252 fescatanum

ZS} niveigu/tana 233° contixana

FEMALE GENITALIA OF EXARTEMA AND PHAECASIOPHORA

FOR EXPLANATION OF PLATE SEE PAGE 199

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 38

2354. osmundana

256 griseoalbana

237 abbicihana LISS auricapitana

FEMALE GENITALIA OF OLETHREUTES

FOR EXPLANATION OF PLATE SEE PAGE 200

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 39

“2A0O consfellatanai

carolana

C42 coruscana O43

FEMALE GENITALIA OF OLETHREUTES

FOR EXPLANATION OF PLATE SEE PAGE 200

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 40

1! ly Wee

Weliae Ae

aa ae
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QAC coshimaculana’

O44 polluxana

\ly

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FEMALE GENITALIA OF OLETHREUTES

FOR EXPLANATION OF PLATE SEE PAGE 200

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 41

250 *

devolana

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FEMALE GENITALIA OF OLETHREUTES

FoR EXPLANATION OF PLATE SEE PAGE 200

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 42

254

punclicostona

258 intermistana

2S, duplex

GENITALIA OF OLETHREUTES, BADEBECIA, AND SCIAPHILA

FOR EXPLANATION OF PLATE SEE PAGE 200

U. S. NATIONAL MUSEUM BULLETIN 132

PL. 43

/ineana

260

\\
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26! GC inosema

ochroleucan@

| ae 2635 separatana

FEMALE GENITALIA OF HEDIA

FoR EXPLANATION OF PLATE SEE PAGE 201

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 44

268
dec eplana

FEMALE GENITALIA OF APHANIA

FOR EXPLANATION OF PLATE SEE PAGE 201

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 45

2U0 youngana

21zZ

albeolana

ZU ; 4 infida
FEMALE GENITALIA OF APHANIA

FOR EXPLANATION OF PLATE SEE PAGE 201

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 46

VE hea Lritarra

2/6 SIIPILIAIIA |

MALE GENITALIA OF DICHRORAMPHA

FOR EXPLANATION OF PLATE SEE PAGE 201

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 47

Ed | ICOTIQVIIA

ZOO leqopardara

wr ie ean

7 ed .
SCILAIA/IA

Zoe

i, EAS a OF

Pee ee eee

MALE GENITALIA OF DICHRORAMPHA

FOR EXPLANATION OF PLATE SEE PAGE 201

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 48

Zoo ye VERA EL SNGMIETA

LOCO DS 2Y oe

UfOrI

‘SS Qe Fr

7 SS 5 Oe
26/ fexanana LIC felicilara

CP b
LOE fourtana

22S bumelianra

MALE GENITALIA OF LASPEYRESIINAE

FOR EXPLANATION OF PLATE SEE PAGE 202

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 49

LIF = ffbertira Pe hed CACLU/EA/IIA

29 packard? BOO iS

SOA COLWELL EQIA

JOZ echipsara

LIE Um/frativa SOS LUIS AVP

MALE GENITALIA OF GRAPHOLITHA

FOR EXPLANATION OF PLATE SEE PAGE 202

54346—26 18

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 50

JOF BNGIESEA/IA 508 mole sta

SOE wrershircrana

JOB

\\ B/O suminesa
IO? duodecemstriala

MALE GENITALIA OF. GRAPHOLITHA AND OFATULENA

FOR EXPLANATION OF PLATE SEE PAGE 202

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 51

“4

VAFUTIONIR

Ulam SarlCeQ072

MALE GENITALIA OF LASPEYRESIA

FOR EXPLANATION OF PLATE SEE PAGE 202

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 52

IIE

Bere.

Saou

Bz/ LletChEY 2078

MALE GENITALIA OF LASPEYRESIA

FOR EXPLANATION OF PLATE SEE PAGE 203

U- S. NATIONAL MUSEUM BULLETIN 132 PL. 53

CV caf S701 OF OSA

He COL OI7IA

SILF JV OSPEl AIA

COPIDA/IA

MALE GENITALIA OF LASPEYRESIA

FOR EXPLANATION OF PLATE SEE PAGE 203

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 54

az a 7 OQ/7G

CPA pormorella

MALE GENITALIA OF LASPEYRESIA AND CARPOCAPSA

FoR EXPLANATION OF PLATE SEE PAGE 203

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 55

See DOMSIP AIA 7 GAM aESaANICTANa

Wye CHIR

MALE GENITALIA OF LESPEYRESIA AND HEDULIA

FoR EXPLANATION OF PLATE SEE PAGE 203

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 56

Dispos S211? 209

TIE QIPCIICAIIA

nic Wa COlOrAVIa

MALE GENITALIA OF LASPEYRESIA

FOR EXPLANATION OF PLATE SEE PAGE 203

U.S. NATIONAL MUSEUM BUECERING i323 PE or

-—

Ceres ie

S57 SSVASIADQIID

SH/ LUPCHIAIS COSI 1T

MALE GENITALIA OF LASPEYRESIA, ECDYTOLOPHA, AND GYMNANDROSOMA

FOR EXFLANATION OF PLATE SEE PAGE 203

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 58

3I4E fsariceosara oS 46 pee

IF. SUSU OE.

IFC chrysea

JSIA9 mortanana

ZFD priap C/7

MALE GENITALIA OF BACTRA AND ENDOTHENIA

FOR EXPLANATION OF PLATE SEE PAGE 204

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 59

325/ daeckeara cae

“4

JIS2 sordiu/enta ee ee bipure. oe

Ios. Lite
B53 melanosticra ETA

MALE GENITALIA OF ENDOTHENIA

FOR EXPLANATION OF PLATE SEE PAGE 204

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 60

FS5E Lorrand

—o7 : Mirtodendraria

FEF COVA UaIIA

AYN St

564 Soe |

MALE GENITALIA OF POLYCHROSIS

FOR EXPLANATION OF PLATE SEE PAGE 204

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 61
rs és
‘ Fi { Ag an
ae
x -

GOS QYUlICOIA JEG awerulanra

: i
= ila,

EE yoy

JOE bandula S72 rholl-uclora | : ; oy

MALE GENITALIA OF POLYCHROSIS

FOR EXPLANATION OF PLATE SEE PAGE 204

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 62

SIF PECELI ANIA

SIP capreana

MALE GENITALIA OF APHANIA

FOR EXPLANATION OF PLATE SEE PAGE 205

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 63

SYA VErKIA/7@

Le! OVIICKICIA

MALE GENITALIA OF APHANIA

FoR EXPLANATION OF PLATE SEE PAGE 205

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 64

Cee apare HICQVIA

Lt ida

FCIVIOLKA/7@

MALE GENITALIA OF APHANIA

FOR EXPLANATION OF PLATE SEE PAGE 205

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 65

JBL SIVIGOSA

FOS a/beolara

IEF ADOST QNIA

MALE GENITALIA OF APHANIA

FOR EXPLANATION OF PLATE SEE PAGE 205

54346—26- 19

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 66

JO8 WIP U AES S

SOD: QSPAE/A172

Gab inea &§ if i

~~ fy?
390 fottlesciana

JOA ou yp Pax IW uwrtcand

MALE GENITALIA OF OLETHREUTINAE

FOR EXPLANATION OF PLATE SEE PAGE 205

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 67

IIF Pon ange

ae WGUIANUS

a foro) harvigiarra

wablbergiana

MALE GENITALIA OF EPISIMUS, PHAECASIOPHORA, CYMOLOMIA, AND ECCOPSIS

FOR EXPLANATION OF PLATE SEE PAGE 205

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 68

AE eI a

AE. SUVSAAIT ASIA

4OO olweceanum
FU/ PMIICT AIIM

MALE GENITALIA OF EXARTEMA

FOR EXPLANATION OF PLATE SEE PAGE 206

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 69

ar -?

QQCEIIDICEU 17 FDS HOGION AIT?

: ) MARE
ie r GOS ferriferariutn

FOP fasclatanum

MALE GENITALIA OF EXARTEMA

FOR EXPLANATION OF PLATE SEE PAGE 206

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 70

SAN

FOE ITEP UPIOMICRLIA

FOP posaocheana

F/O anad-omedevna 4/4 GOH OKT UIA

yrs; : aio} CVA aa
4) — A CTUACE/TIA

GENITALIA OF ZOMARIA, EVORA, TIA, EUMAROZIA, ESIA, AND HEDIA

FOR EXPLANATION OF PLATE SEE PAGE 206

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 7I

4/ 5 F/P cee

F//7 =» eae F770 wong og

FO/ costimaculana

WE = variegana

MALE GENITALIA OF HEDIA AND OLETHREUTES

FOR EXPLANATION OF PLATE SEE PAGE 206

U. S. NATIONAL MUSEUM BUELERING 132) SPI 72

422 griseoa/bara

G23 osr7urdaria ee

FF aaurreagpirara B27

chalybearia

MALE GENITALIA OF OLETHREUTES

FOR EXPLANATION OF PLATE SEE PAGE 206

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 73

926 sordidarna F5/ pee

429 constellanriara (EI Gaclana

| Cte

MALE GENITALIA OF OLETHREUTES

FoR EXPLANATION OF PLATE SEE PAGE 207

U. S. NATIONAL MUSEUM BULLETIN 132

PER 7G!

FTF CESPIT AIA bP GT LEO ihe

= 25 ee
eho Liparvitara CSU ALLE,

436 LEED!

Se ee eee
MALE GENITALIA OF OLETHREUTES

FOR EXPLANATION OF PLATE SEE PAGE 207

U.S. NATIONAL MUSEUM BULLETIN [32 PL. 75

FF0 SCAULZ/A/92 445 SEP PES EIONI OIA

a7 MPP CLTTUSEV ASIA 444 pp asor

44S POMUXQMA

aera jngilletana

MALE GENITALIA OF OLETHREUTES

FOR EXPLANATION OF PLATE SEE PAGE 207

U. S. NATIONAL MUSEUM BULLETIN 132 PL. 76

P96

FOGLE Lowrr7as/7 17D”

FIG Luckelarna

MALE GENITALIA OF OLETHREUTES

FOR EXPLANATION OF PLATE SEE PAGE 207

INDEX

This index includes all the genera, species, and varieties treated in this paper and also groups discussed

in the text when the information pertaining to them is important.

Family and subfamily names are in

small capitals, valid generic names in boldface, valid species and varieties in roman, and synonyms in

italics.

Page
RLUICIC IB TISC Ke ec oa oe kok 171
auido chernald)==2.----- ~~. --- 22-222 - 107
MaMilaneMeMnniCh =... 22. - o222--225--5 05s 89, 94
MCQUONENMOVLOVTICK) 222 os nes oo oo 16
BC Clee mmens — 2-22 5-5. 2-5 115, 118
PAN CHEIMENS) eee ae 168, 171
Pnirmrosiaeinnmien a a 76. 97
aspasiana (McDunnough). 98, 99
galbinea Heinrich- _-_.___- 97, 98
PMO aN ey (VAG Oppel aie ee 114, 119
PUL STU UES revit Gl O(Z\ era FC De aps see 168, 171

albidula Heinrich (variety of buckellana
(he Munnowgh)) eee ee 167, 189
‘albimaculana: (Fernald) 2 2.-2. 2.20.2... 43, 51

albipuncta Heinrich (variety of verutana
LONGED 1a ets ep eat SS a a 82, 84
WLUO;OSCIIVILIN LONE aon nee oo otk 159
albolineana @iearfott)o---- 22... 22) -. 2 5, 106, 107
TA TREE (CLOTS enka ae ae ie a en ee 79
PAT (eNNaAld) =e ee 12
eaimprostanaKearlottess=- 22. <2 - asa econ eee 94
americana (Walsingham) _____.......-.._---- 43, 60
amygdalana (Duponchel)--_-_..-..._____--.-- 21
andromedana (Barnes & McDunnough)._- 112,113
anglescanai(Keartott)e-.- 22 -- =. eae 27, 31
antiquanal(Hiibner)!) == 2282 2228 100
apateticana (McDunnough)..._.__...-_-_- 114, 120
PA hanin veld bner ses en 76, 113
afficticia Heinrich____._- 115, 118
albeolana (Zeller) __....- 114, 119

apateticana (McDun-
NOW) ea eM 114, 120
capreana (Hiibner)-_--.__- 115
deceptana (Kearfott).... 115, 120
deceptana (McDunnough). 120

dextrana (McDun-
MOUPH) poe saa ae 115, 121
frigidana (Packard) ____- 115, 116
Junerea (Meyrick). ------- 115
infida Heinrich----_- 114, 115, 121
inundana (Schiffermiiller) 120
moeschleri (Kennel) .___--- 116
removana (Kearfott).... 114, 122
scriptana (Hiibner) ----- 113, 119
semifasciana (Haworth)... 113
sororculana (Zetterstedt).. 119
strigosa Heinrich______-- 115, 118
tertiana (McDunnough) 115, 117

youngana (McDun-
Lala be 5) Suge ei ee 115, 116
appendiceum Zeller-__..............-..-... 135, 157
approximana (Heinrich)__.._.........----- 109, 110
arbutelia (uinmsdens) i soeee 228 2s 166

54346—26——20

Page

areuellar(Clerck))s-c see = 2 ose ee 130, 166, 167
angutanus (Clemens) 22.2 sa eee aes 79
Ang y Tana CENT) seco cee eee eee 20, 23
argyrociana (Zeller) i 22.22 eee ee 175
PAN TUTOTIOCE TLL Dela = - ne ee ee 166
ariemesiane (Zeller). 88
ariculatana (Kearfott) _..--___-- = 5r
ARUNCAND ACCAtlO lta 8 oan ee ee ee 89, 95>
aspasiana (McDunnough)_-_-_-____.-_-___--__ 98, 997
astrolozanal(Zellen) wees ee ee 168, 174
atrodentanum (Fernald) _....-...-._.______ 132, 141
augmentanus) (Zeller) -2— ea 79, 80
auricapitana (Walsingham)__________._____ 168, 170
MiuyichaiccangeRileysa= ae ee ee 67
UTERUS TLIO ZO eae en eee ee eee 12
BactrayStephens aan see a lo eee 76, 82
furfurana (Haworth) -_____ 82, 83

lanceolana (Hiibner)_-__ 81, 82,83

maiorina Heinrich____.___- 82, 86
priapeia Heinrich_____- 82, 83, 86
Scirpamna (Herrich-
Schacter) 2225s aa ees 86
sinistra Heinrich__._...___ 82, 87
verutana Zeller_...___..___ 83, 84
verutana albipuncta Hein-
Tienes. 2 ee eee 82, 84
verutana chrysea Hein-
Tigh s222 2 ete = ee 83, 85
BadebeciaytHeinrich=s= 2222s eee 76, 124
campestrana (Zeller)... 125

urticana (Hiibner)_.. 124, 125

Bal piss Walsingham. cents eee 3, 19
bananas (BUSCK) S222 ee ot ee ee ee 11,14
bentleyans (Curtis) asses ste ee ee eee 183
bicoloranum MeDunnough_-___________--- 131, 144
bipartitanan(Clemens) tess) 22 seen eee 167, 180°
bittanaCBusek) S= ssa ae ase hn ee eee eee 10, 13

Dlandula*Heinriche ee "=a s2s-* ae ene 89, 96°

bolandanum McDunnough. ____-_--_-------_ 137
DONA Re Delns a 6acee ee ee eee 186)
botranayhernal dae: soit ee eek eee 90°
botrana (Schiffermiiller)_..._.......--..-__-- 87, 88:
bowmanana (McDunnough)-_----_--------- 169, 186:
Brachytaenia Stephens_--.--.-.----------.--- 113:
Ipracteatanai(heriald) sess een e aes aa eee 44-
branderiana (Linnaeus) --......---------.---- 123:
brevirostratum Heinrich._...........---.-. 133, 154!
DYitanas (BUSCK ea eee ne ae ee 11, 12:
brunneopurpuratum Heinrich-----_------- 133, 151’
buckellana (MeDunnough).--_---------- 117, 169, 188:
bumeltana Heinnich= sees eee eeee ae 8:
caeruleana Walsingham. ...-..---.---..----- 27,31
caesialbana:(Zeller)2222--ccceccssoau eee a Le

209

910 INDEX

Page Page
campesinana: (Zeller) sas = sees eee 125 | Dichrorampha Guenée banana (Busck)---_--- 11, 14
candanalHonbes: a3 ese = see ee ee De 44, 54 bittana (Busck) - ----- 10, 13
Capltanal (BUS) poe ss see ae ee 10, 11 britana (Busck) -.---- 11,12
capreana: (EtG bner) 222 eee 115 capitana (Busck) ----- 10, 11
CarduanseBuscks.sees = ae eee 89, 96 dana (Kearfott) __-..- 10, 16
carolana (McDunnough).--_--------------- 167, 180 incanana (Clemens)-- 10, 13
Carpocapsa /Trietschkes 2 2 oe eee 6, 69 kana (Busck)--..------ 10, 11

pomonana Treitschke 70
pomonella (Linnaeus) 5;

69, 70
pomonella simpsoni
((Busck) Pe seaseaeen 70, 71
CORY CEL SALINE) se eres pee nee eee ee ae 55
caryanal (ite) <2 ==) = 2224-52555. - keene 44, 55
castorana (McDunnough)_.-_..-.---.-------- 182
Gel DOME ONel eee san aa ene 166
GeralasStepneusse. sss5) 2 -— ==. = scone eee 41
cespitana(MUnnen) a5) oe 168, 179
chalybeana (Walsingham) (variety of side-

Hania pure SCHKe). os s0 anon a ae 168, 172
wbionosema (Zeller) <— 2-2 s-=_ ee ees 162, 164
christophana (Moschler)_.-.......-----.----- 21
chrysea Heinrich (variety of verutana Zeller) 83, 85
chrysotypa (Meyrick)= =. ----- -=-4=-a0--soao= 37
CUTROS) GVICY TICK) ==" == ees 2 eee eee 56
clavanum) (Walken) 2-2 2-2 = 2 55. sea 131, 143
coloranay (Kearlott) p= ss -22=— =a en ee nee eee 43, 61
compositella (Fabricius) -...-......--.------- 26
conchanan(Hubnen)=ssess-222--2- seen eee ee 166
concinnanum (Clemens) -_-------.---------- 132, 158
conditana (Walsingham) -..._...-..-----.. 101, 104
confixana;: (Walker) (22-4255 25s) oo eee 126, 127
consanguinana (Walsingham)-_.-.--._-------- 163
constellatana, (Zeller) 22-2225 <=) -a2 = eee 168, 174
COTO LOTS AVY AUK GT) ee ee ee 163
conhversana Walsingham: -- 2022-7 222 27, 33
COMAaN UME Clint Cheass ee ee eee 132, 142
cornutana (Dyar) (variety of bracteatana

(Hernia) ese See nek ene as 44, 45
eorollanas(ELubneY) ease eee aoe enon eeeeee= 41
COLORANG Car lottae se ee eae eae 174
coruscana:(Glemens) eee ee eee 168, 175
Corylanum)((Hermald)je= 222522222 sone e— 132, 150
costimaculana (Fernald) _.......-.......... 167, 187
crispana (Clemens) (Thiodia) -_.....--...--- 48
CuUpida (Mey TICK) Sn ee ae eee ee cere 33
Cuwpressana CKcarott) ass sone oe eee 42, 56
evyanana:(Murtfeldt) 22-22. 22-- eee 162, 165
eyclopianay Heinrichs: sess) eee nee 88, 97
Gymolomia Busch. sso ee enn essence aes 129

hartigiana (Ratzeburg)... 129

eypripediana Morbes:- = -— = =) ae 89, 92
daeckeanay(Kearfott) a= 3-2 eee 101, 104
dalecarhnanay(Gulenee) ea. a a eee 166
danai(Kearfott) 2.22.2 2 ease eee 10, 16
@andand: (Kari Oth) oes eee 53
aealband\(Wialkker)) 2-3 eer ek dai lor
deceptana (Kearfott) ......-...-.-.5....... 115, 120
deceptana (McDunnough) -_-.-.-_---------_.- 120
mecisana: (Walker) == 3) a Sate a ee 159
Aeprecatoria Heinrichis= seo caesar 168, 177
AesovaniwwmeH einricha 42 a2 - se ase ee 72
Mevotana: Kearfott.-22- 22522 oe ee ee 167, 188
dextrana (McDunnough)__-._...._-_..___.. 115, 121
Dichrorampha Guenée_______..___-_---__---- 6,9
aequorea (Meyrick) --_- 16

alpinana (Fernald) - -- 12

aurisignana Zeller. --- 12

leopardana (Busck) -- 10, 16
nigromaculana (Kear-

fObb) he eee eae 13
petiverella (Linnaeus) 9
piperana (Busck)_..--. 11,15
planiloqua (Meyrick) - 11
plumbagana_ (Treit-

schke) 20285 sete 9

plumbana (Fernald) -. 15

plumbana (Scopoli)._ 9,15
radicicolana Walsing-

AMS = ae 11, 14
radicolana (Fernald) -- 14
sedatana (Busck)_---- 10, 15

simulana (Clemens)... 11, 12
sordescens (Meyrick) - 14

dilutijuscana Kearlott.--s=-s->--— ee 181
dilutifuscana (Walsingham) -.....--------- 167, 181
adunidiang Chermald) esos eee eee 162
distema, Grote: -cs-s52 554-52 =o a eee eee 36
dorsana (abricius) eats 22s se a eee eee 26
duodecemstriata (Walsingham)-__.-_...._-.-- 39, 40
duplex (Walsingham) 222-2 so5- =a 124
dyarana (searfott)is=ee- 282-2 - ose ee 28, 38.
Eccopsis Zeller ss.ee== = ee ee eee 129
wahlbergiana Zeller___...---- 129

Ecdytolopha) Zellemes ot eeen eee terse ee 3, 4, 6, 73
insiticiana Zeller___._- 5, 73, 74

insulicola (Meyrick) ---. 75

islandana (Kearfott).... 74, 75

mana (Kearfott).......- 74, 75

thaliastis (Meyrick)-..-- 75

eclipsana) Zeller: <= 252- =< occa ee sac eee ee 27, 36
edwardsiana:(Kiearfott)=22.2222-55--5-- eee 28, 37
electrofuscum Heinrich...............-.... 133, 139
Endopsia\GQuenee. 22s. -- 255-25. se--- =~ 2 =e 41
Endothenia Stephens_-_.___..._.......-- 3, 4, 76, 100

antiquana (Hiibner)_. 100
antiquana nubilana

(Clemens) - ---.-- 101, 105
conditana (Walsing-

ham). S222s=22225 101, 104
daeckeana (Kear-

fobt) a 24-=2 101, 104
flavillana (Dyar)----- 103
fullerea (Riley)-.-.--. 103

gentianana (Hiibner) 100
hebesana (Walker). 101, 103
inezpertana (Walker) 103
infuscata Heinrich. 101, 105

melanosticta (Wal-

singham) ...-.... 101, 103
montanana (Kear-

1Otb) Maes See 101
rubipunctana (Kear-

fott) j2aniss.22u 101, 102
sordulenta Hein-

Tich: anaes 101, 102
vetulana (Walsing-

Nam) soa sees 105

eoleuca! (Meyrick) 2-222ee2 soe eee eee eee 31

INDEX OTP
Page Page
Ephippiphora Duponchel_------------------- 26 | Exartema Clemens inornatanum Clem-
Episimus Walsingham-_-__--.------------- 3, 4, 76, 78 ens. 2 sS225 See 132, 142
allutana (Zeller)_----- 79 malanum (Fernald)__ 131, 157
argutanus (Clemens)- 79 meanderana (Walker)... 155
augmentanus (Zel- mediopartitum MHein-
HOR) Petes e a ae 79, 80 Bichwwres ec eek hams 132, 143
hamameliella (Clem- melanomesum Hein-
(=S01s) eee ee ere 79 Fich 3-22 eee 132, 153
nesiotes Walsingham_ 78 merrickanum Kear-
transferrana (Walker) 78 fott2s4absSe Fas 132, 149
tyrius Heinrich---_-- 79, 81 micantana (Fobes) ----- 153
erotellarébempieh hese ee. 2. ob 2-22 43, 61 monetiferanum Riley._ 129,
EsiamEeinrich 2 eee ne sale 3, 4, 76, 109 131, 135
approximana (Heinrich)_--. 109, 110 myricana (Kearfott).... 152
inelgoda; heinrich 2822-5. 3 Les Ses es 6, 23 nananum McDun-
texanana (Walsing- nough=<=22===ee=- 133, 156
: 131345 6) etre ee 23, 24 nigranum Heinrich_-.. 131,
IPUCOSMIN AT 2 oak SSA ey 4 132, 148
Kumaroviavkeinrichts)-s202. 2-22 2222..222.. 76, 110 nitidanum Clemens.-.. 128»
malachitana (Zeller)... 110, 111 132, 136
Guphorbianasbreyers.222----ss2hee- 98 nitidanum Zeller ------- 139
PspileStephens t= - eas eee 26 nortana (Kearfott) - -_.- 153
evcrae Heinrich ss eee 3, 76, 189 ochrosuffusanum Hein-
hemidesma (Zeller)__..--- 189, 190 jake eee 132, 156
exaeresimum Heinrich___-.-..-.------------- 160 olivaceanum (Fer-
Exartema Clemens-_--_-..------------------ 78, 128 eG) 133, 137
albofaciatum Zeller...-- 159 ornatana (Kearfott).... 148
appendiceum Zeller__ 135, 157 permundanum Clem-
atrodentanum (Fer- emS---------------- 183, 155
maid) eee 132, 141 punctanum Walsing-
bicoloranum McDun- GE 132, 141
MOUs he Nee sass 131, 144 quadrifidum Zeller__- 132, 145
bolandanum McDun- quebecense Heinrich_---- 156
moughesgs siete 137 trusticanum McDun-
brevirostratum Hein- nough----.-------- 133, 139
rich: 2 neo 133, 154 sciotanum Heinrich_- 132, 147
brunneopurpuratum sericoranum Walsing-
Heinrich_.--_------ 133, 151 an 133, 135, 152
clavanum (Walker)__ 131, 143 submissanum McDun-
concinnanum (Clem- nough----_-------- 131, 156
nS) ees ate aa 132, 158 subnubilum Heinrich 134, 138

2oncinnanum termina-
num McDunnough 131, 158

cornanum Heinrich-- 132, 142

corylanum (Fernald) 132, 150

decisana (Walker) -.---- 159
electrofuscum Hein-

mi Cheyne. 29-22 2522 525 133, 139
exaeresimum Hein-

riehs 2 2242s s222-- 133, 160
exoletum Zeller------ 132, 144
fagigemmeanum Cham-

Wersehsoos22=e-o se. 133, 151

fasciatanum Clemens 135, 159
ferriferanum (Walker) 131, 161
ferrugineanum Riley- 132, 151
foedanum (Clemens). 133, 136
footianum (Fernald)_ 132, 140
fraternanum McDun-

moOughe-==----=s-—— 135, 138
furfuranum McDun-

nOuUg Mes ees ass==—= 133, 137
gaylussaciana (Kearfott) 155
gratiosana (Clemens)... 161
hippocastanum Kear-

fot) Se ~--.-=-- 133, 149

tenebricum Heinrich. 131, 144
tilianum Heinrich_-- 1382, 146

trepidulum Heinrich_-_ 182,
140, 147

troglodanum McDun-
Rouen 225 ee 133, 159
usticana (Zeller) -._---- 161

valdanum MecDun-
novgh== 2) --cs=—5—4 135, 153

versicoloranum Clem-
ONS Senso eee asa 135, 154
zellerianum (Fernald). 133,
139
exoletumeczellors Soy iiwik. feo eee se eee eae 132, 144
fagigemmeanum Chambers-.--------------- 133, 151
fan Ckcariott)easeeeens oe - 27, 33
fasciatanum Clemens- --------------------- 135, 159
felicitana/ (Heinrich) S22225--22---=---------—— 21, 22
ferriferanum (Walker). -..----------------- 131, 161
ferrolineana ( Walker) ------------------------ 175
ferrugineanum Riley_--------------------- 132, 151
fia briana (Haw Olu) jesse e === =a eee 21
flavicollis (Walsingham) -.-----.------------- 42, 60
flaviiiana) OD yan) essen =a ae een 103
flavofasciana (West wood and Humphreys) -- 179

212 | INDEX
Page Page
ffletcherana (Kearfott).___.------------------ 42,56 | Grapholitha Treitschke sawndersana (Kear-
Afiexanay (Zeller) 22.5 sss ees oS a es 20, 21 fot) Bi zzctn se ee 39
Aexiloguadaeimnich=ssseeeeen ese 42, 59 scitana (Walker) _.- 36
foedantmy(Clemens) =e sesen eS =e 133, 136 tristrigana
footianuml Ghemald)Ss2ss2225_----=-----=- 132, 140 (Clemens) -_.....- 27,39
fraternanum McDunnough---.-_------------ 135, 138 vana (Kearfott) __-- 31
frigidana,@hackard) 2222s see aaa === 115, 116 vancouverana
fullencai(Riley))o2 == 2 = ee ee eee 103 (Kearfott)=.--=-_ 37
Julvifrontana (Packard)---------------------- 185 vitrana (Walsing-
fiunebrana.(ireitsehke))-2225—----=---2--->--— 26 Man) See 27, 32
iyumerea'\(Meyrick)2 22-2 s-s==- 2 === = 115 wana (Kearfott) --- 33
furturana) (ela worth) eee ae enone 82, 83 zanthospora (Mey-
furfuranum McDunnough----------------- 133, 137 rick) =<: ees 31
apscapanm(aeller)=ssessaseee=———- ee == 182 zana (Kearfott)--_- 31
galaxanaiearfott=2--ss"=5=----=—---- === 168,173 | gratiosana (Clemens) ___.---...-----------=-- 161
galbineasbleinriche <2 ees. == - 22a eo 97,98 | griseoalbana (Walsingham) ---------------- 167, 169
gallaesaliciana (Riley) ------_:--------------- 42,58 | groenlandicana (Bang-Hass)____-------------- 187
gallicolana(Zeller)=--2- 25-2 25 <== = == 21 | Gymnandrosoma Dyar-_.-----..---------- 3, 4, 6, 71
garacana, (dearfott)s2= 22522 2---- === -===-==— 43, 49 desotanum Heinrich. 72
gaylussaciana (Kearfott) --.------------------ 155 punctidiscanum Dyar 71,72
gerntianans, (Eiibnene==- =. = = 100 | hamameliella (Clemens) _.__------------------ 79
giaeiana: (Moshler) ss sses8 See: -=---+-- ==. 167,182 | hartigiana (Ratzeburg) ---........---.------- 129
glitrananaikKearfottie == se s2Ss_ ea aa one 173: |; hebesana: (Walker) >. = 25-2... =.=. ele 101, 103
GodithatHeimniche > oe) eee 638i") Hreidiiian ii bnenese sacs ) ee 78, 161
bumeliana Heinrich_--.---- 8 chionosema (Zeller) __...-- 162, 164
PoranayKearlotts=s =a. sa ae enna 191 consanguinana (Walsingham) 163
erandiculaseinniches=-—s=2=---—-----=2=a== 44, 54 contrariana (Walker) -.------ 163
Grapholitha Treitschke2=2.._-.--.--..-.-- 6, 26 eyanana (Murtfeldt)_-.-_- 162, 165
angleseana (Kear- dimidiana (Fernald) _-------- 162
1Obt) se 27, 31 lineana (Fernald) _-_-._---- 162, 165
eaeruleana W al- nimbatana (Clemens) -_------- 163
singham-.-------- 27, 31 ochroleucana (Hiibner)--.- 162, 163
chrysotypa (Mey- salicella (Linnaeus) - -------- 161
rick)/=—2=-------= 37 separatana (Kearfott) .------ 162
compositella (Fa- variegana (Hiibner) ------- 162, 164
jbrictus)=2==222--- 26h |b Heanliawlcinrich= === sere 6, 65
conversana Wal- injectiva Heinrich-------- 65
singham.-.---.-- 27,33 | Hemerosia Stephens .-----.---------------- 20
cupida (Meyrick)-. 33 | hemidesma (Zeller)_.---------------------- 189, 190
distema (Grote)-.-- 36 | Hemimene Fernald and Authors (not
dorsana (Fabricius) 26 Ju bner) ose te See ee be ee 9
dyarana (Kearfott) 28,38 | Hemimene Hiibner_--_------------------- 3, 6, 20
eclipsana (Zeller) -_ 27, 36 amygdalana (Dupon-
edwardsiana (Kear- chel)eee ee eee 21
f0tt)Es~a2e a= ae 28, 37 argyrana (Hiibner) ----- 20, 21
eoleuca (Meyrick) - 31 christophana (Mésch-
fana (Kearfott)_... 27,33 lenjeie ee So oe sees 21
funebrana (Treit- felicitana (Heinrich)__-- 21, 22
senke)2=22=---— = 26 fimbriana (Haworth) _-- 21
imitativa Heinrich 27, 34 flexana (Zeller) - -.------ 20, 21
interstinctana gallicolana (Zeller)_-._-- 21
(Clemens) ------- 28, 36 juliana (Curtis) -------- 21
jungiella (Clerck) _- 26 ocliferia Heinrich ------ 21
lana (Kearfott)- 27, 28, 37 oxycedrana (Milliére) -- 21
libertina Heinrich. 27, 29 paula Heinrich---_---.--- 21, 23
lunatana (Walsing- populana (Fabricius) --- 20, 21
ham) -.--S2-S2ee. 27, 35 regiana (Zeller) _-------- 21
molesta (Busck) - 5, 27, 28 rhediella (Clerck)------- 20, 21
oenochroa (Mey- signifera Heinrich----.-- 21, 22
Tick) ieees aeeu 33 spiniana (Duponchel) - - 21
packardi (Zeller). 5, 27, 29 splendidulana (Guenée) - 21
placerana (Kear- trauniana (Schiffermiil-
fott) eee te ae 37 er) heeites here 20, 21
prunivora hippocastanum Kearfott------------------ 133, 149
@Wialsh) 22222 52730) |; HuldavcHelnricheseseswess == oases 76, 108
pyricolana (Murt- impudens (Walsingham)- 108, 109
feldt)=s-." 2222 29) |-imitativaibeinni ches ]-==sessse——= == eee 27, 34

INDEX 213
Page Page
impudens (Walsingham) - --.-------------- 108,109 | Laspeyresia Hiibner multilineana (Kearfott) 43, 50
incanana (Clemens) - ------------------------ 10, 13 nigricana (Stephens) _____ 5,
inexpertana (Walker) .----------------------- 103 41, 44, 53
Hretile Gy 18@sbor la oe So ee 114, 115, 121 ninana (Dyar)__-__-_- 42, 60
infuscate (eleinrich)==2-2--22---2--—---—- 101, 105 novimundi Heinrich - -- 53
roeyeqsyns) 18 Crh rat Aleta = 43, 63 obnisa Heinrich_______ 43, 47
progeny 18 G)bebstelol. 5 Se ee 43, 50 pallidibasalis Heinrich. 44
imjectiverHloinnich sen o ana asssaaene asec 65 parmatana (Clemens)_ 43, 47
vba ayo kaistsy 18 Revb rate blo ee 44, 46 piperana (Kearfott) _ 5, 43, 63
inornatanum Clemens..------------------- 132, 142 populana Busck_--____ 42, 52
inquadrana (Walsingham) -_.....------------ 191 prosperana (Kearfott). 42, 57
inquietana (Walker) ---------------------- 169, 186 rana Forbes-_--------- 44, 46
inquilina) (Xearfott)= == 222 -----.----------=- 67 ratifera (Meyrick) ____- 53
iusiticianaielers==-ss2"-—---=—-----—-- === 5, 73, 74 septicola (Meyrick) -_-_- 49
instrutana (Clemens) - ----------------------- 179 servillana (Duponchel) ® 41
ANS WLUCOLOKONLCY RICK) a eea=-—=2---5---->--==5- 75 succedana (W alsing-
intermistana (Clemens) - ---.--.----------- 169, 184 ham) 2232 Geen 57
interruptolineana (Fernald) --------------- 111,112 tana (Kearfott)_______ 42, 56
interstinctana (Clemens) ---.---------------- 28, 36 toreuta (Grote) ------- 43, 62
inundana (Schiffermiiller) --.---..----------- 120 youngana (Kearfott)__ 44, 52
islandana @kearfott)ieo. 32622528 es T4760 |p ASPRMEESHNAM 5° Je SseOl 2 See alse eee 5,6
fuilianai(@netis)Heses- sean ae eae 21 | latifasciana (Haworth) ___..........-.-.----- 129
jungiellal(Clerck)S=ss"s--2e2-ss2=—-=----55< 26 | latiferreanus (Walsingham) ------_.------- 5, 66, 67
enti (BeISGka eee eo ee Se See SOS 10510) |elautans| (Clemens) -=-2=-"- = eee 25
kreithneriana (Hornig) -.---.---------------- SRiailantivscnia Heimrich= 2222 ee ae 42, 58
lacunansa Duponchel=2ses2-- 22-5. -----=-_=- O8mlsleopardana (Buseck) => 2 eee aeeee eee eee 10, 16
fanankcarfott) = sees ene= aa sa-ee a aaae eS DTe28e3it | lel CObASIS BUSCK =a. 2- = asaaoae eee aca nen ee 42, 57
Jianceolana (Hiibner)==---2--------------= 81) 82.83|olibertina, Heinrich: -22-- 2222. oa asenenan eee 27, 29
larrcanarybuscke ss se eas sees 43-454 lneans(Hernald)s222--=—-- == sos e ee ea 162, 165
larimana (Walsingham). -_--.---------------- 43°48)\\ Lipoptycha luederers2s=--242---_----= === == 9, 10
Taspeyresia Hiibner_----=- -. --.=------.- 6; 41) |) lriodendrana Kearfott2s:<-.=---_----------- 89
albimaculana (Fer- obesia.Guentes*s 222 2a2022 2. 22h oe 88
nald)!.-5-2r a= Sees 43, 51 artemesiana (Zeller) -------- 88
americana (W alsing- kreithneriana (Hornig) - ---- 88
Ham) eos eee 43, 60 permixtana (Hiibner)------- 88
articulatana (Kearfott)- 51 Loxoterma) Buseck ase Jae ek ee 129
bracteatana (Fernald) - 44 latifasciana (Haworth)-- 129
bracteatana cornutana luminosa\Heinrichv22322822 = 2. 22-5 == 40
(Divan) eae ee 44°45 | lunatana Woalsingham=-2=---.----_------=-= 27, 35
candana Forbes_-_---.- 44°54 | maculana (Fernald) ---.----=------_-----=-—— 18
caryae (Shimer) - ------ 5b) |) “magnolana Kearlottesseee2=- === aaa 89
earyana (Fitch) ------. 44" boys meaiornna Hein chas=see=seee= = =o eee 82, 86
cirrhas (Meyrick) - - --- 56 | major (Walsingham) (variety of puncti-
colorana (Kearfott)-..- 43, 61 costana, (Wialker))22s-2- + 2o2a225 222 e 168, 177
corollana (Hiibner) --- Ale iemalachitana) (Zeller) ss 2s o2 52 = 2 a 110, 111
cupressana (Kearfott). 42,56 | malanum (Fernald) ----------------------- 131, 157
dandana (Kearfott) -._- Sou Mani cantott) sss eee eee ena eee 74, 75
erotella (Heinrich) .-.. 43,61 | meanderana (Walker)------------------------ 155
flavicollis (Walsing- mediopartitum Heinrich----._.------------ 132, 143
ham) eae ee ae 42,60 | melanomesum Heinrich-----.-------------- 132, 153
fletcherana (Kearfott). 42,56 | melanosticta (Walsingham)----....-------- 101, 103
flexiloqua Heinrich... 42,59 | Melissopus Riley -.----..-------------------- 6, 66
gallaesaliciana (Riley). 42, 58 aurichalceana Riley------ 67
garacana (Kearfott)__. 43, 49 inquilina (Kearfott) -.--- 67
grandicula Heinrich... 44, 54 latiferreanus (Walsing-
ingens Heinrich-___--- 43, 63 Nam) =2c-sess=s ones 5, 66, 67
a ingrata Heinrich __--_- 43,50 | membrosa Heinrich------.------------------ 43, 49
inopiosa Heinrich - ---- 44,46 | mengelana (Fernald) ---------------------- 169, 187
laricana Busck-_-_----- 43,45 | merrickanum Kearfott.---.--..------------ 132, 149
larimana (Walsing- mi canai(eh ben) pssst eee ae aaa a eee! 166
Nam) {eS see 43,48 | micantana (Forbes)-.-.----------------------- 153
lautiuscula Heinrich.. 42,58 | miscitata Heinrich-.....--.------------------ 43, 64
leucobasis Busck_----- 42.57 | Mizodia Guenée-..----......---.------------ 166
membrosa Heinrich... 43,49 | moeschleri (Kennel)--..--.------------------- 116
miscitata Heinrich.... 43,64 | molesta (Busck).....---------------------- 5, 27, 28

9 1 4 TNDEX
Page Page
monetiferanum Riley..........-------_- 129,131,135 | Olethreutes Hiibner costimaculana (Fer-
monotropanavelemmrich=s-s===-—-=-2--=- 22 -= 89, 91 nagld),2S2s22 se & 167, 187
montanana (Kearfott)es=2-=-2-.922-= 2-2 = 101 dalecarliana (Guenée) 166
minltilineana (earfott) ses - 2-222 el 43, 50 dealbana (Walker)_-.. 182
MVULTITULS (ER ACKATG)) eeseaes tee we ee 176 deprecatoria Hein-
mutabilana (Clemens) - -------------------- 126, 127 Fiche = y2 aac 168, 177
myricanardeantott) josie 2 = 2s 152 devotana Kearfott_ 167, 188
nananum McDunnough..--.---.---------- 133, 156 dilutifuscana Kear-
MesiotesaWalsinghameeeser es) 23s he 78 fottz= = eee say 181
NigranumsHeimnich sess =e 131, 132, 148 dilutifuseana (W al-
nigricanay(Stephens) ssssss2e2 ----+-----= 5, 41, 44, 53 Singham) S225" 167, 181
nigromaculana (Kearfott)....---------------- 13 Jerrolineana (Walker). 175
mimbatand) (Clemens) -sesese2<20-2-022---=--— 163 flavofasciana (West-
Minanay CD yan) masts ewes ss 5 so ase 42, 60 wood and Humph-
nitidanum Clemens----.-------------- 128, 132, 136 TOYS) Leese Ss; Bee 179
mitidanim Zellers) ween ee SoS toe 139 fulvifrontana (Pack-
Nivelguttana Grote=s-eess. 2 eo ee ao 126, 127 ard) mse eh aie 185
nordeggana (McDunnough) --------------- 169, 183 fuscalbana (Zeller)... 182
mortama(Kearfotth)eee 2 ease - == 22 oe 153 galaxana Kearfott._ 168, 173
Mound eleinniche seen tess = - oo ee oe 53 glaciana (Moéschler) 167, 182
nubilana (Clemens) (variety of antiquana glitranana Kearfott_.. 173
GH pre) = 2 eae eee ee aie 101, 105 griseoalbana (Wal-
Obmisaybleinrich = 2-2 sseee ee as See 43, 47 singham)/2222_-== 167, 169
ochrolewcana, (Hailbner)=22-2----- =2222 233 162, 163 groenlandicana (Bang-
Ochromedianag Kearlotts 2222 sae nee ee ee 170 ifiass)saes siete oes 187
ochrosuffusanum Heinrich_-_-.-------.---- 132, 150 inquietana (Walk-
ocliferia Heinrich--_--- Fete oh Whale. Ais 21 er) dbese Ue 169, 186
oenochroa (Meyrick) ------------ ees Loe 33 instrutana (Clemens). 179
Ofatulenal Heinrich==-- 22222. 2 a eee ee 6, 39 intermistana (Clem-
duodecemstriata (Wal- nS) ee wy 2S 169, 184
singham) pees 39, 40 mengelana (Fer-
luminosa Heinrich. -_- 40 mald)22es eh 169, 187
Olethreutes Hiibner= 422-8 Ss Ss - 78, 166 micana (Hiibner)_--. 166
abiciliana Busck -- --- 171 murina (Packard)... 176
agilana (Clemens) - 168, 171 nordeggana (McDun-
albiciliana (Fer- mough)=eeee ase 169, 183
Mald) eeeae s 168, 171 ochromediana Kear-
arbutella (Linnaeus). 166 fOthe t-te ne 170
arcuella (Clerck) ----- 130, osmundana (Fer-
166, 167 mald) = sees 168, 170
argyroelana (Zeller)-- 175 poana (Zeller) _.------ 179
astrologana (Zeller) 168, 174 polluxana (McDun-
auricapitana (Wal- mOuUgh) seesaw a= 167, 180
Ssingham) 2222252 168, 170 primariana (Walker). 185
bentleyana (Curtis)_-. 183 puncticostana (Wal-
bipartitana (Clem- ep) naa sree 168, 176
@nS) 22-2. GU ee 167, 180 puncticostana major
boreana Rebel-_-_------ 186 (Walsingham) --- 168, 177
bowmanana (Me- rivulana (Scopoli).. 166, 183
Dunnough)------ 169, 186 roseomaculana (Her-
buckellana (McDun- rich-Schaefer) _----- 188
nough)_-_-.-- 117, 169, 188 sehulziana (Fabri-
buekellana albidula clus) 2222 23 166, 169, 183
eimrich=== 22222 167, 189 septentrionana (Cur-
caesialbana (Zeller)... 180 tis) eee eee 168, 185
earolana (MecDun- septentrionana (Mosch-
nough) 23s 167, 180 len) lee eS 186
castorana (McDun- siderana (Treitschke). 172
nough)s=222 Slane 182 siderana chalybeana
ecespitana (Htib- (Walsingham) _-_-- 168,172
ner) == a 168, 179 similisana (Walker)--- 180
conchana (Hiibner)_-. 166 sordidana (McDun-
constellatana (Zel- MOUGN) Sean eeeen ee = 168, 173
ler))=22 Se Seeeke 22 168, 174 striana (Denis and
coronana Kearfott..-. 174 Schiffermiiller) _._--- 166
coruscana (Clem- tessellana (Packard)... 184

ens) 22. eeies oe 168, 175

turfosana (Moschler)-. 184

Page
OLETHRRU TINA oe on ee ee ie 4,76
Olivaceanum (Fernald)-_-..-.-.------------- 133, 137
Opadig\ Guenbevs saa ae 26
ornatana (Kearfott)_..-.....------.--=.---.-- 148
Orthotaenia Stainton (not Stephens) -----.-_- 100
Orthotaenia-Stephons..- .2-. 2-2. 22- 22-5... 166
osmundana (Fernald) --..-..-----..-.------ 168, 170
oxycedrana (Milliére).........-...--.-.-.--. 21
Packardi Zeller cece. s ee LS eS 5, 27, 29
Palle ll bergeeee se eae ot cc Bee eee cee 20
pallidibasalis Heinrich--_----_---------------- 44
IPAM Meneset DNEleee es sees a aac eee OS 20
parmatana.(Clemens) =-225 2222) Pbk = Beets 43, 47
Palla welmrnen ces oaoe ea os Slt Se 21, 23
perductana Cwalker) = 2cc-1n.1 221 eee 127
EN UG NG CACNEY) pe cesses el 25
FEV TION TETAS COTES NS Sa gE gS 123
permixtana (Hubner). ....20i22522ce 222-22. 88
permundanum Clemens.-------.---------- 133, 155
petiverella (Linnaeus) -.-.---.--...--.------- 9
Phaecasiophora Grote______.-...-.--------- 78, 126
confixana (Walker). 126,
127

mutabilana (Clem-
ens) s/o. Sete 126, 127
niveiguttana Grote. 126,
127
perductana (Walker) 127
PAtirisEEbU NERS Veen lee Se SE 166
Phthoroblastis Lederer - -...------------------ 20
ARCEMEGBUSCKS acct eee cc tcc oS 107
piperans (BuUSsCk)...-<.....--s beeen ee 11,15
miperana.(cearfott)'. occ SSS 5, 43, 63
miacerann (Rseariott) hat sete «= = 2 37
planiloqua (Meyrick) -......-.--.------------ 11
plumbagana (Treitschke) - ..........-------- 9
pianonne Ch ermal) — oe 2 oc 15
plumbana) (Seepoli)isp2ce0ee28% - 22 <2 =---- 9,15
plommeriana. (Busck)/=-2-5 ------- 2 .5-222-=- 19
AIDEN CLION) eee re eee et eee eS 179
polluxana (McDunnough)---...----...-.-. 167, 180
PolychrosispRAgOnO.o222>=-5 tack ese- === 76, 87
aemulana Heinrich. -._. 89, 94
ambrosiana Kearfott - -- 94
aruncana Kearfott-____ 89, 95
blandula Heinrich. ___- 89, 96
botrana Fernald ------- 90

botrana (Schiffermiil-
len) 2s-2222 se 87, 88
carduana Busck-__-.-- 89, 96
eyclopiana Heinrich--._ 88, 97
cypripediana Forbes__. 89, 92
liriodendrana Kearfott. 89
magnoliana Kearfott--- 89
monotropana Heinrich 89, 91
rhoifructana Kearfott_. 89, 92
signifera Meyrick_--_--- 93
slingerlandana Kearfott 89, 95
spiraeifoliana Heinrich. 89, 93
vernoniana Kearfott_.. 89, 94
viteana (Clemens)-.- 5, 89, 90
vitivorana (Packard) --- 90
yaracana Kearfott---__- 89, 93
pomonana Treitschke...-=--.---.---=-=------ 70
pomonelay (uinnsAels))=—— 2 =-5=2 026 ---5-—- 5, 69, 70

MopulanapBuschkse ses. koe = Sa 42, 52

Page

populana: (Fabricius) £20 22222 222 eee 20, 21
priapeis Heinrich Y= Ss see 82, 83, 86
primariang, (Walker)== 2022-2.) eee ee 185
prosperana (Kearfott)-....-.-.... 22 ss 42, 57
provans, Kearfott...22..-- es See 191
prunivora;(Walsh) =2.222-.-.222 ee 5, 27, 30
Pseudotomia Stephens. ..........------.----- 20
punctanum Walsingham-.-_....._-_______. 132, 141
puncticostana (Walker). ........._..-__.-- 168, 176
punctidiscanum) Dyari. 22-2) ae ee 71, 72
pyrtcolana.(Murtieldt)..... 0 ee 29
Pyrodes' Quenéexs =< =33.5) ee ee 20
quadrifidum.Zoller....----- ===. 132, 145
quebecense Heinrichs: 29 Soe 2 ee 156
radicicolana Walsingham...-..........-.---- 11,14
radicolang (Hernald).2..22 Sees 14
rana Forbes s3-2222)---U Deets eee 44, 46
ratijera (Meyrick). 2<-<2:=--.222-22). Se 53
regiana (Zeller): 22--<+.-2 Rea ee 21
Femovana) Gcearfott)=o.ce22=-2e tees 114, 122
rhediella (Wlerck) oss 2o ee ee 20, 21
rhotfructana:Kearlott.--2.22.22222-2 Paes 89, 92
Rieuls Heinrich: <2 =. -.2) see 6,18
maculana (Fernald) -_.-___._- 18

rivulana (Scopoli).--__-__- eget te seers eas 166, 183
rosaochreana (Kearfoot) -........-.-------. 112, 113
roseomaculana (Herrich-Schaefer) ___----_---- 188
Rorana Stephens 22222452252 see 166
rubipunctans (Kearfott) =~ 2-2--- == 528 101, 102
rusticanum McDunnough---_.-------_----- 133, 139
sabastianniae Riley:=.224 3 eee 70
salicella:(iinnaeus)*2<=-=-252-e ee 161
Saltitans Westwood: 22s. see eee eee 70
Satronia Heinrich=\=- <==" <3: eee 6,17
tantilla Heinrich________- 17

saundcrsana (Kearfott)_.-._------------------ 39
schulziana (Fabricius) -....------------ 166, 169, 183
Selaphila “Treitschkes= 2-2-2 = 2 =e eae 76, 123

branderiana (Linnaeus) 123
duplex (Walsingham).. 124

thallasane (MecDun-
NOUN) ese ee 124
wahlbomiane (Lin-

TAC) = -22 oa 2ocaeeeene 123

SClOLANUIM) ELeinri Chis = ae ene ae aoe en 132, 147
scirpana Herrich-Schaefer---_-------------..--- 86
scitanaGWalken)as-2+- 4222 === See = nano aoa 36
seriptana (hibner)oa2e---s2s-ses-- aa eaeee 113, 119
Sedatana, (Buseck) S02 28 —a-s ese ee ane eee 10, 15
‘Selenodes) Guentée--2-- -= 5 222-2 osscessene ee 166
semifasciana (Haworth)-_-.--.---------------- 113
separatana (Kearfott)_.......---.------------ 162
septentrionana (Curtis)_--.---------------- 168, 185
septentrionana (Moschler) -------------------- 186
seplicola (Meyrick) = a= 2-2-2 =-==== aeee ae 49
Sereda\Heinrich:--22.- 4) = eee 6, 25
lautana (Clemens) - -------- 25

perfluana (Zeller) ----.------ 25

sericoranum Walsingham_------------- 133, 135, 152
Sericoris: Dreitsehke:- 22222 22s ee eee 166
servillana (Duponchel) -.-------------------- 41
siderana, (CU rettschKe) 2225-2) 92 —e a ee ene 172
SioniferayEelHe] Chisa= ass see eae 21, 22
sigmijen® IMOVTICKS = ee nese 93
BiMIsAMNG |W AlKeL) ee nese es aoe ae 180

216 INDEX
Page Page
simpsoni (Busck) (variety of pomonella (Lin- toreutay (Grote) eso o es es ne ey 43, 62
MIAOLES)) ese eee aes Ia ah PUNO leh 70,71 | transferrana (Walker) --__-------------.------ 78
Simulana(@lemiens) ese ae ees 11,12 | trauniana (Schiffermiiller)_......-._..._..... 20, 21
SINISETA CELINE Cheese eet neeees 82,87 | trepidulum Heinrich-_-__-_........._-.. 132, 140, 147
slingerlandana Kearfott__.-.-..-------.------ 89595) |) tristrigana) (@lemens) 2522.2 2 ee 27, 39
SORMESCENS MIC VTICK) ooo ne = oe ee as 14 | troglodanum McDunnough.-__....--..___- 133, 159
sordidana (McDunnough)-___-------------- 168.173 tunfosana (Mischler) 2-222 eee 184
sordulenta Heinrich: — 22222222 TOT AOD ie Gye LUS EM Gir ri Chae er a noe 79, 81
sororeculana (Zetterstedt) ....-.----_-.------- 19};|| unticana:(Hubner) 2-222 ee ee 124, 125
spiniana (Duponchel).....-..--------.--=-+-- Die |; usticang (Zeller) ==22-522-s-2s— eee 161
spiraeifoliana Heinrich__.-..-.-=2-2==-2--===- 89,93 | valdanum McDunnough-----......-..--.. 135, 153
splendidulana (Guenée) -..------------------ 21) || vane: (Keartott) =-<- 2- a e e 31
Stigmonota Guenee:.- 2522-5 ate ee 26.1) vancouverana: (Neariott) non.) pene ee 37
striana (Denis and Sdhiffermiiller)_.....-__-- 166. |) wablegansa) (Ehubner) oss. = 22 aoe 162, 164
strigosalemrichs22 = 2-3 = ano an tee ee 115,118; || vernoniana Kearfott__.-.-.-- =u. 222-22 89, 94
Strophedra Herrich-Schaeler__.--.------------ 20 | versicoloranum Clemens------.-------.---- 135, 154
submissanum McDonnough- ---.---------- 1315156) ||) ‘verutana-Zellers--2-= = 2 eee 83, 84
subnubilum) Heinrich. -----22-2- esse 134,188 | vetulana (Walsingham) _._-_.........--.-.--- 105
succedana (Walsingham) .-......-.--2<-s=3.-- ‘67¢4|, Niteana(@lemens)=-25"- = 2en 2 sed Pa ee 5, 89, 90
Talponiaseveinrich) =- ees. - 2s ase eee nes 6.19) |haitivorana:. (Packard) =o ~ = - 2 eee eee eee 90
plummeriana (Busck)-_._- 19. |} vatranaWialsingham-* -_ 9625. soz cee ee 27, 32
tanay(Kearfott) <2 22-22 =2- 5-2 eS eee 42,56, | vulgana (McDunnough)-__._-_........---.-.. 108
Taniva celeinrich oso 2 ons ooo oko oo eee 769106") wahlbergianayZelleri--2-22--2 == S22 aceon 129
abietana (Fernald)-_--.-._--- 107 | wahlbomiana (Linnaeus) -..----.----.-------- 123
albolineana \(Kearfott)) = 15,106,107, || wanai@Kearfott)--- 2 ese2- 2.5 ee 33
DICELe (BUSCK) 2) ase 107 | wellingtoniana Kearfott_.................-.-- 191
Tantillaseeinrl Chis 2s *ecs yoo Se | zanthospora (Meyrick)2-----—-]—-- eae eee 31
Lenebpricum Hemrich: <2-¢-s22-22-2--e55- 131,144 || yaracanaKearfotts2222 S222 eee see 89, 93
terminanum McDunnough (variety of con- youngana: (Kearfott)s222= sass = eee 44, 52
cinnanum (Clemens) )222_2- 32 --2.-<-4_- 131,158 | youngana (McDunnough)----------------- 115, 116
tertiana (McDunnough)-.----.-----....-.. 15 U7, |} zane (Kearfott) 22-22" se Se eee ee 31
LESSCLLOMAN CL ACKALG) sae sea eese ee anene eens 184 | zellerianum (Fernald) ----.----.-----.----- 133, 139
texanana (Walsingham) 2- 22-2222 —-- 22-2. 23004541, {| Zomaria Heinrich=--- 2-22-22 ---sseen eee eae 76, 111
thaiasits! (Meyrick) otesenss se sees eee 75 andromedana (Barnes and
thallasana (McDunnough)--._--=------------ 124 McDunnough).----.--- 112, 113
Tiaeinrich==_ esse ase el eee eee 76, 108 interruptolineana (F er-
vulgana (McDunnough)------- 108 nald) 2-2 2.5.2622542 111, 112
tiliantimpllemnrich= ca css" s_seene-= aoe eee 132, 146 rosaochreana (Kearfott). 112, 113

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